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Articles 1 - 30 of 205
Full-Text Articles in Ecology and Evolutionary Biology
Assessing The Responses Of Adult, Juvenile, And Larval Fish Assemblages To The Closure Of The Mississippi River Gulf Outlet, An Artificial Tidal Pass In Southeastern Louisiana, Rebecca Weatherall Cope
Assessing The Responses Of Adult, Juvenile, And Larval Fish Assemblages To The Closure Of The Mississippi River Gulf Outlet, An Artificial Tidal Pass In Southeastern Louisiana, Rebecca Weatherall Cope
University of New Orleans Theses and Dissertations
The creation of the Mississippi River Gulf Outlet (MRGO) in 1963 by the U.S. Army Corps of Engineers (USACE) represents one of the most catastrophic, anthropogenic stressors ever to impact the Lake Pontchartrain estuary in southeastern Louisiana, USA. The artificial tidal pass provided a direct route from the Gulf of Mexico to New Orleans. It allowed for high saline waters to enter Lake Pontchartrain, resulting in detrimental changes to the biotic community of the Lake and surrounding wetland areas. In July 2009, the USACE closed the MRGO in hopes of restoring natural ecosystems. This study assesses changes in the adult, …
Statistical And Comparative Phylogeography Of Mexican Freshwater Taxa In Extreme Aquatic Environments, Lyndon M. Coghill
Statistical And Comparative Phylogeography Of Mexican Freshwater Taxa In Extreme Aquatic Environments, Lyndon M. Coghill
University of New Orleans Theses and Dissertations
Phylogeography aims to understand the processes that underlie the distribution of genetic variation within and among closely related species. Although the means by which this goal might be achieved differ considerably from those that spawned the field some thirty years ago, the foundation and conceptual breakthroughs made by Avise are nonetheless the same and are as relevant today as they were two decades ago. Namely, patterns of neutral genetic variation among individuals carry the signature of a species’ demographic past, and the spatial and temporal environmental heterogeneity across a species’ geographic range can influence patterns of evolutionary change. Aquatic systems …
Soil Development And Vegetation Response To Removal Of A Small Dam, Lassen Volcanic National Park, California, Stephanie Kay Rohdy
Soil Development And Vegetation Response To Removal Of A Small Dam, Lassen Volcanic National Park, California, Stephanie Kay Rohdy
Dissertations and Theses
Dam removal is increasing as dams age, yet little is understood about the pedogenic response to dam removal. This study reports on the edaphic changes of reservoir sediments and vegetation cover one year following removal of a small earthen dam in Northern California. In August 2012, I sampled surface sediments from the former Dream Lake reservoir in Lassen Volcanic National Park, and compared their physical and chemical properties with soil samples from two reference sites. I also estimated percent cover of vegetation in the former reservoir and reference sites. My results show that the reservoir was under reduced conditions, as …
Wood Decomposition In A Warmer World, Emily Elizabeth Austin
Wood Decomposition In A Warmer World, Emily Elizabeth Austin
Doctoral Dissertations
Climatic warming is altering species distributions and ecosystem functions across the globe. Wood is an important carbon pool and the fungal communities in wood are relatively simple compared to those in soil. These factors make decomposing wood an ideal system for exploring the influence of decomposer community on the response of decomposition to warming. My research has focused on the effects of warming wood decomposition rates and wood decomposing communities. Using field and lab- based manipulative experiments and field observations I explore the influence of tree species, wood decomposition stage, geography and warming on fungal community structure and activity. In …
Census And Mapping Of Chorro Creek Bog Thistle In Reservoir Canyon, San Luis Obispo, Ca, Tyler Michael Lutz
Census And Mapping Of Chorro Creek Bog Thistle In Reservoir Canyon, San Luis Obispo, Ca, Tyler Michael Lutz
Biological Sciences
Chorro Creek bog thistle (Cirsium fontinale var. obispoense) is a federally endangered variety of Fountain thistle endemic to western San Luis Obispo County. The U.S. Fish and Wildlife Service knows of nineteen populations, many with multiple colonies. A population was discovered in the Reservoir Canyon Natural Reserve in 2001, but has not been monitored or described since the time of its discovery. In fall of 2013, a census of the population was performed, the four colonies were mapped, and a floristic survey was conducted. A field experiment was initiated to determine if reducing the riparian canopy coverage can …
