Ecology and Evolutionary Biology Commons^™

Reevaluation Of The Described Subspecies Of Euphydryas Phaeton (Drury, 1773) With A Replacement Name For Melitaea Phaeton Schausi (Clark, 1927), Harry Pavulaan

The Taxonomic Report of the International Lepidoptera Survey

The present paper reevaluates the subspecific standing of the nominotypical and three described subspecies of Euphydryas phaeton (Drury, 1773). The nominotypical subspecies phaeton occupies the mid-Atlantic region, with undefined zones of contact with described subspecies borealis (F. Chermock & R. Chermock, 1940) to the north, and subspecies schausi (Clark, 1927) to the south. Nominotypical phaeton is an intermediate phenotype between borealis and schausi, which are each noticeably different from each other but both reasonably similar to intermediate (nominotypical) phaeton. Both borealis and schausi were synonymized under nominotypical phaeton since about time of their descriptions, by authors and list …

New Synonymies And Notes In Criodion Audinet-Serville (Coleoptera: Cerambycidae: Cerambycinae), Antonio Santos-Silva, Sergio Devesa

Insecta Mundi

Criodion angustatum Buquet, 1852 and C. pilosum Lucas, 1857 (Coleoptera: Cerambycidae: Cerambycinae) are synonymized with C. tomentosum Audinet-Serville, 1834. The holotypes of C. angustatum and C. hirsutum, and syntypes of C. pilosum and of C. tuberculatum Gahan, 1892 are illustrated for the first time. One of the type localities of C. tuberculatum is corrected.

Currently, Criodion Audinet-Serville, 1834 includes 13 species distributed from Guatemala to southern South America (Monné 2021; Tavakilian and Chevillotte 2021). Despite the relatively small number of species, they encompass several problems with their identification. For example, the specimens often identified as C. murinum Nonfried, 1895 …

Ishtarella Thailandica Martens, New Genus, New Species (Hymenoptera: Braconidae: Aphidiinae) Of Aphid Parasitoid From Thailand, With A Country Checklist Of Aphidiinae, Abigail P. Martens, Matthew L. Buffington, Donald L.J. Quicke, Marisa Raweearamwong, Buntika A. Butcher, Paul J. Johnson

Insecta Mundi

Ishtarella Martens new genus (Hymenoptera: Braconidae: Aphidiinae) and I. thailandica Martens new species are described and illustrated from Doi Phu Kha National Park, Nan Province, Thailand. The genus is assigned to the tribe Aphidiini, subtribe Trioxina. Based on morphology, Ishtarella appears most closely related to Binodoxys Mackauer. An updated checklist of the aphidiine fauna of Thailand, based on published records, is presented.

Parasitoid wasps in the subfamily Aphidiinae (Hymenoptera: Braconidae) comprise about 60 genera and more than 650 species worldwide (Yu et al. 2016). The Aphidiinae of Thailand were all but unknown prior to a Thai­land Inventory Group for Entomological …

Ardella Magnaemirabilis (Coleoptera: Scarabaeidae: Melolonthinae: Ardellini), A New Scarabaeoid Species, Genus And Tribe From The Southwestern United States, M. J. Paulsen

Insecta Mundi

An interesting new taxon of scarabaeoid beetle is described from western Texas, USA. The remark­ably small chafer, Ardella magnaemirabilis new genus, new species (Coleoptera: Scarabaeidae: Melolonthinae: Ardellini) is described and compared to other diminutive scarabaeoid genera from drier habitats of the southwestern USA. The tribe Ardellini, new tribe, is erected to accommodate the unusual genus.

After a few centuries with entomologists scouring the continent, the discovery of unknown Nearctic scarab gen­era is no longer commonplace. The western United States, in particular, is biogeographically diverse with pockets of endemism that makes such discoveries still possible, driving collectors to continue sampling …

Two New Species Of Deromecus Solier From Guatemala (Coleoptera: Elateridae: Pomachiliini), E. R. Fuller

Insecta Mundi

Deromecus georginas Fuller, new species, and Deromecus longitarsis Fuller, new species (Cole­optera: Elateridae: Pomachiliini) are described from cloud forest habitat on eastern coastal volcanoes in Quetzaltenango Department, Guatemala, and compared with Deromecus trivittatus Champion from Mexico.

