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Full-Text Articles in Ecology and Evolutionary Biology

Mammalian & Avian Community Response To African Elephant (Loxodonta Africana) Habitat Modification In Southeastern Kenya, Dakota Vaccaro Aug 2023

Mammalian & Avian Community Response To African Elephant (Loxodonta Africana) Habitat Modification In Southeastern Kenya, Dakota Vaccaro

Masters Theses & Specialist Projects

African elephants (Loxodonta africana) play a significant role in the modification of their habitat, foremost by decreasing woody vegetation cover and density and in some parts of Africa high elephant densities damage small sanctuaries and preserves. For wildlife that depend on forested patches, this increased modification could lead to displacement and/or decreases in abundance. If suitable habitat is not available elsewhere, elephants and other wildlife may move closer to humans in search of resources, which can result in increased human-wildlife conflict. Evaluating this impact was one of the goals of the Elephants and Sustainable Agriculture in Kenya (ESAK) …


The Systems Measurement Of Mammalian Biotas, Part Two, Charles H. Smith, Patrick Georges, Ngoc Nguyen Jan 2023

The Systems Measurement Of Mammalian Biotas, Part Two, Charles H. Smith, Patrick Georges, Ngoc Nguyen

Faculty/Staff Personal Papers

For a recent publication, the authors identified a seven-region model of mammal family distribution patterns, in which each unit contributes equally to the system’s overall statistical characteristics of diversity, despite its individual units having measurably different levels of diversity and endemism. This systemization presents a highly efficient descriptive model that can possibly be interpreted as a form of natural classification. An additional analysis of the same mode is described here, in which the seven-region model of the distribution of mammal families’ spatial affinities is shown to closely approach a most-probable-state arrangement, as assessed through combinatorics, raising some important questions about …


Bibliography, Bruce A. Schulte Jan 2023

Bibliography, Bruce A. Schulte

Faculty/Staff Personal Papers

Bibliography of publications by Bruce Schulte.


Alleviating Human-Elephant Conflict Through Deterrent Fences And Environmental Monitoring In Southern Kenya, Sophia Carmen Corde Apr 2022

Alleviating Human-Elephant Conflict Through Deterrent Fences And Environmental Monitoring In Southern Kenya, Sophia Carmen Corde

Masters Theses & Specialist Projects

Human-wildlife conflict is present across the world. In areas where human settlements overlap with elephant habitats, human-elephant conflict can result from crop raiding events, compromising farmers’ food and economic security, and putting humans and elephants in danger through farmer retaliation. Elephants raid crops primarily at night, when detection by humans is lowest, and during the dry season, as crops are developing towards harvest and natural forage quality drops. People living in these areas facing HEC have developed mitigation strategies to lessen the impacts and move towards coexistence. As a team member on the Elephants and Sustainable Agriculture in Kenya project, …


Changes In The Zooplankton Community In Barren River Lake (South Central Ky) Between 2008 And 2020, Laurel Philpott Jan 2022

Changes In The Zooplankton Community In Barren River Lake (South Central Ky) Between 2008 And 2020, Laurel Philpott

Mahurin Honors College Capstone Experience/Thesis Projects

Zooplankton are small aquatic animals that serve an important role in transferring energy from phytoplankton to higher trophic levels. The zooplankton community composition in Barren River Lake from 2008 and 2020 was compared. A field study was conducted to not only describe the seasonal population dynamics of zooplankton in Barren River Lake, but also to determine if an invasive zooplankton, Daphnia lumholtzi, was present. While D. lumholtzi was found throughout Barren River Lake in 2008, it was absent from both resampled sites in 2019 and 2020. This, along with erratic patterns of emergence in several other zooplankton species, indicates …


Effects Of Melatonin Implantation On The Activity Levels Of Captive Snow Buntings (Plectrophenax Nivalis), Zoë Ward Jan 2022

Effects Of Melatonin Implantation On The Activity Levels Of Captive Snow Buntings (Plectrophenax Nivalis), Zoë Ward

Mahurin Honors College Capstone Experience/Thesis Projects

In vertebrates, melatonin regulates many components of the circadian rhythm; depending on the organism’s location and time of year, the pineal gland will produce melatonin accordingly, controlling the sleep-wake cycle and breeding physiologies. However, locations at the poles in which the sun is above the horizon for multiple days at a time pose unique challenges for the organisms living there. The goal of this study was to examine how the implantation of melatonin in an arctic-breeding songbird affects its activity levels and circadian rhythm. The subject of this study, the snow bunting (Plectrophenax nivalis), is a small songbird …