Homing Behavior In Response To Displacement And Orientation Of The Northern Diamondback Terrapin (Malaclemys Terrapin Terrapin) In Barnegat Bay, New Jersey, Nicole M. Lainhart
Homing Behavior In Response To Displacement And Orientation Of The Northern Diamondback Terrapin (Malaclemys Terrapin Terrapin) In Barnegat Bay, New Jersey, Nicole M. Lainhart
Biology Theses
Increasing urbanization of the Barnegat Bayestuary in New Jersey has subjected northern diamondback terrapins to substantial habitat loss. Understanding whether terrapins have homing behavior, and determining the types of orientation cues they use to aid in this behavior, is important for conservation management. To test their homing behavior, nine non-gravid female terrapins were outfitted with biotelemetry tracking devices and data loggers and were displaced 4 km north and/or south. Eight of nine terrapins successfully returned home; the one terrapin that did not return home was inadvertently captured in a crab pot. Urbanization and shoreline development of the north displacement location …
Lake Michigan Hydrodynamics: Mysis And Larval Fish Interactions, Yutta Wang
Lake Michigan Hydrodynamics: Mysis And Larval Fish Interactions, Yutta Wang
Theses and Dissertations
I studied the interactions between Lake Michigan hydrodynamics (the spring thermal bar) and Mysis, deepwater sculpin, and burbot larvae. The thermal bar is a zone of sinking 4º С water that separates warmer inshore water from colder offshore water. Mysis was a major bycatch of sampling for larval fishes. The density of Mysis did not differ statistically between inshore (about 6º С) and offshore of the thermal bar, but the percentage of Mysis that were newborns was significantly higher inshore (P = 0.007). These "early start" coastal Mysis may have an advantage in growth and survival, but with the risk …
Influence Of Genetic Variation In The Biotic Environment On Phenotypic Variation In A Plant-Feeding Insect, Darren Rebar
Influence Of Genetic Variation In The Biotic Environment On Phenotypic Variation In A Plant-Feeding Insect, Darren Rebar
Theses and Dissertations
While many species spend much of their lives in close association with other organisms, only recently have biologists started to explore the implications of the biotic nature of environments for their role as causes of variation in phenotypes. This means that the genotypes of individuals that constitute the biotic environment may influence the phenotypes of individuals that live in that environment. These are called indirect genetic effects (IGEs) when they occur between conspecifics, and interspecific indirect genetic effects (IIGEs) when they occur between heterospecifics. However, the impact of genetic variation in biotic environments remains largely unknown. I used a member …
Evaluating The Influence Of Environmental Factors On The Rate Of Extra-Pair Matings In Tropical And Temperate Populations Of The House Wren (Troglodytes Aedon), Kaitlin Claire Mckenney
Evaluating The Influence Of Environmental Factors On The Rate Of Extra-Pair Matings In Tropical And Temperate Populations Of The House Wren (Troglodytes Aedon), Kaitlin Claire Mckenney
Theses and Dissertations
Considerable variation exists in the rate of extra-pair matings (EPMs) in birds. Environmental variability likely influences EPM rates within species, but the effects of local environmental factors on EPM rates are largely unpredictable. To determine whether broad-scale environmental factors might be better predictors of EPM rates within species, we quantified levels of extra-pair paternity in the house wren (Troglodytes aedon) in four populations spanning a range of latitude, elevation, and primary productivity (measured by actual evapotranspiration rates). Our results indicated an intermediate and variable level of EPM among populations (6 -31% extra-pair young) that was not significantly affected by 3 …
Sources Of Variability In Agronomic Weed Seed Predation: Time, Space, Habitat, And Hyperpredation, Sonja K. Birthisel
Sources Of Variability In Agronomic Weed Seed Predation: Time, Space, Habitat, And Hyperpredation, Sonja K. Birthisel
Electronic Theses and Dissertations
Weed seed predation is an ecosystem service that benefits farmers by decreasing seedbank inputs, thereby reducing weed pressure in subsequent growing seasons. Seed predation can be considerable, but is highly variable. Sources of variability may include time, space, habitat, and trophic interactions such as hyperpredation. Two experiments were conducted to measure the impacts of these sources of variability on weed seed predation rates in Maine mixed vegetable agroecosystems.