Se describen la neuva especie Deromecus georginas Fuller y la neuva especie Deromecus longitarsis Fuller (Coleoptera: Elateridae: Pomachiliini), del hábitat del bosque nuboso, de los volcanes oriental de la costa, en el Departmento de Quetzaltenango, Guatemala, y se comparan con Deromecus trivittatus Cham­pion, de México.

The Elateridae are a conspicuous and diverse group of beetles in Central America (Champion 1894–1897). Adults of luminous …

Occurrence Of An Exotic Whitefly, Siphoninus Phillyreae (Haliday) (Hemiptera: Aleyrodidae) In South Korea And Its Potential Pathway Analysis, Seon Woo Lee, Hyun Jik Lee, Jun-Ran Kim, Soo-Jung Suh

Insecta Mundi

The ash whitefly, Siphoninus phillyreae (Haliday) (Hemiptera: Aleyrodidae), was first found in southern areas of South Korea during September 2019. During the survey of 2021, no additional specimens of the ash whitefly were found other than the previously reported regions. Attempts to trace a possible path­way for exotic ash whitefly suggested that this species probably followed pathways of illegal importation of infested plants from China or Japan and was unintentionally introduced into South Korea based on a survey-based study. This whitefly has not been intercepted at ports of entry to South Korea on imported plant material between 1999 and 2019 …

Nesting Behavior, Ecology, And Functional Morphology Of The Trapdoor Spider-Hunting Spider Wasp Aporus (Plectraporus) Hirsutus (Banks) (Hymenoptera: Pompilidae), Frank E. Kurczewski, Alice J. Abela, Rick C. West

Insecta Mundi

Macrophotographs in series taken by Alice Abela on sandy coastal dunes in Santa Barbara and San Luis Obispo Counties, CA in 2010–2021 supplement and enhance F. X. Williams (1928) study of the ecol­ogy and nesting behavior of the trapdoor spider-hunting spider wasp Aporus (Plectraporus) hirsutus (Banks) (Hymenoptera: Pompilidae: Aporini). Abela’s macrophotographs and observations provide new details of adult wasp feeding, functional morphology, hunting, digging and prey transport, and host spider trapdoor, entrance, burrow structure, host capture and escape activity. Newly reported host records from this study and online photographs expand A. hirsutus host selection in the large wafer-lid …

New Larval Host Plants And Ecological Observations On North American Cerambycidae (Coleoptera), Josef Vlasak, Katerina Vlasakova

Insecta Mundi

New larval hosts are presented for 152 North American Cerambycidae (Coleoptera). Larval host plants of Aneflus calvatus Horn in Leng, Aneflomorpha arizonica Linsley, Aneflomorpha fisheri Linsley, Ane­flomorpha opacicornis Linsley, Anelaphus magnipunctatus (Knull), Atylostagma glabra Schaeffer, Cacophrissus pauper Bates, Elytroleptus limpianus Skiles and Chemsak, Obrium rubidum LeConte, and Stenelytrana splen­dens (Knull) are reported for the first time. The types of wood utilized by larvae of Leptorhabdium pictum (Haldeman), Sphenostethus taslei (Buquet), Typocerus lugubris (Say), and Xestoleptura octonotata (Say) are discussed. Notes on larval morphology and larval habits of Aneflus spp. and Stenaspis solitaria (Say), includ­ing a correction of a published host …

Micromentignatha Geberti, A New Tiger Beetle Species From Australia (Coleoptera: Cicindelidae), Jürgen Wiesner

Insecta Mundi

Micromentignatha geberti (Coleoptera: Cicindelidae), new species, is described. It is distinguished from the other members of the genus by its larger body size and shape of elytral maculation.

Due to the kindness of Christian Kerkering, I was able to study specimens of a tiger beetle species he recently acquired. It proved to be new to science and is described herein. It is a member of Micromentignatha Sumlin, 1981, which raises the number of species in this genus to four (Wiesner 2020: 327). A key to distinguish these congeners as well as a catalog of the genus are given herein.

S5e11: Why Might Maine Lose Two Species Of Songbirds?, Ron Lisnet, Katherine Ruskin, Brian Olsen

The Maine Question

Maine may lose two tidal marsh songbird species in the next few decades. Saltmarsh sparrows face extinction, while Acadian Nelson’s sparrows are threatened with extirpation — localized eradication with the possibility of survival elsewhere. Their populations along the Eastern Seaboard have been declining as sea level rise destroys their habitats and, according to a new University of Maine-led study, mercury exposure inhibits their reproduction.