Morphological Correlates Of Auditory Sensitivity In The Inner Ear Of Two Species Of Invasive Carp, Tyler Patty Jan 2020

Morphological Correlates Of Auditory Sensitivity In The Inner Ear Of Two Species Of Invasive Carp, Tyler Patty

Mahurin Honors College Capstone Experience/Thesis Projects

Silver (Hypopthalmichthys molitrix) and bighead (H. nobilis) carp, are invasive species that have negative impacts upon ecosystems. H. molitrix is known to jump completely out of the water in response to broadband sounds, however, this is not observed in H. nobilis. Preliminary experiments reveal that sounds can be used to modify the behavior of carps. Thus, understanding the hearing abilities of these species is important in order to design appropriate acoustical deterrents. Fish heads were preserved in 4% paraformaldehyde and the inner ears dissected and photographed under a light microscope in order to describe the …


An Investigation On The Cover Preference Of The Mountain Madtom (Noturus Eleutherus), Mariah Slaughter Jan 2020

An Investigation On The Cover Preference Of The Mountain Madtom (Noturus Eleutherus), Mariah Slaughter

Mahurin Honors College Capstone Experience/Thesis Projects

Madtom catfish, members of the genus Noturus, are common in the waters of the Southeastern US. A previous study observed that madtoms in the Green River, Kentucky, preferred to shelter within old mussel shells compared to under or next to rocks. A laboratory study on the Carolina Madtom (Noturus furiosus), found that they did not utilize mussel shells and preferred rocks as cover. I conducted a similar laboratory study to determine which cover options the Mountain Madtoms (Noturus eleutherus) prefer. Cover preference was determined by offering the madtoms shelter options (rocks or mussel shells) in …


Tourism And African Elephant (Loxodonta Africana) Behavior In Zambezi National Park, Zimbabwe, Brigit Rooney Jul 2019

Tourism And African Elephant (Loxodonta Africana) Behavior In Zambezi National Park, Zimbabwe, Brigit Rooney

Masters Theses & Specialist Projects

As charismatic megafauna and a flagship species, African elephants (Loxodonta africana) are vital to the African tourist economy. Conversely, high levels of wildlife tourism can induce behavioral shifts that push desired animals into less frequented areas and disrupt natural behaviors. In order to examine this trade-off, tourism levels and African elephant behaviors were studied in Zambezi National Park (ZNP) near Victoria Falls, Zimbabwe. Over the course of 14 weeks, in-person observations and camera traps in ZNP were used to collect geographic, demographic, and behavioral data from elephant sightings. As a proxy for human presence, geo-locational data were collected for each …


Documenting Marine Mammal Behavior And Evaluating The Benefits And Consequences Of Viewing Marine Mammals In Southcentral Alaska, Lauren E. Mccaslin Jul 2019

Documenting Marine Mammal Behavior And Evaluating The Benefits And Consequences Of Viewing Marine Mammals In Southcentral Alaska, Lauren E. Mccaslin

Masters Theses & Specialist Projects

Marine mammals are in a precarious conservation position because of anthropogenic impacts and historic perceptions that they are a consumable commodity. In light of changing abiotic conditions, further evaluation is needed on the habitat use, behavior, and interactions among marine mammals. Conservation legislation has helped protect species, but the greatest ground swelling may be the advent of the commercial whale watching industry. The feeding grounds in Alaskan waters have made this area a prime tourism location, and these nutrient-rich waters have resulted in a confluence of marine mammal species, including the appealing and abundant humpback whale (Megaptera novaeangliae) that may …


Investigations Of Biotremors In The Veiled Chameleon (Chamaeleo Calyptratus), Kathryn C. Laslie Jul 2018

Investigations Of Biotremors In The Veiled Chameleon (Chamaeleo Calyptratus), Kathryn C. Laslie