Chapter One of this thesis describes a series of landscape-level field experiments conducted to quantify the effects of time, space, and habitat on seed predation rates. Seed assays, with and without vertebrate …
Effect Of Predator Removal On Greater Sage-Grouse (Centrocercus Urophasianus) Ecology In The Bighorn Basin Conservation Area Of Wyoming, Elizabeth Kari Orning
Effect Of Predator Removal On Greater Sage-Grouse (Centrocercus Urophasianus) Ecology In The Bighorn Basin Conservation Area Of Wyoming, Elizabeth Kari Orning
All Graduate Theses and Dissertations, Spring 1920 to Summer 2023
The decline of greater sage-grouse distribution and population densities across western North America has led conservation, research, and management objectives to focus efforts on understanding sage-grouse populations across their range. The purpose of this study was to gain a better understanding of direct and indirect predation effects on hen survival and nest success of sage-grouse. The project was conducted in Hot Springs and Park Counties in the Bighorn Basin of Wyoming. The study had three main objectives: 1) obtain and quantify the types and impacts of predators on sage-grouse hen survival and nest success, 2) compare the effect predator removals …
Assessing Growth Response To Climate Controls In A Great Basin Artemisia Tridentata Plant Community, Lorenzo F. Apodaca
Assessing Growth Response To Climate Controls In A Great Basin Artemisia Tridentata Plant Community, Lorenzo F. Apodaca
UNLV Theses, Dissertations, Professional Papers, and Capstones
An assessment of the growth response of key vegetative species to climatic variability is vital to identifying possible local impacts on ecosystems faced with imminent climate change. With current climate projections in Nevada predicting a shift to an even more arid climate with greater year-to-year variability, the imperative exists to identify the effects of specific climatic controls on plant growth and to research methods to assess large-scale vegetative changes, especially in more remote areas where readily available data sets may be lacking. This study utilized annual growth ring indices constructed from big sagebrush (Artemisia tridentatassp.tridentata) stems collected in Spring Valley, …
Relationships Between Landscape Factors And Crayfish Density And The Interacting Effects Of Grazing And Increased Sedimentation On Algal Biomass, Katheryn Haley Rose
Relationships Between Landscape Factors And Crayfish Density And The Interacting Effects Of Grazing And Increased Sedimentation On Algal Biomass, Katheryn Haley Rose
Graduate Theses and Dissertations
In recent years in the United States there has been a rapid expansion in anthropogenic sources of sediment in streams including construction, agriculture, and drilling for natural gas. Potential effects land disturbance associated with activities from natural gas development on aquatic biota in surrounding streams have not yet been well documented. An increase in inorganic sediment in streams can be detrimental to organisms through a variety of mechanisms including alteration of dominant substrate type, higher turbidity resulting in lower visibility, and burial of food resources such as algae and detritus. Increasing sedimentation in stream environments through anthropogenic disturbance is a …
Distribution, Ecology, And Trophic Relationships Of A Colonial Waterbird: The Double-Crested Cormorant, Kate Sheehan
Distribution, Ecology, And Trophic Relationships Of A Colonial Waterbird: The Double-Crested Cormorant, Kate Sheehan
All Dissertations
The realized distribution of an organism is dependent on the environmental characteristics of the landscape, biological interactions within the communities in which it lives, and geographic barriers to dispersal. Changes in habitat can influence landscape characteristics, and consequently, the distribution of organisms within the landscape. The Double-crested Cormorant ( Phalacrocorax auritus ) is a piscivorous waterbird implicated in human-wildlife conflicts at aquaculture facilities and natural aquatic systems where they compete for resources (fish) with anglers and commercial fishing guides. Impacts of P. auritus on aquatic systems result from their consumption of fish stocks and their contamination of water and soil …
Distribution And Abundance Of Anopheles Spp. In The Lower Rio Grande Valley, South Texas, Norma Hermelinda Martinez
Distribution And Abundance Of Anopheles Spp. In The Lower Rio Grande Valley, South Texas, Norma Hermelinda Martinez
Theses and Dissertations - UTB/UTPA
