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Articles 1 - 24 of 24
Full-Text Articles in Biodiversity
Two New Species Of Byrrhinus Motschulsky, 1858 (Coleoptera, Limnichidae, Limnichinae) From Negros, Philippines, Emmanuel D. Delocado, Hendrik Freitag
Two New Species Of Byrrhinus Motschulsky, 1858 (Coleoptera, Limnichidae, Limnichinae) From Negros, Philippines, Emmanuel D. Delocado, Hendrik Freitag
Biology Faculty Publications
Two new species of Limnichidae beetles, Byrrhinus negrosensis sp. nov. and Byrrhinus villarini sp. nov., are described from the Island of Negros in the Philippines. The adult specimens of the new species can be differentiated by patterns of body punctation, colour and orientation of elytral pubescence, posterolateral angle of pronotum, tarsomere length ratio and aedeagal form. Two clades, representing the two new species, were retrieved in the Maximum Likelihood gene tree using the 3’-end of the COI gene. Maximum genetic divergence within B. negrosensis sp. nov. and B. villarini sp. nov. were recorded to be 2.3% and 1.3%, respectively, while …
Taxonomic Revision Of Agraphydrus Régimbart , 1903 V. Philippine Species And Their First Dna Barcodes (Coleoptera: Hydrophilidae: Acidocerinae), Hendrik Freitag, Albrecht Komarek
Taxonomic Revision Of Agraphydrus Régimbart , 1903 V. Philippine Species And Their First Dna Barcodes (Coleoptera: Hydrophilidae: Acidocerinae), Hendrik Freitag, Albrecht Komarek
Biology Faculty Publications
The genus Agraphydrus RÉGIMBART , 1903 (Coleoptera: Hydrophilidae) of the Republic of the Philippines is taxonomically reviewed aided by COI mtDNA barcodes. Nine new species are described, all currently regarded as endemic to the Philippines: A. abrasus sp.n., A. ampullatus sp.n., A. batak sp.n., A. brevilobatus sp.n., A. occultus sp.n., A. palawanensis sp.n., A. pelingeni sp.n., A. tenuipalpis sp.n., and A. zetteli sp.n. Agraphydrus coomani (O RCHYMONT , 1927), widespread in the Oriental and Australasian regions, is recorded from the Philippines for the first time. COI sequences and thorough morphological examination suggest that this and further species represent a group …
A New Genus And Two New Species Of Unarmed Hymenolepidid Cestodes (Cestoda: Hymenolepididae) From Geomyid Rodents In Mexico And Costa Rica, Scott Lyell Gardner, Altangerel Tsogtsaikhan Dursahinhan, Mariel Campbell, S. Elizabeth Rácz
A New Genus And Two New Species Of Unarmed Hymenolepidid Cestodes (Cestoda: Hymenolepididae) From Geomyid Rodents In Mexico And Costa Rica, Scott Lyell Gardner, Altangerel Tsogtsaikhan Dursahinhan, Mariel Campbell, S. Elizabeth Rácz
Scott L. Gardner Publications
Two new cestodes of the family Hymenolepididae are described from two species of rodents of the family Geomyidae collected in Mexico and Costa Rica. One new species of Hymenolepis is described from Cratogeomys planiceps Merriam 1895 from near Toluca, Mexico and another that we allocate to a new genus is described from Heterogeomys heterodus (Peters, 1865) from near Irazú Volcano, Costa Rica. Hymenolepis s. str. includes those Hymenolepididae with an apical organ, with no hooks on suckers or apical organ, and three testes. Hobergia irazuensis n. gen., n. sp. includes a hymenolepidid with an apical organ, unarmed scolex, small …
More Than Meets The Eye: A Taxonomic Revision Of The Nearctic Species In The Eye-Gnat Genus Amiota Loew (Diptera: Drosophilidae), Lance E. Jones
More Than Meets The Eye: A Taxonomic Revision Of The Nearctic Species In The Eye-Gnat Genus Amiota Loew (Diptera: Drosophilidae), Lance E. Jones
Dissertations and Theses
Thorough biotic inventories are still needed even in families as seemingly well-studied as fruit flies (Diptera: Drosophilidae). Drosophilidae have had an outsize effect on the history of biology because of their pivotal role in the study of genetics, development, and evolution. Even with more than a century of intensive study, large gaps remain in our understanding of diversity in this fly family, including well-studied areas such as North America. This thesis presents a taxonomic revision of the Nearctic species of the genus Amiota Loew. Twenty-three species are identified as being new to science for publication at a later date. This …
