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Full-Text Articles in Forest Sciences

Stabilization Of Pfas-Contaminated Soil With Sewage Sludge- And Wood-Based Biochar Sorbents, Erlend Sørmo, Clara Benedikte Mader Lade, Julie Zhang, Alexandros G. Asimakopoulos, Geir Wold Åsli, Michel Hubert, Aleksandar I. Goranov, Hans Peter H. Arp, Gerard Cornelissen Jan 2024

Stabilization Of Pfas-Contaminated Soil With Sewage Sludge- And Wood-Based Biochar Sorbents, Erlend Sørmo, Clara Benedikte Mader Lade, Julie Zhang, Alexandros G. Asimakopoulos, Geir Wold Åsli, Michel Hubert, Aleksandar I. Goranov, Hans Peter H. Arp, Gerard Cornelissen

Chemistry & Biochemistry Faculty Publications

Sustainable and effective remediation technologies for the treatment of soil contaminated with per- and polyfluoroalkyl substances (PFAS) are greatly needed. This study investigated the effects of waste-based biochars on the leaching of PFAS from a sandy soil with a low total organic carbon content (TOC) of 0.57 ± 0.04% impacted by PFAS from aqueous film forming foam (AFFF) dispersed at a former fire-fighting facility. Six different biochars (pyrolyzed at 700–900°C) were tested, made from clean wood chips (CWC), waste timber (WT), activated waste timber (aWT), two digested sewage sludges (DSS-1 and DSS-2) and de-watered raw sewage sludge (DWSS). Up-flow column …


Influence Of Abiotic Drivers And Plant Community Interactions On Bald Cypress (Taxodium Distichum) Seedlings: Implications For Restoration, Victoria Ellis Aug 2023

Influence Of Abiotic Drivers And Plant Community Interactions On Bald Cypress (Taxodium Distichum) Seedlings: Implications For Restoration, Victoria Ellis

Biological Sciences Theses & Dissertations

Forested freshwater wetlands are valuable ecosystems that provide habitat for numerous species, sequester carbon, and act as sinks for excess water and nutrients. Historically, these ecosystems have been heavily degraded by anthropogenic activities leading to loss of ecosystem services and a desire to restore freshwater forested wetland habitat. Thus, science-backed approaches for the restoration of freshwater forested wetlands are necessary to ensure restoration goals are met. This body of research employed the Stress Gradient Hypothesis to test whether a multi-species planting approach using Juncus effusus (L.) (soft rush) could facilitate the survival of Taxodium distichum (L.) Rich. (bald cypress) seedlings …


Dendrochronology In The Blackwater Ecological Preserve, Sarah Durham, Arvind Bhuta Jan 2023

Dendrochronology In The Blackwater Ecological Preserve, Sarah Durham, Arvind Bhuta

College of Sciences Posters

The Blackwater Ecological Preserve (BEP), located in southeastern Virginia, holds significant ecological importance as it contains the northernmost intact community of Pinus palustris, longleaf pine. At the BEP, we cored longleaf pine and recorded sample data including tree height and diameter at breast height (DBH). Two sites on the BEP, the 20’s series and 40’s series, were found to correlate within, but not between, sites. Statistical and qualitative analyses were conducted to find differences between the sites significant enough to affect correlation within a chronology. This data is being incorporated into a larger dendrochronology and climate analysis study on …


Territory Inheritance And The Evolution Of Cooperative Breeding In The Acorn Woodpecker, Walter D. Koenig, Joseph Haydock, Hannah L. Dugdale, Eric L. Walters Jan 2023

Territory Inheritance And The Evolution Of Cooperative Breeding In The Acorn Woodpecker, Walter D. Koenig, Joseph Haydock, Hannah L. Dugdale, Eric L. Walters

Biological Sciences Faculty Publications

There are two main hypotheses for why offspring in cooperatively breeding taxa delay dispersal and remain on their natal territory rather than disperse. First, ecological constraints may force offspring to remain on their natal territory until a reproductive opportunity presents itself in an otherwise saturated habitat. Alternatively, delaying dispersal and helping kin may increase an offspring's inclusive fitness. One means by which offspring might enhance their direct fitness by delaying dispersal is by inheriting breeding status on their natal territory. Such territory inheritance regularly occurs in acorn woodpeckers, Melanerpes formicivorus, a species whose social groups consist of a cooperatively …