This study investigated the relationship between Anopheles abundance, collection sites and environmental variables in the Lower Rio Grande Valley, South Texas. In addition, species composition in the Lower Rio Grande Valley was determined in 2011. A total of 6772 female mosquitoes were identified to six genera and 27 species. The most prevalent genera collected were Culex (53.9%), Ochlerotatus (25.6%) and Aedes (13.6%). Anopheles mosquitoes were collected using resting boxes during summer 2012 at multiple locations along the Lower Rio Grande Valley. ArcGIS was used to identify land cover characteristics and nearest water sources at mosquito collection sites. Estero Llano Grande …
Conserving The Land Of The Giants: Critical Threats To Asian Elephant Habitat In Sri Lankan Protected Areas, Christie Sampson
Conserving The Land Of The Giants: Critical Threats To Asian Elephant Habitat In Sri Lankan Protected Areas, Christie Sampson
All Theses
Grasslands habitats are hypothesized to be a critical resource for the endangered Asian elephant (Elephas maximus) throughout its range. However, this hypothesis has not been rigorously tested. My study examined elephant habitat selection in Sri Lanka to determine the importance of grassland versus other local habitats, and how livestock abundance, fire, and the invasive plant Lantana camara affect the relative abundance of elephants within habitats. My research was conducted in two protected areas in Sri Lanka, Udawalawe National Park (UNP) and Hurulu Forest Reserve (HR). I used distance-sampling on a total of 50.8 km of dung line-transects in the four …
An Assessment Of Habitat Suitability For Pronghorn Populations Of The Central Valley Region Of California, Virginia Burroughs
An Assessment Of Habitat Suitability For Pronghorn Populations Of The Central Valley Region Of California, Virginia Burroughs
Master's Theses
Efforts to reintroduce and maintain populations of pronghorn (Antilocapra americana) to the California Central Valley, specifically the Carrizo Plain National Monument (CPNM) and the Mojave Desert (Antelope Valley) portion of Tejon Ranch, have largely been unsuccessful due to dwindling numbers of translocated animals. The objective of this study was to improve upon previous models for the CPNM using aerial survey data and then apply the model to the Tejon Ranch. Aerial survey data collected from 2000-2010 on the CPNM was used to establish “use” and “non-use” areas in the model. Model variables included vegetation type (forest, shrub, grassland, …
Population Demographics And Genetic Structure Of Black Bears In Coastal Louisiana, Jesse Charles Troxler
Population Demographics And Genetic Structure Of Black Bears In Coastal Louisiana, Jesse Charles Troxler
Masters Theses
The range and abundance of the Louisiana black bear (Ursus americanus luteolus) were greatly diminished during the 20th century. This subspecies was reduced to 3 small, isolated subpopulations in Louisiana as bottomland hardwood habitat was converted to agriculture. These bears were listed as threatened by the U.S. Fish and Wildlife Service in 1992 and a recovery plan was published in 1995. Recovery requires estimates of population parameters to evaluate current population status and future viability. I conducted a mark-recapture study from 2010 to 2012 to estimate demographic parameters of the coastal population of Louisiana black bears. Because inbreeding …
Time-Dependent Random Effect Poisson Random Field Model For Polymorphism Within And Between Two Related Species, Shilei Zhou
Time-Dependent Random Effect Poisson Random Field Model For Polymorphism Within And Between Two Related Species, Shilei Zhou
UNLV Theses, Dissertations, Professional Papers, and Capstones
Molecular evolution is partially driven by mutation, selection, random genetic drift, or combination of the three factors. To quantify the magnitude of these genetic forces, a previously developed time-dependent fixed effect Poisson random field model provides powerful likelihood and Bayesian estimates of mutation rate, selection coefficient, and species divergence time. The assumption of the fixed effect model that selection intensity is constant within a genetic locus but varies across genes is obviously biologically unrealistic, but it serves the original purpose of making statistical inference about selection and divergence between two related species they are individually at mutation-selection-drift inequilibrium. By relaxing …
Ichnology And Paleoecology Of The Jurassic Aztec Sandstone, Heather Marie Stoller
Ichnology And Paleoecology Of The Jurassic Aztec Sandstone, Heather Marie Stoller
UNLV Theses, Dissertations, Professional Papers, and Capstones
In this study I describe and interpret tracks and trackways of the Jurassic Aztec Sandstone of southern Nevada and southern California. This study involved mapping of all known tracks and trackways, including foot length, stride length, and trackway width. Photogrammetric data, collected by Bureau of Land Management scientists, were utilized for several trackways in Red Rock Canyon National Conservation Area.