Repositories For Taxonomic Data: Where We Are And What Is Missing, Aurélian Miralles, Teddy Bruy, Katherine Wolcott, Mark D. Scherz, Dominik Begerow, Bank Beszteri, Michael Bonkowski, Janine Felden, Birgit Gemeinholzer, Frank Glaw, Frank Oliver Glöckner, Oliver Hawlitschek, Ivaylo Kostadinov, Tim W. Nattkemper, Christian Printzen, Jasmin Renz, Nataliya Rybalka, Marc Stadler, Tanja Weibulat, Thomas Wilke, Susanne S. Renner, Miguel Vences
Repositories For Taxonomic Data: Where We Are And What Is Missing, Aurélian Miralles, Teddy Bruy, Katherine Wolcott, Mark D. Scherz, Dominik Begerow, Bank Beszteri, Michael Bonkowski, Janine Felden, Birgit Gemeinholzer, Frank Glaw, Frank Oliver Glöckner, Oliver Hawlitschek, Ivaylo Kostadinov, Tim W. Nattkemper, Christian Printzen, Jasmin Renz, Nataliya Rybalka, Marc Stadler, Tanja Weibulat, Thomas Wilke, Susanne S. Renner, Miguel Vences
Harold W. Manter Laboratory: Library Materials
Natural history collections are leading successful large-scale projects of specimen digitization (images, metadata, DNA barcodes), thereby transforming taxonomy into a big data science. Yet, little effort has been directed towards safeguarding and subsequently mobilizing the considerable amount of original data generated during the process of naming 15,000–20,000 species every year. From the perspective of alpha-taxonomists, we provide a review of the properties and diversity of taxonomic data, assess their volume and use, and establish criteria for optimizing data repositories. We surveyed 4,113 alpha-taxonomic studies in representative journals for 2002, 2010, and 2018, and found an increasing yet comparatively limited use …
Temperate Eurasian Origins Of Hawaiian Chenopodium (Amaranthaceae) Plus Description Of A New Species Endemic To Moloka‘I, Jason T. Cantley, Angela J. Mcdonnell, J Branson, S R. Long, W Garnett, Christopher T. Martine
Temperate Eurasian Origins Of Hawaiian Chenopodium (Amaranthaceae) Plus Description Of A New Species Endemic To Moloka‘I, Jason T. Cantley, Angela J. Mcdonnell, J Branson, S R. Long, W Garnett, Christopher T. Martine
Faculty Journal Articles
Chenopodium taxa of Hawai‘i are tetraploids distinguished from other members of the circumglobally distributed genus by minute morphological characters. Because of these reasons, the geographic origin of Hawaiian Chenopodium has remained unclear. Across the Hawaiian Archipelago, Chenopodium taxa are morphologically variable and grow in highly disparate xeric habitats, especially in terms of precipitation, temperature, wind, salt spray, and solar irradiation. Habitats include dry subalpine shrublands, sandy beach strand of atolls in the Northwest Hawaiian Islands, dry forests, and precipitously tall sea cliffs of northwestern Moloka‘i. From the Moloka‘i sea cliffs, which are battered by high energy winds, salt spray, and …
A New Petrophilous Tiger Beetle From The Trans-Pecos Region Of Texas And Revised Key To The Genus Amblycheila (Coleoptera, Carabidae, Cicindelinae), Daniel P. Duran, Stephen J. Roman
A New Petrophilous Tiger Beetle From The Trans-Pecos Region Of Texas And Revised Key To The Genus Amblycheila (Coleoptera, Carabidae, Cicindelinae), Daniel P. Duran, Stephen J. Roman
School of Earth & Environment Faculty Scholarship
A new rock-dwelling (petrophilous) tiger beetle, Amblycheila katzi Duran & Roman, sp. nov. is described from calcareous canyons and steep hillsides in the Trans-Pecos region of western Texas. It is distinguished from all other Amblycheila based on multiple morphological characters, biogeography, and ecology. A revised key to the genus Amblycheila is provided.