Wetlands In Our Backyard: A Review Of Wetland Types In Virginia State Parks, Kirsten Bauer, Benjamin K. Campbell Oct 2022

Wetlands In Our Backyard: A Review Of Wetland Types In Virginia State Parks, Kirsten Bauer, Benjamin K. Campbell

Virginia Journal of Science

Wetlands constitute a significant component of Virginia’s natural resources and heritage. Though historically they have been discounted—and often denigrated—the exceptional value of wetlands is currently growing in recognition and appreciation. In addition to the value provided by extracted resources and ecological regulation, wetlands also offer people the opportunity to enrich themselves through cultural, educational, and recreational pursuits. The state parks of Virginia provide access to a variety of ecosystems, including a wide array of wetland types. In this review, we document the diversity of wetlands in Virginia State Parks through a typology that groups wetland systems into the three principal …


Remote Detection Of Disturbance From Motorized Vehicle Use In Appalachian Wetlands, Walter Smith Oct 2021

Remote Detection Of Disturbance From Motorized Vehicle Use In Appalachian Wetlands, Walter Smith

Virginia Journal of Science

Wetland disturbance from motorized vehicle use is a growing concern across the Appalachian coalfields of southwestern Virginia and portions of adjacent states, particularly as both extractive industries and outdoor recreation development expand in regional communities. However, few attempts have been made in this region or elsewhere to adapt approaches that can assist researchers and land managers in remotely identifying and monitoring wetland habitats disturbed by motorized vehicle use. A comparative analysis of wetlands impacted and unimpacted by off-road vehicle activity at a public recreation area in Tazewell County, Virginia was conducted to determine if and how a common, satellite-derived index …


Camera Trap Survey Suggests Forestry And Prescribed Burns Attract Wildlife, But May Not Enhance Diversity, Melissa S. Vilgats, Ryan Ott, Stephanie S. Coster Oct 2021

Camera Trap Survey Suggests Forestry And Prescribed Burns Attract Wildlife, But May Not Enhance Diversity, Melissa S. Vilgats, Ryan Ott, Stephanie S. Coster

Virginia Journal of Science

This study explored whether habitat management techniques such as forest thinning and burning promoted biodiversity. Fifteen camera trap stations were established at Fort A.P. Hill in Bowling Green, VA across forest stands with low, medium, and high basal area. Camera traps were deployed for a total of 532 trap nights, and trap success and species diversity were calculated using Shannon’s index. At each site, the distance to trafficable roadways and water sources, vegetation composition, and the percent groundcover, canopy cover, and understory were measured. The cameras captured nine species and recorded a total of 398 trap events. Linear regression was …


Focus Stacking Images Of Morphological Character States For Differentiating The Adults Of Ixodes Affinis And Ixodes Scapularis (Acari: Ixodidae) In Areas Of Sympatry, Robyn M. Nadolny, Marcée Toliver, Holly D. Gaff, John G. Snodgrass, Richard G. Robbins Jan 2021

Focus Stacking Images Of Morphological Character States For Differentiating The Adults Of Ixodes Affinis And Ixodes Scapularis (Acari: Ixodidae) In Areas Of Sympatry, Robyn M. Nadolny, Marcée Toliver, Holly D. Gaff, John G. Snodgrass, Richard G. Robbins

Biological Sciences Faculty Publications

Adult females and males of Ixodes affinis and Ixodes scapularis are illustrated by focus stacking image photography, and morphological character states are described that reliably differentiate the two species. In conjunction with other environmental cues, such as the questing phenology of adults, these characteristics will enable the rapid identification of adults of either sex along the southern Coastal Plain of the United States, where these species are sympatric.