In the final episode of Season 5 of “The Maine Question,” Kate Ruskin, a lecturer in ecology and environmental science at UMaine who spearheaded the recent mercury exposure study, and Brian Olsen, who is now an …

Uncoupled Seasonal Variability Of Transparent Exopolymer And Coomassie Stainable Particles In Coastal Mediterranean Waters: Insights Into Sources And Driving Mechanisms, Marina Zamanillo, Eva Ortega-Retuerta, Carolina Cisternas-Novoa, Cèlia Marrasé, Carles Pelejero, Josep Pascual, Josep M. Gasol, Anja Engel, Rafel Simó

Research outputs 2014 to 2021

Transparent exopolymer particles (TEP) and Coomassie stainable particles (CSP) are gel-like particles, ubiquitous in the ocean, that affect important biogeochemical processes including organic carbon cycling by planktonic food webs. Despite much research on both groups of particles (especially TEP) over many years, whether they exist as distinctly stainable fractions of the same particles or as independent particles, each with different driving factors, remains unclear. To address this question, we examined the temporal dynamics of TEP and CSP over 2 complete seasonal cycles at 2 coastal sites in the Northwestern Mediterranean Sea, the Blanes Bay Microbial Observatory (BBMO) and the L’Estartit …

Soil Microbial Community Dynamics In Response To Prescribed Extreme Fires Following Juniperus Virginiana Invasion In The Loess Canyons Of Nebraska, Julie A. Fowler

School of Natural Resources: Dissertations, Theses, and Student Research

In Nebraska and other regions of the Great Plains, the conifer Juniperus virginiana (eastern redcedar) is converting grasslands to dense woodlands. This is driven by the interacting drivers of fire suppression, altered grazing regimes, climate change and other anthropogenic factors, impacting the provisioning of ecosystem services. This vegetation state transition modifies water resource regulation and biogeochemical cycles leading to altered edaphic properties including soil microbial community composition. To restore these grasslands and control J. virginiana spread, prescribed extreme burns are implemented as a management tool through local prescribed burn associations. We hypothesized that the alternative state transition to dense J. …

Northern Saw-Whet Owl Autumn Migration In Eastern Nebraska: Results From A Three-Year Banding Study, Stephen J. Brenner, Joel G. Jorgensen

Nebraska Bird Review

Prior to 2019, records of NSWOs in Nebraska during fall were limited. There were fewer than five accepted reports in eastern Nebraska since the 1950s (Silcock and Jorgensen, 2021). The Hitchcock banding station along the Missouri River bluffs in nearby western Iowa has captured, banded, and released 20-50 birds during autumn migration for a number of years (J. Toll, personal communication). Furthermore, a single season banding study in central Nebraska during fall 2004 captured 14 NSWOs (Kim 2005), suggesting the species may migrate annually over much of the state in fall, or at least in more years than previously thought. …

The Nebraska Bird Review, Volume 89 Number 4, December 2021

Nebraska Bird Review

Fall Field Report, August - November 2021 by W. Ross Silcock

Book Review: Nest Records of Nebraska Birds by Wayne J. Mollhoff Reviewed by W. Ross Silcock

Northern Saw-whet Owl Autumn Migration in Eastern Nebraska: Results from a Three-year Banding Study by Stephen J. Brenner and Joel G. Jorgensen

Mute Swan Observation at Gavins Point Dam Area, Cedar/Knox Co., Nebraska by Caleb Strand and Mark A. Brogie

Book Review: The Sandhill Crane State: A Naturalist's Guide to Nebraska by Paul Johnsgard, Reviewed by Ruth Stearns

Index to Volume 89

Subscription and Organization Information

Mute Swan Observation At Gavins Point Dam Area, Cedar/Knox Co., Nebraska, Caleb Strand, Mark A. Brogie

Nebraska Bird Review

The Nebraska Ornithologists’ Union Records Committee (NOURC) considers Mute Swan (Cygnus olor) to be Accidental in Nebraska (NOURC 2017) with only two accepted records. A review of Mute Swan status in the Midwest and Nebraska, how individual records in Nebraska are evaluated to determine wild provenance, and a review of Nebraska reports can be found in “The Mute Swan in Nebraska: History and Current Status” (Brogie et al. 2016).