Solanum Plastisexum, An Enigmatic New Bush Tomato From The Australian Monsoon Tropics Exhibiting Breeding System Fluidity., Angela J. Mcdonnell, Heather B. Wetreich, Jason T. Cantley, Peter Jobson, Christopher T. Martine
Solanum Plastisexum, An Enigmatic New Bush Tomato From The Australian Monsoon Tropics Exhibiting Breeding System Fluidity., Angela J. Mcdonnell, Heather B. Wetreich, Jason T. Cantley, Peter Jobson, Christopher T. Martine
Faculty Journal Articles
A bush tomato that has evaded classification by solanologists for decades has been identified and is described as a new species belonging to the Australian “Solanum dioicum group” of the Ord Victoria Plain biogeographic region in the monsoon tropics of the Northern Territory. Although now recognised to be andromonoecious, S. plastisexum Martine & McDonnell, sp. nov. exhibits multiple reproductive phenotypes, with solitary perfect flowers, a few staminate flowers or with cymes composed of a basal hermaphrodite and an extended rachis of several to many staminate flowers. When in fruit, the distal rachis may abcise and drop. A member of …
Cryptic Diversity In The North American Dromochorus Tiger Beetles (Coleoptera: Carabidae: Cicindelinae): A Congruence-Based Method For Species Discovery, Daniel P. Duran, David P. Herrman, Stephen J. Roman, Rodger A. Gwiazdowski, Jennifer A. Drummond, Glen R. Hood, Scott P. Egan
Cryptic Diversity In The North American Dromochorus Tiger Beetles (Coleoptera: Carabidae: Cicindelinae): A Congruence-Based Method For Species Discovery, Daniel P. Duran, David P. Herrman, Stephen J. Roman, Rodger A. Gwiazdowski, Jennifer A. Drummond, Glen R. Hood, Scott P. Egan
School of Earth & Environment Faculty Scholarship
A fundamental problem in biodiversity science is determining the number of species in any taxon, and there is a growing awareness that cryptic diversity contributes to this problem – even in well-studied groups. Discovering cryptic species requires several lines of evidence to elucidate congruent patterns across data-types, and distinguish unrecognized species. Tiger beetles are among the most well-studied insect groups; yet few new North American species have been described since the mid-20th century, suggesting that that the number of morphologically distinct species is reaching an asymptote. We explore the possibility that more species exist in the fauna as cryptic species, …
Dna Barcoding As A Tool For Species Discovery And Documentation In The Superfamily Ichneumonoidea, Sarah Meierotto
Dna Barcoding As A Tool For Species Discovery And Documentation In The Superfamily Ichneumonoidea, Sarah Meierotto
Theses and Dissertations--Entomology
Changes to traditional taxonomic methods to incorporate new technologies and methods have already improved the quality of species hypotheses, but more work can be done to improve the speed of new species documentation. The mitochondrial COI DNA barcode has been successfully used to identify species with high accuracy since the early 2000s, and has been used in conjunction with morphological examinations and other DNA markers to discover and delimit new species. This thesis explores the application of DNA barcodes as the primary data for delimitation and diagnosis of new species of ichneumonoids.