More than three hundred tracks belonging to five ichnotaxa were documented within the Aztec Sandstone, including about 165 tridactylGrallator, 250 tetradactyl (four-toed)Brasilichniumtracks, and 7 arthropod trackways ofOctopodichnusandPaleohelcura. Four of the five ichnotaxa were not previously reported from the Aztec Sandstone. …
Determining Parentage And The Effects Of Relatedness On Play Partner Preference In Belding's Ground Squirrels, Jessica Weidenbach
Determining Parentage And The Effects Of Relatedness On Play Partner Preference In Belding's Ground Squirrels, Jessica Weidenbach
Master's Theses
I investigated 1) multiple paternity in Belding’s ground squirrels (Urocitellus beldingi) and 2) social partner preferences in juvenile U. beldingi. Prior work with blood allozymes demonstrated multiple paternity in U. beldingi litters. I evaluated paternity using microsatellite DNA analysis, which is more accurate than examining blood allozymes. My results indicate that multiple paternity in U. beldingi is more extensive than previously shown, occurring in about 90% of litters with more than one juvenile, and averaging 2.95 fathers in multiply sired litters. I also evaluated the hypothesis that play and other social behavior promotes bonding among juvenile female …
Body Size Variation In Two Adjacent Populations Of Black Kingsnakes (Lampropeltis Nigra) In East Tennessee, Jesse Weber, Joshua Ennen
Body Size Variation In Two Adjacent Populations Of Black Kingsnakes (Lampropeltis Nigra) In East Tennessee, Jesse Weber, Joshua Ennen
Chancellor’s Honors Program Projects
No abstract provided.
Reproductive Biology And Speciation In Drosophila Pseudoobscura, Josh H.M. Alpern
Reproductive Biology And Speciation In Drosophila Pseudoobscura, Josh H.M. Alpern
Electronic Thesis and Dissertation Repository
Sexual selection is an important force driving the evolution of reproductive traits, including sperm morphology and mating behaviour. Divergent sexual selection among populations can eventually lead to errors in spermatogenesis in inter-population hybrids, and subsequently speciation. In Chapter 2, I identify a novel sperm class and how its proportion in the ejaculate is adjusted when Drosophila pseudoobscura males are exposed to competition. In Chapter 3, I assess how competition causes both males and females to adjust their mating behaviour. In Chapter 4, I characterize interspecific hybrid spermatogenic breakdown from two closely-related sub-species. While the genetics of hybrid sterility has been …
Iron-Regulated Cyanobacterial Predominance And Siderophore Production In Oligotrophic Freshwater Lakes, Ryan J. Sorichetti
Iron-Regulated Cyanobacterial Predominance And Siderophore Production In Oligotrophic Freshwater Lakes, Ryan J. Sorichetti
Electronic Thesis and Dissertation Repository
The frequency and intensity of cyanobacterial blooms (cyanoblooms) is increasing globally. Contrary to existing phosphorus (P) and nitrogen (N) paradigms describing cyanobloom proliferation in eutrophic (nutrient-rich) freshwater lakes, many of the recent cyanobloom reports pertain to oligotrophic (nutrient-poor) freshwater lakes with no prior history of cyanobloom occurrence. There exists a critical research need to re-visit existing conceptual models, identify regulating factors currently unaccounted for and improve our ability to effectively detect and measure cyanobacterial toxins (cyanotoxins) in lakes. Iron (Fe) is required in nearly all pathways of cyanobacterial macronutrient use, though its direct role in regulating cyanobacterial biomass is not …
Soil Freezing Effects On A Grass-Dominated Old Field Ecosystem Under Current And Future Rates Of Atmospheric Nitrogen Deposition, Mat Vankoughnett
Soil Freezing Effects On A Grass-Dominated Old Field Ecosystem Under Current And Future Rates Of Atmospheric Nitrogen Deposition, Mat Vankoughnett
Electronic Thesis and Dissertation Repository
Climate change is expected to alter the intensity and dynamics of soil freezing as a result of increased air temperatures and reduced snow cover. Soil freezing can influence ecosystem nitrogen (N) cycling by damaging plants and soil microorganisms, but little is known about how soil freezing effects on ecosystem N cycling may combine or interact with increased atmospheric N deposition, which is also expected to exert a strong influence on terrestrial ecosystems in the coming decades. The objective of my thesis was to examine the combined and possibly the interactive effects of climate induced changes in soil freezing and N …
The Role Of Morphology In Diet And Flower Visitation By Five Species Of Cuban Flower-Visiting Bats, Lindsey Clairmont