The following is an account of the third NOURC accepted record for Mute Swan in Nebraska. On 11 June 2021 we observed two sub-adult Mute Swans on Lake Yankton, Cedar County, …

Review Of Nest Records Of Nebraska Birds By Wayne J. Mollhoff, W. Ross Silcock

Nebraska Bird Review

This book summarizes a lifetime of investigation of Nebraska’s breeding birds by Wayne Mollhoff. Even though I say the book summarizes a lifetime of investigation, I would not be surprised if Wayne resurfaces in a few years with a follow-up of some sort, intrepid investigator that he is. Over the years Wayne has spent innumerable days in the field climbing trees in full forestry gear, wading and kayaking marshes and rivers, hiking over prairies, and bushwhacking through dense forests (there are still a few in Nebraska). But this isn’t the totality of Wayne’s talents; he is a determined researcher with …

Review Of The Sandhill Crane State: A Naturalist’S Guide To Nebraska By Paul Johnsgard, Ruth Stearns

Nebraska Bird Review

Dr. Paul Austin Johnsgard left us one last gift, in this reference guide to all places natural in Nebraska. Published posthumously, and available for purchase or free via UNL’s Digital Commons, this book arrived at my doorstep like a veritable “seed catalog” of Nebraska’s places of wonder to delight and inspire fair weather adventures just before the long winter set in. He wants us to explore the state as he had done, to be excited by all the out-ofdoors Nebraska has to offer.

Index To Volume 89

Nebraska Bird Review

Adams, Marie 52

to

Zonotrichia leucophrys gambelii 168 leucophrys 75, 168-169 oriantha 75, 169

Fall Field Report, August-November 2021, W. Ross Silcock

Nebraska Bird Review

INTRODUCTION Recent years have seen increasing reports of birds lingering quite late, and this fall was no exception. Notable groups exhibiting tardy departures were shorebirds and vireos. Tardy shorebirds were record late Solitary Sandpipers, near-record late Dunlin, and late Long-billed Curlew, Ruddy Turnstone, Least and Semipalmated Sandpipers, Long-billed Dowitcher, and Wilson’s Phalarope. Among the vireos, all regular species were at least tardy; record late was a Red-eyed Vireo, a Warbling Vireo was record late for the Panhandle, and a Blue-headed Vireo was second latest on record. Record late was a Barn Swallow, and a Mountain Bluebird was record late in …

Expanding The Conservation Genomics Toolbox: Incorporating Structural Variants To Enhance Genomic Studies For Species Of Conservation Concern, Stephanie J. Galla

Biology Faculty Publications and Presentations

Structural variants (SVs) are large rearrangements (>50 bp) within the genome that impact gene function and the content and structure of chromosomes. As a result, SVs are a significant source of functional genomic variation, that is, variation at genomic regions underpinning phenotype differences, that can have large effects on individual and population fitness. While there are increasing opportunities to investigate functional genomic variation in threatened species via single nucleotide polymorphism (SNP) data sets, SVs remain understudied despite their potential influence on fitness traits of conservation interest. In this future-focused Opinion, we contend that characterizing SVs offers the conservation genomics …

Primnoa Pacifica Reproduction In Shallow Versus Deep Habitats Of Glacier Bay National Park And Preserve, Alaska, Ciara N. Larence

Honors College

Primnoa pacifica is a species of deep-sea cold-water coral that can be found in Glacier Bay National Park and Preserve in Alaska. These colonies are important to their ecosystem as they provide habitat for other species living in this area. One thing that makes P. pacifica important to study is that the species displays deep-water emergence. This is a phenomenon where species normally found in deep waters can exist in shallower waters, allowing easier access for research. The purpose of this thesis was to determine if depth effects the reproduction of male P. pacifica colonies. Two colonies from deep depths …

The Use Of Singlebeam Echo-Sounder Depth Data To Produce Demersal Fish Distribution Models That Are Comparable To Models Produced Using Multibeam Echo-Sounder Depth, Marcela Montserrat Landero Figueroa, Miles J. G. Parsons, Benjamin J. Saunders, Ben Radford, Chandra Salgado-Kent, Iain M. Parnum