The genera Zelomorpha and Hemichoma are revised and …
Three New Species Of Grouvellinus Champion, 1923 From Maliau Basin, Sabah, Borneo, Discovered By Citizen Scientists During The First Taxon Expedition (Insecta, Coleoptera, Elmidae), Hendrik Freitag, Iva Njunjic, Clister V. Pangantihon
Three New Species Of Grouvellinus Champion, 1923 From Maliau Basin, Sabah, Borneo, Discovered By Citizen Scientists During The First Taxon Expedition (Insecta, Coleoptera, Elmidae), Hendrik Freitag, Iva Njunjic, Clister V. Pangantihon
Biology Faculty Publications
Further results are presented of the first field course at Maliau Basin, Malaysian Borneo organized by Taxon Expeditions, an organization which enables citizen scientists to be directly involved in taxonomic discoveries. Three new species of the aquatic beetle genus Grouvellinus Champion, 1923, namely G. leonardodicaprioi sp. n., G. andrekuipersi sp. n., and G. quest sp. n. were collected jointly by the citizen scientists and taxonomists during the fieldwork in Maliau Basin. Material was mainly sampled from sandstone bottom rocks of blackwater streams at altitudes between 900 m and 1,000 m using fine-meshed hand-nets. The genus is widely distributed …
Marine Tardigrades From South Carolina, Usa, Paul J. Bartels, Lilvia J. Bradbury, Diane R. Nelson
Marine Tardigrades From South Carolina, Usa, Paul J. Bartels, Lilvia J. Bradbury, Diane R. Nelson
Journal of the South Carolina Academy of Science
Until now, there have been only three studies of marine tardigrades from South Carolina, USA. The Fall 2015 invertebrate zoology class from Warren Wilson College collected four sediment samples from Huntington Beach State Park: shallow beach sand, deep beach sand, shallow salt marsh sediment, and deep salt marsh sediment. No tardigrades were found in the salt marsh, but two species were found in the beach samples. Batillipes pennaki Marcus, 1946 was found in shallow and deep beach sand, and three specimens of a potentially new species of Stygarctus Schulz, 1951 were found in the deep beach sand at groundwater level. …
An Endemic Commensal Leucothoid Discovered In The Tunicate Cnemidocarpa Bicornuta, From New Zealand (Crustacea, Amphipoda), Kaitlyn M. Brucker
An Endemic Commensal Leucothoid Discovered In The Tunicate Cnemidocarpa Bicornuta, From New Zealand (Crustacea, Amphipoda), Kaitlyn M. Brucker
HCNSO Student Theses and Dissertations
Precise descriptions and comprehensive taxonomies of species and their ecology are essential in monitoring changes in marine biodiversity at multiple spatial scales. A currently undescribed species of commensal amphipod in the genus Leucothoe is reported from New Zealand, collected from the endemic tunicate Cnemidocarpa bicornuta. This species differs from others in the genus in having a one-articulate first maxilla palp and an apically produced tuberculate lobe on the inner margin of the outer plate of the maxilliped. Previous taxonomic surveys in New Zealand waters did not document this species, indicating that it may be a recent arrival. This research …
Two New Species Of Geosesarma De Man, 1892 (Crustacea: Brachyura: Sesarmidae) From Palawan, The Philippines, Hendrik Freitag, Marivene Manuel-Santos, Peter K.L Ng
Two New Species Of Geosesarma De Man, 1892 (Crustacea: Brachyura: Sesarmidae) From Palawan, The Philippines, Hendrik Freitag, Marivene Manuel-Santos, Peter K.L Ng
Biology Faculty Publications
Two new and relatively large species of semi-terrestrial sesarmid crabs of the genus Geosesarma De Man, 1892, are described from Palawan Island, Philippines. They are distinguishable from congeners by the characteristic structure of their carapace, chelipeds, ambulatory legs and male first gonopods. Aspects of their ecology are briefly discussed
Hymenolepis Folkertsi N. Sp. (Eucestoda: Hymenolepididae) In The Oldfield Mouse Peromyscus Polionotus (Wagner) (Rodentia: Cricetidae: Neotominae) From The Southeastern Nearctic With Comments On Tapeworm Faunal Diversity Among Deer Mice, Arseny A. Makarikov, Todd N. Nims, Kurt E. Galbreath, Eric P. Hoberg
Hymenolepis Folkertsi N. Sp. (Eucestoda: Hymenolepididae) In The Oldfield Mouse Peromyscus Polionotus (Wagner) (Rodentia: Cricetidae: Neotominae) From The Southeastern Nearctic With Comments On Tapeworm Faunal Diversity Among Deer Mice, Arseny A. Makarikov, Todd N. Nims, Kurt E. Galbreath, Eric P. Hoberg
Harold W. Manter Laboratory of Parasitology: Faculty and Staff Publications
A previously unrecognized species of hymenolepidid cestode attributable to Hymenolepis is described based on specimens in Peromyscus polionotus, oldfield mouse, from Georgia, United States, near the southeastern coast of continental North America. Specimens of Hymenolepis folkertsi n. sp. differ from those attributed to most other species in the genus by having testes arranged in a triangle and a scolex with a prominent rostrum-like protrusion. The newly recognized species is further distinguished by the relative position and length of the cirrus sac, shape of seminal receptacle, and relative size of external seminal vesicle and seminal receptacle. Hymenolepidid cestodes have sporadically …