Masters Theses & Specialist Projects

While substrate-borne vibrations are utilized by different reptile species, true conspecific communication via biotremors has not yet been demonstrated in reptiles. This study follows a preliminary report that the veiled chameleon (Chamaeleo calyptratus) could produce biotremors in communicative contexts. I tested chameleon behavioral sensitivity to vibrations by placing them on a dowel attached to a shaker emitting vibrations of 25, 50, 150, 300, and 600 Hz and then measured their changes in velocity before and after the stimulus. I then paired chameleons in various social contexts [anthropogenic disturbance (human disruption of animal); dominance (malemale; female-female C. calyptratus); courtship (male-female C. …


An Evaluation Of Deterrent Methods Utilized To Prevent Crop Raiding By African Elephants (Loxodonta Africana) In The Kasigau Wildlife Corridor, Kenya, Rebecca Lynn Von Hagen Jul 2018

An Evaluation Of Deterrent Methods Utilized To Prevent Crop Raiding By African Elephants (Loxodonta Africana) In The Kasigau Wildlife Corridor, Kenya, Rebecca Lynn Von Hagen

Masters Theses & Specialist Projects

Escalating human elephant conflict (HEC) continues to be a contributing factor

towards elephant decline, and crop raiding is the most common form of negative

human-elephant interactions. For communities that cannot reverse or prevent crop

raiding, it is necessary to contain HEC events through deterrent measures. Few

deterrent measures exist that combine practicality and affordability while also

preventing habituation by elephants. This project focused on comparing the efficacy of

deterrent methods to assess which was the most successful at preventing elephants

from entering crops in the farming community of Sasenyi, Kenya. In this paired-control

study, four deterrent methods were evaluated: acacia …


Examination Of Enrichment Using Space And Food For African Elephants At The Sdzsp, Charlotte Hacker, Lance Miller, Bruce A. Schulte Jan 2018

Examination Of Enrichment Using Space And Food For African Elephants At The Sdzsp, Charlotte Hacker, Lance Miller, Bruce A. Schulte

Biology Faculty Publications

Concern for elephant welfare in zoological facilities has prompted a number of exhibit and management modifications, including those involving enrichment. Knowledge of how these changes impact indicators of welfare, such as elephant movement and behaviour, is crucial for continued improvement of elephant husbandry and care. The present study used observations and GPS-collected data to determine the effects of available space and food presentation on the walking distance and behaviour of thirteen African elephants (Loxodonta africana), which had a dominance structure ascertained by animal care staff at the San Diego Zoo Safari Park (SDZSP). This facility has two exhibits of approximately …


Substrate-Borne Communication In Chameleons: Do Vibrations Induce Behavioral Changes?, Emily J. Hamilton Sep 2017

Substrate-Borne Communication In Chameleons: Do Vibrations Induce Behavioral Changes?, Emily J. Hamilton

Mahurin Honors College Capstone Experience/Thesis Projects

Understanding the modes of communication used by a species is essential to understanding their ecology, behavior, and evolution. Substrate-borne vibrations have been reported to be produced by the veiled chameleon (Chamaeleo calyptratus), possibly implemented by use of a gular pouch. We found that veiled chameleons produced vibrations under dominance and mating behavioral contexts. We tested the sensitivity of veiled chameleons to vibrations by placing chameleons, one at a time, on a wooden dowel attached to a permanent magnetic shaker and recording each chameleon’s behavior before, during, and after a three-pulse vibrational stimulus of 25, 50, 150, 300, or 600 Hz. …


Body Shape Diversification Of Pecos Pupfish (Cyprinodon Pecosensis) On Varying Habitats As Evaluated By Geometric Morphometrics, Qianna Xu Apr 2017

Body Shape Diversification Of Pecos Pupfish (Cyprinodon Pecosensis) On Varying Habitats As Evaluated By Geometric Morphometrics, Qianna Xu

Masters Theses & Specialist Projects

During the 19th and 20th centuries, alterations to the Pecos River in New Mexico and Texas, USA due to anthropogenic activities, including damning and river channelization, vast water extraction for irrigation, as well as pollution of associated habitats, have greatly impacted the fish fauna within the drainage. One of the endemic fish species, the Pecos pupfish (Cyprinodon pecosensis), might be the most affected. Historically abundant and widespread large populations have been disrupted and became a series of small isolated subpopulations that persist at a few highly fragmented habitats restricted to a small area in southern New Mexico. The connectivity among …


The Effects Of Drought On Diets Of Apex Predators In The South African Lowveld Inferred By Fecal Hair Analysis, Shelby Wade Sep 2016