The Role Of Morphology In Diet And Flower Visitation By Five Species Of Cuban Flower-Visiting Bats, Lindsey Clairmont
Electronic Thesis and Dissertation Repository
Pollinator and flower morphology are important factors in structuring the plant-pollinator relationship. A pollinator’s morphology may influence aspects of its diet. Flower-visiting bats are an important pollinator group but very little is known about what influences their interactions with food plants. I examined the role of morphology in the partitioning of food resources for five species of flower-visiting Cuban bats. I analyzed cranial traits and body size to examine differences among species and to determine the degree of morphological specialization for flower-feeding for each species. I also collected dietary data from guano and used acoustic monitoring to assess bat activity …
The Effects Of Anthropogenic Noise On The Foraging Efficiency Of A Gleaning Bat And The Activity Levels Of A Natural Bat Assemblage, Jessie Patrice Bunkley
The Effects Of Anthropogenic Noise On The Foraging Efficiency Of A Gleaning Bat And The Activity Levels Of A Natural Bat Assemblage, Jessie Patrice Bunkley
Boise State University Theses and Dissertations
Anthropogenic noise is prevalent across many landscapes, posing a threat of disturbance to countless human and wildlife populations. Studies have revealed that a variety of organisms are negatively affected by an increasingly loud soundscape, including auditory predators such as bats. Bats that exhibit a gleaning hunting strategy passively listen for low frequency, prey-produced sounds. Anthropogenic noise often falls within the same spectral range as important prey cues, potentially masking these signals. I investigated the effects of two sources of anthropogenic noise, traffic and gas compressor station noise, on the foraging efficiency of the North American gleaning bat Antrozous pallidus in …
Thermal Adaptation Of Life History Traits In The Drosophila Melanogaster Group, Christopher James Austin
Thermal Adaptation Of Life History Traits In The Drosophila Melanogaster Group, Christopher James Austin
Electronic Thesis and Dissertation Repository
Thermal adaptation is typically detected by examining the tolerance to extreme temperatures in a few populations within a single life stage. However, the extent to which adaptation occurs among many different populations might depend on the tolerance of multiple life stages and the average temperature range that the population experiences. Here, I examined adaptation to local temperature conditions in four species of fruit flies, including a cosmopolitan species, Drosophila melanogaster, and three species with geographically small-sized ranges, D. nepalensis, D. sechellia, and D. mauritiana. The cosmopolitan species showed adaptation to native temperatures during the larval and adult …
The Role Of Resource Predictability In The Metapopulation Dynamics Of Insects, Byju Nambidiyattil Govindan
The Role Of Resource Predictability In The Metapopulation Dynamics Of Insects, Byju Nambidiyattil Govindan
Open Access Dissertations
The metapopulation paradigm has emerged as an important tool to understand the dynamics of species living in fragmented landscapes. In this dissertation, I investigate the unpredictable nature of resource availability for species living in human-dominated heterogeneous and dynamic landscapes in the context of its consequences for long-term regional persistence of species. In particular, I test theoretical advancements in metapopulation ecology following a two-pronged approach - via experiments in the lab and observations in the field - using insects. In chapter 1, I introduce the concept of metapopulation ecology in the context of its relevance for dynamics of species living in …
Long-Term Disturbance Histories Of Managed And Natural Mixed-Hardwood Forests Of Central Indiana, Robert Charles Morrissey
Long-Term Disturbance Histories Of Managed And Natural Mixed-Hardwood Forests Of Central Indiana, Robert Charles Morrissey
Open Access Dissertations
Over the course of the last century, we have observed changes in forest composition and structure related to changes in disturbance regimes. Reduced frequency and severity of disturbance events has allowed for the ingrowth of shade-tolerant, mesophytic species into forest understories; the result is shifting species compositions and changes in forest structure. In some cases, the loss of masting species, such as oak (Quercus spp.) and hickory (Carya spp.), may also accompany a shift in forest function and further accommodate other species shifts among mammal, insect, and bird populations. Management efforts have been suggested as a possible means …