Ficotylus Laselvae N. Sp. (Tylenchomorpha: Anguinidae) Associated With Ficus Colubrinae In Costa Rica, Robin M. Giblin-Davis, Natsumi Kanzaki, Kerrie A. Davies, Weimin Ye, Yongsan Zeng, Barbara J. Center, Alejandro Esquivel, Thomas O. Powers
Ficotylus Laselvae N. Sp. (Tylenchomorpha: Anguinidae) Associated With Ficus Colubrinae In Costa Rica, Robin M. Giblin-Davis, Natsumi Kanzaki, Kerrie A. Davies, Weimin Ye, Yongsan Zeng, Barbara J. Center, Alejandro Esquivel, Thomas O. Powers
Department of Plant Pathology: Faculty Publications
Ficotylus laselvae n. sp. was recovered from under the bracts of figs (syconia) of Ficus colubrinae from La Selva, Costa Rica, during a survey of nematode rainforest biodiversity and is described herein. This is only the second report of an association between the nematode suborder Tylenchina and the sycones of figs. Previous reports of most nematode associates of the sycones of figs have been from the lumen and involved transmission by female fig wasp pollinators (Agaonidae) during pollination/oviposition (e.g., Schistonchus and Parasitodiplogaster spp.). The association between F. laselvae n. sp. and Ficus colubrinae may involve an invertebrate host, …
Revision Of Anacaena Thomson, 1859 Xi. Republic Of The Philippines (Coleoptera: Hydrophilidae), Hendrik Freitag
Revision Of Anacaena Thomson, 1859 Xi. Republic Of The Philippines (Coleoptera: Hydrophilidae), Hendrik Freitag
Biology Faculty Publications
The species of Anacaena THOMSON, 1859 (Coleoptera: Hydrophilidae) of the Republic of the Philippines are revised. Fifteen new species are described: Anacaena albay sp.n., A. amplocomata sp.n., A. apo sp.n., A. balabag sp.n., A. cordillera sp.n., A. davao sp.n., A. destructa sp.n., A. emergens sp.n., A. hemisphaerica sp.n., A. levistriata sp.n., A. philippina sp.n., A. princesa sp.n., A. quezona sp.n., A. sulcata sp.n., and A. zamboangana. All species are endemic to the Philippines, twelve are aquatic, the habitat of three species is unknown. All Philippine species are morphologically similar to other species of the eastern Oriental Region and probably do …
The Scorpions Of Hainan Island, China (Arachnida: Scorpiones), Zhi-Yong Di, Zhi-Jian Cao, Ying-Liang Wu, Lin Zhu, Hui Liu, Wen-Xin Li
The Scorpions Of Hainan Island, China (Arachnida: Scorpiones), Zhi-Yong Di, Zhi-Jian Cao, Ying-Liang Wu, Lin Zhu, Hui Liu, Wen-Xin Li
Euscorpius
The redescriptions and illustrations of three species, Isometrus (Isometrus) maculatus (DeGeer, 1778), Lychas mucronatus (Fabricius, 1798) (Buthidae), and Liocheles australasiae (Fabricius, 1775) (Hemiscorpiidae) from Hainan Island, China are presented. Distribution data and updated key of Hainan scorpions are provided.
Review Of The Subgenus Pandinurus Fet, 1997 With Descriptions Of Three New Species (Scorpiones, Scorpionidae, Pandinus)., František Kovařík
Review Of The Subgenus Pandinurus Fet, 1997 With Descriptions Of Three New Species (Scorpiones, Scorpionidae, Pandinus)., František Kovařík
Euscorpius
Pandinus (Pandinurus) awashensis sp. n. from Ethiopia, P. (P.) somalilandus sp. n. from Somaliland, and P. (P.) lowei sp. n. from Democratic Republic of Congo are described and compared with other species of the subgenus. P. (P.) sudanicus Hirst, 1911 stat. n. previously considered a synonym of P. (P.) magrettii Borelli, 1901 is recognized as a valid species. New data on taxonomic characters and distribution of the subgenus Pandinurus are presented. Also presented are a map of distribution, photos of the localities and a key to species using hitherto unpublished characters of the tarsomere spination formula and sexual …
Ancyronyx Erichson, 1847 (Coleoptera, Elmidae) From Mindoro, Philippines, With Description Of The Larvae And Two New Species Using Dna Sequences For The Assignment Of The Developmental Stages, Hendrik Freitag
Biology Faculty Publications
Ancyronyx buhid sp. n. and Ancyronyx tamaraw sp. n. are described based on adults and larvae, matched using their cox1 or cob DNA sequence data. Additional records of Ancyronyx schillhammeri Jäch, 1994 and Ancyronyx minerva Freitag & Jäch, 2007 from Mindoro are listed. The previously unknown larva of Ancyronyx schillhammeri is also described here, aided by cox1 data. The new species and larval stages are described in detail and illustrated by SEM and stacked microscopic images. Keys to the adult and larval Ancyronyx species of Mindoro and an updated checklist of Philippine Ancyronyx species are provided. The usefulness as bioindicators, …
A New Intertidal Shore Bug (Heteroptera: Saldidae) From The Philippines, Hendrik Freitag, Herbert Zettel, Clister V. Pangantihon
A New Intertidal Shore Bug (Heteroptera: Saldidae) From The Philippines, Hendrik Freitag, Herbert Zettel, Clister V. Pangantihon
Biology Faculty Publications
Salduncula carmencitae sp.n. is described, its habitus and the paramere and parandria of the male are illustrated. This shore bug was found on Mindanao Island, the Philippines, in the intertidal zone. Habitat and behaviour of the species are briefly described. The distinguishing diagnostic characters of species of Salduncula Brown, 1954 are discussed.