The Effects Of Drought On Diets Of Apex Predators In The South African Lowveld Inferred By Fecal Hair Analysis, Shelby Wade

Mahurin Honors College Capstone Experience/Thesis Projects

To properly manage offtake quotas and conservation efforts, Balule Nature Reserve (South Africa) instituted a study in 2014 to determine prey species selection by megapredators. In 2015, Balule Nature Reserve received about 190 mm less rainfall between the months of January and June than in 2014 (116 mm less than average). This study compares the diets of lions and hyaenas between 2014 and 2015. Prey species consumed were determined by fecal analysis, and results were compared to prey availability. Sixteen, 1 km2 plots were chosen from the 400 km2 Reserve. Between June and August 2015, we walked three, 1 km …


Impacts Of Habitat Fragmentation On The Cranial Morphology Of A Threatened Desert Fish (Cyprinodon Pecosensis), Michael Chaise Gilbert Jul 2016

Impacts Of Habitat Fragmentation On The Cranial Morphology Of A Threatened Desert Fish (Cyprinodon Pecosensis), Michael Chaise Gilbert

Masters Theses & Specialist Projects

Drastic alterations to the North American Southwest’s hydrology have highly influenced resident fish communities. In New Mexico and Texas, the Pecos River has been severely altered as a result of water manipulation, isolating backwaters and various habitats that were once connected to the main river. Cyprinodon pecosensis (Pecos pupfish) has been highly impacted due to the effects of anthropogenic water manipulation, as well as species introductions. Cyprinodon pecosensis populations have become isolated and scattered, residing in sinkholes, remnant lakes, and static backwaters, thus creating numerous micropopulations. The purpose of this study was to assess the morphological variation in cranial features …


Global Phylogeny Of The Water Penny Beetles Using Both Molecular And Morphological Evidence (Co, Mathew Vincent Wood Apr 2016

Global Phylogeny Of The Water Penny Beetles Using Both Molecular And Morphological Evidence (Co, Mathew Vincent Wood

Masters Theses & Specialist Projects

The Psephenidae is a family of freshwater beetles usually found in swift streams worldwide. Their unique disc shaped and flattened larvae have made this a group of interest for scientists for centuries. Morphologically, this family has been relatively well researched, and systematically the family is fairly well known and supported as monophyletic. One issue with Psephenidae, and with many other insect groups, is the lack of the molecular phylogenetic analyses to test morphology hypothesizes.

After successfully sequencing these two genes from species representing nearly all of the known genera, the data were analyzed using both Bayesian and parsimony methods. Analyses …


Wire Netting Reduces African Elephant (Loxodonta Africana) Impact To Selected Trees In South Africa, Kelly Derham, Michelle Henley, Bruce A. Schulte Jan 2016

Wire Netting Reduces African Elephant (Loxodonta Africana) Impact To Selected Trees In South Africa, Kelly Derham, Michelle Henley, Bruce A. Schulte

Biology Faculty Publications

African elephants (Loxodonta africana) are ecosystem engineers in that they substantially alter the environment through their unique foraging and feeding habits. At high densities, elephants potentially have negative impacts on the environment, specifically for large trees. Because of this, recent increases of elephants in the Associated Private Nature Reserves (APNR) on the western boundary of the Kruger National Park (KNP), South Africa, have caused concern regarding the survival of several tree species. Our objective was to assess the effectiveness of wrapping protective wire netting around the trunk of the tree for preventing and reducing bark stripping, branch breaking, …


Efficacy Of An Electronic Scarecrow On 4 Mammalian Crop-Raiders In Limpopo Province, South Africa, Merrie Renee Richardson Aug 2014

Efficacy Of An Electronic Scarecrow On 4 Mammalian Crop-Raiders In Limpopo Province, South Africa, Merrie Renee Richardson

Masters Theses & Specialist Projects

In South Africa, 2 primate species, Chacma baboons (Papio ursinus) and vervet monkeys (Chlorocebus pygerythrus), and 2 nocturnal mammals, Cape porcupine (Hystrix africaeaustralis) and bushbuck (Tragelaphus scriptus), are among many species of crop raiders. Currently, cost-effective, non-lethal solutions are lacking. From June through December 2012, I installed novel electronic scarecrows on two commercial citrus orchards and a private reserve and used video-recording remote cameras to assess cropraiders’ reactions to them in Limpopo Province, South Africa. I used focal animal sampling data from treatment and control group animals to examine differences in activity …