Dietary Protein Content And Digestibility Influences Discrimination Of Amino Acid Nitrogen Isotope Values In A Terrestrial Omnivorous Mammal, John P. Whiteman, Mauriel Rodriguez Curras, Kelli L. Feeser, Seth D. Newsome Jan 2021

Dietary Protein Content And Digestibility Influences Discrimination Of Amino Acid Nitrogen Isotope Values In A Terrestrial Omnivorous Mammal, John P. Whiteman, Mauriel Rodriguez Curras, Kelli L. Feeser, Seth D. Newsome

Biological Sciences Faculty Publications

RATIONALE: Ecologists increasingly determine the δ15N values of amino acids (AA) in animal tissue; "source" AA typically exhibit minor variation between diet and consumer, while "trophic" AA have increased δ15N values in consumers. Thus, trophic-source δ15N offsets (i.e., Δ15NT-S) reflect trophic position in a food web. However, even minor variation in δ15Nsource AA values may influence the magnitude of offset that represents a trophic step, known as the trophic discrimination factor (i.e., TDFT-S). Diet digestibility and protein content can influence the δ15N values …


Assessment Of Factors Influencing Migratory Landbird Use Of Forested Stopover Sites Along The Delmarva Peninsula During Autumn Migration, J. Andrew Arnold Dec 2020

Assessment Of Factors Influencing Migratory Landbird Use Of Forested Stopover Sites Along The Delmarva Peninsula During Autumn Migration, J. Andrew Arnold

Biological Sciences Theses & Dissertations

Autumn migration is a time when billions of birds move from breeding grounds in North America to wintering grounds in Central and South America, with many individuals relying on stopover habitats en route for resting and refueling purposes. These stopover sites are critical to the survival of the hundreds of species of migratory landbirds that migrate annually, and thus identifying important stopover sites is a high priority for conserving such taxa. The Delmarva Peninsula; a coastal region of Delaware, Maryland, and Virginia along the mid-Atlantic flyway; consists of forested habitats with ample food and shelter that likely serves as quality …


Artificial Perches As A Technique For Enhancing Tropical Forest Restoration: A Case Study From The Dominican Republic, Spencer Schubert, Ally S. Lahey, Ashley R. Weisman, Eric L. Walters Apr 2020

Artificial Perches As A Technique For Enhancing Tropical Forest Restoration: A Case Study From The Dominican Republic, Spencer Schubert, Ally S. Lahey, Ashley R. Weisman, Eric L. Walters

College of Sciences Posters

Recovering secondary forests on degraded agricultural lands represents a promising opportunity to offset global carbon emissions as well as increasing local biodiversity and ecosystem services. In the insular tropical forests of the Caribbean, frugivorous birds are the primary seed dispersers for most native woody plants and have a large influence on regeneration dynamics during forest succession. In 2017, we initiated an experimental forest restoration program incorporating artificial perches on private farms within the Rio Yaque del Norte watershed in La Vega province, Dominican Republic. Five restoration plots (0.15–0.25 ha) were constructed in pastures near deforested streams. In each plot, 6-12 …


Apparent Resilience To Fire Of Native Bee (Hymenoptera: Apoidea) Communities From Upland Longleaf Pine Forests In Louisiana And Mississippi, Sara A. Simmons, Janice L. Bossart Jan 2020

Apparent Resilience To Fire Of Native Bee (Hymenoptera: Apoidea) Communities From Upland Longleaf Pine Forests In Louisiana And Mississippi, Sara A. Simmons, Janice L. Bossart

Biological Sciences Faculty Publications

Controlled burning is an essential tool for restoration and management of Pinus palustris (Longleaf Pine) habitats, yet effects of controlled burning on insect species, including pollinators, are rarely considered in conservation planning. We used blue vane traps to sample native bees (Hymenoptera: Apoidea) at recently burned and unburned sites in 2 Longleaf Pine upland forests in Mississippi and Louisiana. Our objective was to quantify short-term effects of controlled burns given fire-return intervals of 1-2 years are now regularly employed to manage Longleaf Pine woodlands. We sampled during 2016 and 2017 and collected 1777 native bees, representing 43 species. Recent fire …


Habitat Saturation Results In Joint-Nesting Female Coalitions In A Social Bird, Sahas Barve, Walter D. Koenig, Joseph Haydock, Eric L. Walters Jan 2019

Habitat Saturation Results In Joint-Nesting Female Coalitions In A Social Bird, Sahas Barve, Walter D. Koenig, Joseph Haydock, Eric L. Walters

Biological Sciences Faculty Publications

Joint nesting by females and cooperative polyandry—cooperatively breeding groups with a male-biased breeder sex ratio—are little-understood, rare breeding systems. We tested alternative hypotheses of factors potentially driving these phenomena in a population of joint-nesting acorn woodpeckers (Melanerpes formicivorus). During periods of high population density and thus low independent breeding opportunities, acorn woodpecker females formed joint-nesting coalitions with close kin. Coalitions were typically associated with groups with a male bias. We found strong evidence for both inter- and intrasexual conflict, as joint nesting conferred a fitness benefit to some males, a significant fitness cost to females, and no gain in per …