Research outputs 2014 to 2021

Seafloor characteristics can help in the prediction of fish distribution, which is required for fisheries and conservation management. Despite this, only 5%–10% of the world's seafloor has been mapped at high resolution, as it is a time-consuming and expensive process. Multibeam echo-sounders (MBES) can produce high-resolution bathymetry and a broad swath coverage of the seafloor, but require greater financial and technical resources for operation and data analysis than singlebeam echo-sounders (SBES). In contrast, SBES provide comparatively limited spatial coverage, as only a single measurement is made from directly under the vessel. Thus, producing a continuous map requires interpolation to fill …

American Bullfrog (Lithobates Catesbeianus) Diet In Uruguay Compared With Other Invasive Populations In Southern South America, Gabriel Laufer, Noelia Gobel, Mauro Berazategui, Matías Zarucki, Sofía Cortizas, Alvaro Soutullo, Claudio Martinez Debat, Rafael O. De Sá

Biology Faculty Publications

Between 2000-2020, more than ten new populations of the invasive American bullfrog (Lithobates catesbeianus) have been reported in the southern cone of South America. We studied the stomach contents of 126 bullfrogs from a population at an early invasion stage in Uruguay (Acegua, Cerro Largo Department). We observed a rich diet, with extensive prey volume range (1 mm3 to more than 7 000 mm(3)); the most frequent items were Hymenoptera (19.6%), Coleoptera (16.4%), Amphipoda (13.3%), Anura (8.9%) and Heteroptera (8.7%). Despite some overlap, differences were observed in volume (chi(2) = 54.6, p <0.001, d.f. = 2) and prey quantity (F = 8.1, p <0.001, d.f. = 79) between males, females, and juveniles. Juveniles showed significantly higher consumption of terrestrial prey by count (82% of their total ingestion) than adults (29% for males and 32% for females) (chi(2) = 28.5, p <0.001, d.f. = 2). Adults, especially females, showed a high frequency of cannibalism (33% of their total ingestion; chi(2) = 20.9, p <0.001, d.f. = 2). Comparing our data with other bullfrog regional studies, we found great plasticity in trophic habits and differences in the incidence of cannibalism (higher incidence in the populations of Acegua, Uruguay, and Buenos Aires, Argentina). These differences could be related to local biodiversity, but also could be affected by the invasion phase. Cannibalism frequency was higher in small bullfrog populations, where it could be favoring the establishment success. This shift in foraging strategies during the invasion process had been insufficiently evaluated in amphibians. Knowing the ecological determinants for the invasion by bullfrogs can be useful to the development of management strategies.

American Burying Beetle, Plant Richness, And Soil Property Responses To Collapse Of Juniperus Virginiana Woodlands With Fire, Alison Ludwig

Department of Agronomy and Horticulture: Dissertations, Theses, and Student Research

Grasslands are declining in the Great Plains due to land use changes, woody plant encroachment, and loss of historic fire cycles. Prescribed burn associations have utilized prescribed fire to collapse invading woodlands and allow the restoration of grasslands. This fire is considered “extreme” because it is capable of changing the structure and function of an ecosystem. Our study site is the Loess Canyons Experimental Landscape, a long-term, ecoregion-scale experiment to apply prescribed fire across the region to restore grasslands. The Nebraska Natural Legacy Project established the Loess Canyons ecoregion as a Biologically-Unique Landscape in 2005 with the state’s wildlife action …

Effects Of Glacial Stressors On Sperm Maturation In Colonies Of The Red Tree Coral, Primnoa Pacifica, Joshua Lynn

Honors College

The red tree coral, Primnoa pacifica, is a large, colony forming species of cold- water coral which is often an important habitat for many commercially important species of fish and crab. This keystone species is long lived and found at much shallower depths in the fjords of Glacier Bay National Park (GBNP) than elsewhere in the northern Pacific Ocean because of the phenomenon known as deep-water emergence. Due to their proximity to tidewater glaciers in GBNP, corals likely have to endure glacial stressors such as freshwater runoff and sedimentation that is not typical of populations in deeper water, which …

Deterring Non-Target Birds From Toxic Bait Sites For Wild Pigs, Nathan P. Snow, Joseph M. Halseth, Justin A. Foster, Michael J. Lavelle, Justin W. Fischer, Michael P. Glow, Ingrid A. Messer, Seth M. Cook, Kurt C. Vercauteren