A Revision Of Erythranthe Montioides And Erythranthe Palmeri (Phrymaceae), With Descriptions Of Five New Species From California And Nevada, Usa, Naomi S. Fraga
A Revision Of Erythranthe Montioides And Erythranthe Palmeri (Phrymaceae), With Descriptions Of Five New Species From California And Nevada, Usa, Naomi S. Fraga
Aliso: A Journal of Systematic and Floristic Botany
The genus Erythranthe (Phrymaceae) continues to be a source of floristic novelty, especially in the American West, as evidenced by recent discoveries presented here. Two species in Erythranthe section Paradantha (E. montioides and E. palmeri) have long been a source of taxonomic confusion. Recent research reveals that a total of ten species have previously been treated as part of these two species. Here I present a revised taxonomy for E. montioides and E. palmeri including clarification of species circumscriptions (E. montioides, E. palmeri), recognition of three species that have previously been treated as synonyms …
A New Species Of Genus Laboulbenia (Laboulbeniales) On Craspedophorus Formosanus (Coleoptera, Carabidae) From Taiwan, With A Note On Laboulbenia Asiatica, Katsuyuki Terada, Meng-Hao Hsu, Wen-Jer Wu
A New Species Of Genus Laboulbenia (Laboulbeniales) On Craspedophorus Formosanus (Coleoptera, Carabidae) From Taiwan, With A Note On Laboulbenia Asiatica, Katsuyuki Terada, Meng-Hao Hsu, Wen-Jer Wu
Aliso: A Journal of Systematic and Floristic Botany
Laboulbenia taiwaniana sp. nov. on Craspedophorus formosanus is described from Taiwan and illustrated with photographs. It is characterized by a long, asymmetrical perithecium with an oblique apex and a pale-colored lower wall, a slender, evenly tapered receptacle with cylindrical cell I and II and trapezoidal cell IV, well developed appendages with black septa concentrated in the basal portion of the appendage system, and especially by black septa on the distal end of cell g. Laboulbenia asiatica, which was described from an Asian carabid---"Casnonia sp."---in 1899 and was illustrated in 1908 by Thaxter, is reviewed and compared …
The Genus Prolixandromyces (Laboulbeniales) In The Old World Prolixandromyces (Laboulbeniales), Alex Weir
The Genus Prolixandromyces (Laboulbeniales) In The Old World Prolixandromyces (Laboulbeniales), Alex Weir
Aliso: A Journal of Systematic and Floristic Botany
Two new species of Prolixandromyces (Laboulbeniales) parasitic on Rhagovelia (Heteroptera: Veliidae) are described and illustrated: Prolixandromyces benjaminii from the Philippine Islands and P. lanceolatus from Africa (Madagascar, Malawi, Nigeria, Sierra Leone, and South Africa) and Asia (Indonesia: Sulawesi, Sumba, Timor; Malaysia: Selangor; and Sri Lanka). In order to accommodate these species the circumscription of Prolixandromyces is emended. The new taxa are compared with the one known species from the Old World, P. triandrus from Spain, and with the five known New World taxa.