The Effects Of Climate On Radial Growth Of Disjunct Northern White Cedar (Thuja Occidentalis L.) In Virginia, Joshua A. Kincaid Oct 2017

The Effects Of Climate On Radial Growth Of Disjunct Northern White Cedar (Thuja Occidentalis L.) In Virginia, Joshua A. Kincaid

Virginia Journal of Science

Understanding the geographic range and growth of species is essential for effective land management in a landscape affected by anthropogenic activity and climate change. Climate change is expected to alter the distribution and growth of many tree species in eastern North America, including northern white cedar (Thuja occidentalis L.). This research examined the effects of climate on radial growth of T. occidentalis in disjunct populations south of its continuous range margin in eastern North America. A T. occidentalis tree-ring chronology was developed and examined for growth-climate interactions. Mean sensitivity of the T. occidentalis chronology was within the range of …


Section Abstracts: Agriculture, Forestry And Aquaculture Apr 2016

Section Abstracts: Agriculture, Forestry And Aquaculture

Virginia Journal of Science

Abstracts of the Agriculture, Forestry and Aquaculture Section for the 94th Annual Virginia Academy of Science Meeting, May 18-20, 2016, at University of Mary Washington, Fredericksburg, VA.


Forest Diversity And Disturbance: Changing Influences And The Future Of Virginia’S Forests, Christine J. Small, James L. Chamberlain Oct 2015

Forest Diversity And Disturbance: Changing Influences And The Future Of Virginia’S Forests, Christine J. Small, James L. Chamberlain

Virginia Journal of Science

The Virginia landscape supports a remarkable diversity of forests, from maritime dune woodlands, swamp forests, and pine savannas of the Atlantic Coastal Plain, to post-agricultural pine-hardwood forests of the Piedmont, to mixed oak, mesophytic, northern hardwood, and high elevation spruce-fir forests across three mountain provinces in western parts of the state. Virginia’s forests also have been profoundly shaped by disturbance. Chestnut blight, hemlock woolly adelgid, emerald ash borer, and other pests have caused declines or functional extirpation of foundation species. Invasive plants like multiflora rose, Oriental bittersweet, and Japanese stiltgrass threaten both disturbed and intact forests. Oaks and other fire-dependent …


Section Abstracts: Agriculture, Forestry And Aquaculture May 2015

Section Abstracts: Agriculture, Forestry And Aquaculture

Virginia Journal of Science

Abstracts of the Agriculture, Forestry and Aquaculture Section for the 93rd Annual Virginia Academy of Science Meeting, May 2015, at James Madison University


Amphibian And Small Mammal Assemblages In A Northern Virginia Forest Before And After Defoliation By Gypsy Moths (Lymantria Dispar), Joseph C. Mitchell Jan 2015

Amphibian And Small Mammal Assemblages In A Northern Virginia Forest Before And After Defoliation By Gypsy Moths (Lymantria Dispar), Joseph C. Mitchell

Virginia Journal of Science

The introduced European gypsy moth (Lymantria dispar) caused substantial defoliation and mortality of oak trees along the North Fork of Quantico Creek in Prince William Forest Park, Prince William County, Virginia, U.S.A., in 1989 and the early 1990s. Results of a drift fence/pitfall study conducted in 1988 were compared to those obtained from the same technique in the same areas in 1993 to elucidate whether the amphibian and small mammal assemblages had changed over time. Number of Lithobates sylvaticus increased significantly in 1993, but the numbers of Lithobates clamitans and Plethodon cinereus were significantly higher in 1988. Total …


Section Abstracts: Agriculture, Forestry, And Aquaculture Apr 2014

Section Abstracts: Agriculture, Forestry, And Aquaculture

Virginia Journal of Science

Abstracts of the Agriculture, Forestry, and Aquaculture Section for the 92nd Annual Virginia Academy of Science Meeting, May 2014, at Virginia Commonwealth University in Richmond, Virginia


Thresholds Of Change In Decomposition Rate Along A Dune/Swale Transect On A Virginia Barrier Island, Dominic J. Graziani Oct 2012