Level Of Uv-B Radiation Influences The Effects Of Glyphosate-Based Herbicide On Fitness Of The Spotted Salamander, Nicholas A. Levis May 2014

Level Of Uv-B Radiation Influences The Effects Of Glyphosate-Based Herbicide On Fitness Of The Spotted Salamander, Nicholas A. Levis

Masters Theses & Specialist Projects

Numerous causes have been implicated in contributing to amphibian population declines since the 1980's, with habitat modification, ultraviolet radiation (UV-B) and environmental contaminants (such as glyphosate-based herbicide) being among the most common. This study identifies the effects of a generic glyphosate-based herbicide (GLY- 4 Plus) on mortality, immune function, body condition, and morphological plasticity of larvae of the spotted salamander (Ambystoma maculatum) under conditions that reflect open and closed canopy light regimes. Larval salamander responses to glyphosate-based herbicide varied depending on UV-B conditions. In general, it appears that an open canopy (i.e. greater UV-B exposure) may confer fitness benefits. In …


Landscape Genetics Of The Marbled Salamander (Ambystoma Opacum) At Mammoth Cave National Park, James Kyle Martin Dec 2013

Landscape Genetics Of The Marbled Salamander (Ambystoma Opacum) At Mammoth Cave National Park, James Kyle Martin

Masters Theses & Specialist Projects

Habitat connectivity is important to maintain in order to prevent loss of genetic diversity, reduce inbreeding depression, and decrease extinction risk in threatened or endangered species. Here I present a landscape genetics study on marbled salamanders (Ambystoma opacum) in highly connected forested habitat at Mammoth Cave National Park. This investigation of gene flow among ponds within a mostly continuous landscape provides data that can be compared with patterns observed in more fragmented landscapes. These comparisons can provide a means of investigating the separate effects of structural and functional habitat connectivity on amphibian genetic population structure. Structural connectivity refers to the …


Comparison Of Creel Survey Data To Traditional Sampling Techniques In Pit-Lake Fisheries Of Muhlenberg County, Kentucky, Derek L. Rupert May 2012

Comparison Of Creel Survey Data To Traditional Sampling Techniques In Pit-Lake Fisheries Of Muhlenberg County, Kentucky, Derek L. Rupert

Masters Theses & Specialist Projects

Populations of largemouth bass, Micropterus salmoides, and bluegill, Lepomis macrochirus, were evaluated from five pit-lakes in Muhlenberg County, Kentucky, to determine if accurate proportional stock density (PSD) data can be obtained from a mandatory creel survey. It was hypothesized that the proportion of stock-to-quality (300-400mm) and quality (+400mm) largemouth bass from four years (2007-2010) of creel survey data would be statistically similar to those generated through on-site sampling in 2011. Fish were collected via a combination of gill netting, seining, hook-and-line fishing, and boat-mounted electro-fishing. In two of the pit-lakes, the sampling-generated length frequency data was not significantly …


Environment-Induced Phenotypic Plasticity In The Teeth Of Hatchery And Wild Largemouth Bass, Micropterus Floridanus, Tamilselvi Selvaraj Aug 2010

Environment-Induced Phenotypic Plasticity In The Teeth Of Hatchery And Wild Largemouth Bass, Micropterus Floridanus, Tamilselvi Selvaraj

Masters Theses & Specialist Projects

Successful stock enhancement of hatchery-reared fish depends heavily on the release of individuals able to demonstrate strong survival skills. Overall survival of fishes is a reflection of a successful blend of physiology, anatomy, and behavior. With fishes being highly phenotypically plastic, the potential exists for all aspects of hatcheryreared fish to vary significantly from their wild counterparts while having potentially adverse effects on their survival after release. Previous analyses have demonstrated significant differences between the feeding behavior of hatchery-reared and wild caught largemouth bass (Micropterus floridanus) in the laboratory, as well as differences in the development of the …


A Comparison Of The Singing Activity Of Carolina Wrens (Thryothorus Ludovicianus)) In Urban And Rural Settings, Shannon R. Trimboli May 2010

A Comparison Of The Singing Activity Of Carolina Wrens (Thryothorus Ludovicianus)) In Urban And Rural Settings, Shannon R. Trimboli

Masters Theses & Specialist Projects

As the earth’s landscape becomes increasingly urbanized, local wildlife must adapt to urban conditions or migrate to areas that are more rural. Urban wildlife face challenges such as direct loss of habitat, competition with non-native species, disturbance due to anthropogenic noise, and micro-climatic changes. Factors such as temperature, relative humidity, and noise affect the acoustical environment and may affect the ability of many animals, including birds, to communicate.