United States Department of Agriculture Wildlife Services: Staff Publications

Toxic baiting of wild pigs (Sus scrofa) is a potential new tool for population control and damage reduction in the US. Field trials testing a prototype toxic bait (HOGGONE 2 containing 5% sodium nitrite [SN]), though, revealed that wild pigs spilled small particles of toxic bait outside of bait stations which subsequently created hazards for non-target species that consumed those particles, primarily passerine birds. To deter non-target birds from consuming particles of spilled bait, we tested four deterrents at mock bait sites (i.e., baited with bird seed) in north-central Colorado, USA during April–May 2020. We found a programable, inflatable deterrent …

Towards A More Comprehensive Understanding Of Wild Pig (Sus Scrofa) Impacts On Agricultural Producers: Insights From A Texas Case Study, Keith M. Carlisle, Nicole Didero, Sophie Mckee, Julie Elser, Stephanie A. Shwiff

United States Department of Agriculture Wildlife Services: Staff Publications

This research investigates the impacts of invasive wild pigs (Sus scrofa Linneaus) on agricultural producers in Texas, with the aim of identifying and describing all categories of wild pig impacts and quantifying the extent of producers’ over- or underestimation of their total wild pig-related costs in 2018, as compared to calculations based upon data subsequently provided by the producers about individual wild pig-related costs and losses. Based on interviews with 23 producers in 16 Texas counties, we identified more than 20 discrete categories of negative impacts and negligible positive impacts associated with wild pigs. Among them were categories that have …

Landscape Transformations Produce Favorable Roosting Conditions For Turkey Vultures And Black Vultures, Jacob E. Hill, Kenneth F. Kellner, Bryan M. Kluever, Michael L. Avery, John S. Humphrey, Eric A. Tillman, Travis L. Devault, Jerrold L. Belant

United States Department of Agriculture Wildlife Services: Staff Publications

Recent increases in turkey vulture (Cathartes aura) and black vulture (Coragyps atratus) populations in North America have been attributed in part to their success adapting to human-modified landscapes. However, the capacity for such landscapes to generate favorable roosting conditions for these species has not been thoroughly investigated. We assessed the role of anthropogenic and natural landscape elements on roosting habitat selection of 11 black and 7 turkey vultures in coastal South Carolina, USA using a GPS satellite transmitter dataset derived from previous research. Our dataset spanned 2006–2012 and contained data from 7916 nights of roosting. Landscape …

Reproductive Success Of Captive-Reared Allegheny Woodrats (Neotoma Magister) Released Into Genetically Depauperate Populations, Miracle M. Davis, Timothy J. Smyser, Scott A. Johnson, Joseph Duchamp, Jeffery L. Larkin, Robert K. Swihart, Jacqueline M. Doyle

United States Department of Agriculture Wildlife Services: Staff Publications

Habitat fragmentation and degradation have led to a proliferation of small and isolated populations that are vulnerable to genetic erosion. Reduction in habitat and concomitant declines in population connectivity can expediate the collapse of species that exist as natural metapopulations. In recent years, Allegheny woodrats (Neotoma magister) have experienced local extirpations and declines in genetic diversity across their range due to disease-related mortality, reduced food availability, and disruption of connectivity among subpopulations. In response, we developed a captive-breeding program to facilitate genetic management of isolated woodrat populations in Indiana. Between 2010 and 2012, 27 captive-reared individuals were released to four …

Economic Estimates Of Wild Hog (Sus Scrofa) Damage And Control Among Young Forest Plantations In Alabama, Micah Fern, Rebecca Barlow, Chris Slootmaker, John Kush, Stephanie Shwiff, Larry Teeter, Jim Armstrong

United States Department of Agriculture Wildlife Services: Staff Publications

Operating as ecological engineers, the increased distribution and abundance of wild hogs (Sus scrofa) has caused considerable socio-economic impacts. The international scope of economic research providing wild hog damage estimates are often confined to agricultural crops, while damage estimates among forest plantations are lacking. In Alabama, private landowners hold the majority of timberland acreage and are less equipped to absorb financial losses from wild hog damage than their industrial counterparts. A survey was conducted to estimate the economic impact of wild hogs, namely costs of damage and control, to privately owned forestlands. The survey was distributed in the …