Thresholds Of Change In Decomposition Rate Along A Dune/Swale Transect On A Virginia Barrier Island, Dominic J. Graziani

Biological Sciences Theses & Dissertations

Aboveground and belowground decomposition rates were determined along a barrier island dune/swale transect located on the Virginia Coast Reserve-Long Term Ecological Research Site using litterbags and wooden dowels. The objective was to determine the influence of fine scale changes in the environment on decomposition to identify any potential thresholds affecting decomposition rate. Wax myrtle (Morella cerifera L. Small) leaves and dowels of southern yellow pine wood were used as standard substrates to evaluate environmental influences on decay. Aboveground (F=6.494, p < 0.0001) and belowground (F=5.705, p < 0.0001) decay rates (yr-1) showed significant variation among litterbag/dowel locations. Aboveground decay rates (yr-1) ranged from 0.339 (Upper Dune station) …


Rats As Forest Pests In Southeastern Virginia: Girdling By The Hispid Cotton Rat Sigmodon Hispidus As A Significant Source Of Mortality Of Loblolly Pines (Pinus Taeda) In A Successional Pine Forest, Robyn M. Nadolny Jul 2011

Rats As Forest Pests In Southeastern Virginia: Girdling By The Hispid Cotton Rat Sigmodon Hispidus As A Significant Source Of Mortality Of Loblolly Pines (Pinus Taeda) In A Successional Pine Forest, Robyn M. Nadolny

Biological Sciences Theses & Dissertations

The hispid cotton rat (Sigmodon hispidus) is a common field rodent throughout the southeastern US, where volunteer loblolly pine trees (Pinus taeda) invade open space and begin the process of ecological succession from field to pine forest. Recent analysis of the diet of S. hispidus indicates that loblolly pine bark is stripped and eaten during the winter months. In this study, we explored the extent of rodent girdling on a 1.23 ha grid in a successional pine forest in southeastern Virginia. During the winter of 2005 we observed damage to 65% of trees in our study area, with 98% …


Section Abstracts: Agriculture, Forestry And Aquaculture Science Apr 2011

Section Abstracts: Agriculture, Forestry And Aquaculture Science

Virginia Journal of Science

Abstracts of the Agriculture, Forestry, and Aquaculture Sciences section for the Virginia Academy of Science 89th Annual Meeting, May 25-27, 2011, University of Richmond, Richmond, Virginia.


Section Abstracts: Agriculture, Forestry, And Aquaculture Science Apr 2010

Section Abstracts: Agriculture, Forestry, And Aquaculture Science

Virginia Journal of Science

Abstracts of the Agriculture, Forestry, and Aquaculture Science Section for the 88th Annual Meeting of the Virginia Academy of Science, May 20-21, 2010, James Madison University, Harrisonburg, VA.


Section Abstracts: Agriculture, Forestry And Aquaculture Science Jul 2009

Section Abstracts: Agriculture, Forestry And Aquaculture Science

Virginia Journal of Science

Abstracts of the Agriculture, Forestry and Aquaculture Section for the 87th Annual Meeting of the Virginia Academy of Science, May 27-29, 2009, Virginia Commonwealth University, Richmond, VA.


Topographic Factors Affecting The Tree Species Composition Of Forests In The Upper Piedmont Of Virginia, Rachael C. Brown, Todd S. Fredericksen Apr 2008

Topographic Factors Affecting The Tree Species Composition Of Forests In The Upper Piedmont Of Virginia, Rachael C. Brown, Todd S. Fredericksen

Virginia Journal of Science

There are many factors that influence forest species composition and many are linked to topographical features. This study, conducted on the Ferrum College campus in the Upper Piedmont Physiographic Province of Virginia revealed three major forest types associated with topographic factors using cluster analysis and detrended correspondence analysis . The first type of forest occurred mostly on northeastern slopes on toe slope topographic positions and was mainly composed of tulip tree (Liriodendron tulipifera) and red maple (Acer rubrum). The second type of forest was found on shoulder and side slope positions and was composed mostly of …


Trees On K-12 School Campuses In Virginia, Jeffrey L. Kirwan, P. Eric Wiseman, John R. Seiler Apr 2007

Trees On K-12 School Campuses In Virginia, Jeffrey L. Kirwan, P. Eric Wiseman, John R. Seiler