Understanding how urbanization affects birds’ singing behavior is critical because singing often plays a vital role in attracting mates and defending territories. In addition, as global climate change occurs it will become …


Elephants - A Conservation Overview, Heidi Riddle, Bruce A. Schulte, Ajay Desai, Laura Van Der Meer Jan 2010

Elephants - A Conservation Overview, Heidi Riddle, Bruce A. Schulte, Ajay Desai, Laura Van Der Meer

Biology Faculty Publications

Loss of habitat is one of the most significant problems facing elephants worldwide, leading to clashes over resources between wildlife and humans where elephants receive the largest part of blame – defined as Human Elephant Conflict (HEC). The sub-Saharan region of Africa contains an approximate population of 500,000 elephants that occupy 37 range countries. The African Elephant (Loxodonta africana) is categorized as Vulnerable in the Red List of Threatened Species; they are listed there as two distinct subspecies: the Savanna Elephant (L.a. africana) and the Forest Elephant (L. a. cyclotis). The Red List of …


On The Zoological Geography Of The Malay Archipelago (1859), Alfred Russel Wallace Nov 2009

On The Zoological Geography Of The Malay Archipelago (1859), Alfred Russel Wallace

Alfred Russel Wallace Classic Writings

No abstract provided.


The Development And Role Of Peripheral Auditory Structures In Otocinclus Affinis, Sri Kiran Kumar Reddy Botta Nov 2009

The Development And Role Of Peripheral Auditory Structures In Otocinclus Affinis, Sri Kiran Kumar Reddy Botta

Masters Theses & Specialist Projects

Loricariidae is a very diverse family of catfishes found primarily in the Amazon River basin. These catfishes have a unique characteristic feature of having fenestrae (holes) in the skull region (compound pterotic bone) adjacent to their bi-lobed swim bladder. Since the swim bladders and the compound pterotic may act as an external ear for hearing in this taxon, I hypothesized that these swim bladders structures have an acoustical functional in the loricariid Otocinclus affinis. In order to understand the development of these structures in O. affinis, I first monitored the ontogeny of the compound pterotic bone by clearing …


Prospecting For Mammalian Chemical Signals Via Solventless Extraction Techniques: An Elephantine Task, Thomas Goodwin, Bruce A. Schulte Jan 2009

Prospecting For Mammalian Chemical Signals Via Solventless Extraction Techniques: An Elephantine Task, Thomas Goodwin, Bruce A. Schulte

Biology Faculty Publications

In contrast to a plethora of known insect pheromones, a paucity of mammalian pheromones has been identified, two of which have been in elephants (Albone, 1984; Brown and Macdonald, 1985; Wyatt, 2003; Burger, 2005). Elephants possess one of the world’s best chemosensory systems, due in no small measure to their prehensile trunk. The trunk is not only the gateway to smelling (primary olfaction), but also the means by which chemical signals are conveyed from their source to the openings of the vomeronasal organ ducts in the roof of the mouth (the flehmen response; secondary olfaction) (Rasmussen, 1999). The late L. …


Investigation Of A Fresh African Elephant Carcass By Conspecifics, Christen Merte, Katie Gough, Bruce A. Schulte Jan 2009

Investigation Of A Fresh African Elephant Carcass By Conspecifics, Christen Merte, Katie Gough, Bruce A. Schulte

Biology Faculty Publications

This examination of elephant bones and ivory indicates that elephants show an elevated level of interest in conspecifics over other dead animals. Elephants do not seem to express special interest in dead kin but rather they appear to have a generalized response to injured, dying and deceased conspecifics (Douglas Hamilton et al. 2006, McComb et al. 2006). The present study reports the behaviour of a group of elephants in response to a euthanized adult male elephant that suffered severe wounds inflicted by a conspecific male. Most of the observations from previous studies have been on females and female group members. …