Virginia Journal of Science

Trees and saplings growing on K-12 school campuses were investigated in 105 school districts across Virginia. There were 2812 trees (>12.5 cm stem diameter at 1.4 m above ground level) inventoried across all campuses. The mean and median campus tree population was 27 and 18, respectively. Loblolly pine (Pinus taeda L.) was the most abundant species, accounting for 11% of all inventoried trees. Red maple (Acer rubrum L.) was the most frequently inventoried species, present on 44% of the campuses. Sapling (trees with 2.5-12.5 cm stem diameter at 1.4 m above ground level) populations were similar …


Response Of Small Mammal Populations To Fescue Hayfield Conversion To Native Warm Season Grasses In Bath County, Virginia, Michael T. Mengak Jan 2004

Response Of Small Mammal Populations To Fescue Hayfield Conversion To Native Warm Season Grasses In Bath County, Virginia, Michael T. Mengak

Virginia Journal of Science

I investigated the effect on small mammal populations of converting an existing fescue (Festuca arundinacea) hayfield to switchgrass (Panicium virgatum) on the George Washington National Forest at Hidden Valley in Bath County, Virginia. Native warm season grasses are thought to provide better habitat than fescue pastures for Northern Bobwhite (Colinus virginianus) and several species of grassland birds as well as herbivorous small mammals. I established one live-trapping grid and conducted trapping (pre-treatment) in both the switchgrass (treatment) and the fescue (control) field in March and May 1997. The treatment field was sprayed with glyphosate …


Longevity Record For A Wild Allegheny Woodrat (Neotama Magister) In West Virginia, Michael T. Mengak, Steven B. Castleberry, W. Mark Ford, Nikki L. Castleberry, Jane L. Rodrigue Oct 2002

Longevity Record For A Wild Allegheny Woodrat (Neotama Magister) In West Virginia, Michael T. Mengak, Steven B. Castleberry, W. Mark Ford, Nikki L. Castleberry, Jane L. Rodrigue

Virginia Journal of Science

The Allegheny woodrat (Neotoma magister) is found throughout much of the central and southern Appalachians and adjacent portions of the Interior Highlands. Allegheny woodrats have declined in the northern portions of their range and are state-listed as threatened, endangered or sensitive species of concern in every state where they occur. Until recently, biologists have had to rely on biological data collected from the closely related eastern woodrat (N. floridana) because of limited research on the Allegheny woodrat. We have been studying the ecology and natural history of woodrats in Virginia and West Virginia since 1990. On …


Decomposition Dynamics In Restored And Naturally Recovering Atlantic White Cedar (Chamaecyparis Thyoides) Wetlands, Edward Ratcliffe Crawford Jan 2002

Decomposition Dynamics In Restored And Naturally Recovering Atlantic White Cedar (Chamaecyparis Thyoides) Wetlands, Edward Ratcliffe Crawford

Biological Sciences Theses & Dissertations

Restoration efforts and ecologically based management practices for Atlantic white cedar wetlands have recently focused on soil organic matter sequestration and aggradation as critical ecosystem functions in the maintenance of this imperiled community type. This work addresses how developmental stage, litter quality and environmental conditions influenced above ground leaf litter and below ground root decay in naturally regenerating and restored Atlantic white cedar wetlands. Using standard litterbags for leaf litter and a modified litterbag technique for roots, decay dynamics of naturally regenerating and restored Atlantic white cedar stands were measured to compare ecosystem development trends with restoration conditions. Effects on …


Ice Storm Damage To Virginia Coastal Plain Forests During The Christmas 1998 Ice Storm, Peter Elstner, Stewart Ware Apr 2001

Ice Storm Damage To Virginia Coastal Plain Forests During The Christmas 1998 Ice Storm, Peter Elstner, Stewart Ware

Virginia Journal of Science

On December 23-25, 1998, a major ice storm struck southeastern Virginia. The storm-deposited glaze ice felled trees and limbs, causing a power outage and highway blockage. Between February and April, 1999, we recorded occurrence, severity, and type of damage to trees over 2.5 cm dbh in nine mostly gently sloping plots in Matoaka Woods at the College of William and Mary. Frequency and severity of damage varied with species and with size of trees. Canopy damage occurred in 75% of large Fagus grandifolia trees, but in only 6% of small Sassafras albidum stems. As a group, small (2.5 to 15 …