Life Sciences Commons^™

Evolutionary Origins Of Secondary Growth - The Periderm Perspective: Integrating Evidence From Fossils And Living Plants, Madison A.K. Lalica

Cal Poly Humboldt theses and projects

Periderm is a structural feature with roles in protection of inner plant tissues and wound healing. Knowledge of periderm occurrences in the fossil record and living lineages outside the seed plants is limited and its evolutionary origins remain poorly explored. Here, I review the known taxonomic distribution of canonical periderm (typical ontogenetic stage) and wound periderm (self-repair mechanism). To this sparse body of data I add new observations and experiments on living plant lineages and new occurrences from the fossil record. One of the latter, documented in the new early euphyllophyte species Nebuloxyla mikmaqiana, joins the oldest known periderm …

Characterizing New Plant Fossils With Woody Growth From The Battery Point Formation Of Quebec (Canada), Emma T. Casselman

Cal Poly Humboldt theses and projects

The oldest woody growth has been recognized in several Early Devonian (ca. 410-395 Ma) euphyllophytes. Their taxonomic diversity is difficult to evaluate due to a lack of quantitative methods for comparing plants based on secondary xylem anatomy. In turn, this hinders understanding of their implications for the evolution of plant development. To develop metrics that quantify secondary xylem anatomy and allow for conclusive comparisons, I investigated extant taxa and Early Devonian fossil woody taxa. I developed multiple metrics for quantifying tracheid size as a function of position in the secondary xylem and tested them on a dataset of extant gymnosperms. …

Analyzing The Shark Paleoecology Of Coastal Georgia From The Miocene And Pliocene Epochs, Joshua Lee Clark, Benjamin Angalet

Georgia Journal of Science

The field of shark paleoecology often yields indecisive conclusions based on the limited fossilization of their anatomical structures, with the exception of their teeth. The majority of the Atlantic coast has been studied regarding the presence of certain prehistoric shark species from the Miocene, Pliocene, and Pleistocene epochs. However, information pertaining to the Georgia coast and understanding its potential community structure is relatively understudied. This study was conducted in which thousands of fossil shark specimens and subsequent marine fauna were collected from dredge spoils created by the US Army Corps of Engineers (USACE): Savannah District. A total of 5,127 fossil …

A Cladoxylopsid With Complex Vascular Architecture From The Early Devonian Battery Point Formation (Québec, Canada), Jessica Chu

Cal Poly Humboldt theses and projects

Cladoxylopsids are seed-free plants that formed the world’s earliest forests and gave rise to horsetails (sphenopsids), but their evolutionary origins are poorly understood. Here, I describe a new Early Devonian cladoxylopsid from the Emsian (400-395 Ma) Battery Point Formation (Québec, Canada). The structural features of this plant indicate taxonomic affinity to cladoxylalean cladoxylopsids and its phylogenetic position supports placement in genus Cladoxylon (the proposed name, Cladoxylon kespekianum sp. nov., to be formalized by peer-reviewed publication). Specimens of this plant are permineralized in calcium carbonate and were studied using the acetate peel technique with light and electron microscopy. Cladoxylon kespekianum is …

Extinct Or Extant? A New Species Of Termitodius Wasmann, 1894, (Coleoptera: Scarabaeidae: Aphodiinae: Rhyparini) With A Short Review Of The Genus, Paul E. Skelley, Julian Clavijo-Bustos, Oliver Keller

Insecta Mundi

A new species of Termitodius Wasmann (Coleoptera: Scarabaeidae: Aphodiinae: Rhyparini) is de­scribed from Colombia, Termitodius woodruffi Skelley, Clavijo-Bustos, and Keller, new species. This species is both extant and abundantly preserved in copal. The genus Termitodius is reviewed with a key and brief accounts to all species.

Una nueva especie de Termitodius Wasmann (Coleoptera: Scarabaeidae: Aphodiinae: Rhyparini) es descrita de Colombia, Termitodius woodruffi Skelley, Clavijo-Bustos, y Keller, nueva especie. Esta especie es existente y abundantemente preservada en copal. El género Termitodius es revisado con una clave y reseñas breves para todas las especies.

The pantropical tribe Rhyparini is represented …

A New Species Of Leptorhyparus Howden, 2003 (Coleoptera: Scarabaeidae: Aphodiinae: Rhyparini) In Amber From The Dominican Republic, With Comments On Extant Species, Paul E. Skelley

Insecta Mundi

A new species of Leptorhyparus Howden, 2003 (Coleoptera: Scarabaeidae: Aphodiinae: Rhyparini) is described, Leptorhyparus quadricornis Skelley, in amber from the Dominican Republic. Extant species of Leptorhyparus are reviewed, with additional collection data and diagnostic characters presented.

Se describe una nueva especie en el género Leptorhyparus Howden, 2003 (Coleoptera: Scarabaeidae: Aphodiinae: Rhyparini), Leptorhyparus quadricornis Skelley, en ámbar de la República Dominicana. Se revisan las especies vivientes del género Leptorhyparus, agregándose datos de colección y caracteres diagnósticos.

New Genus Of Rhyparini (Coleoptera: Scarabaeidae: Aphodiinae) In Amber From The Dominican Republic, Paul E. Skelley

Insecta Mundi

Priscyparus quisqueyensis Skelley, new genus and new species (Coleoptera: Scarabaeidae: Aphodiinae: Rhyparini), is described in amber from the Dominican Republic. An updated key to New World genera of Rhyparini is presented.

Priscyparus quisqueyensis Skelley, género nuevo y especie nueva (Coleoptera: Scarabaeidae: Aphodiinae: Rhyparini), se describe de ámbar de la República Dominicana. La clave a los Rhyparini del Nuevo Mundo es enmendada para incluir el nuevo género.

There are only three fossil taxa of Rhyparini (Coleoptera: Scrabaeidae: Aphodiinae) preserved in copal or amber from the New World. There have been brief listings of a Termitodius Wasmann in Colombian copal (Penney and …

Ecological Controls On The Campanian Distribution Of Hesperornis (Aves: Hesperornithiformes) In The Western Interior Seaway, Blake Chapman

Master's Theses

The epicontinental Western Interior Seaway (WIS) of Late Cretaceous North America provided a unique marine habitat for cephalopods, fish, marine reptiles, and the foot-propelled diving seabird Hesperornis. While several predator-prey relationships among Hesperornis or other hesperornithiforms and other WIS animals have been hypothesized based on gut contents, bite marks, and coprolites/colonites, ecological relationships have not been quantitatively tested. Paleontological species distribution modeling (SDM) studies have focused on extinct non-marine taxa and marine invertebrates, with only two marine vertebrate studies of extant taxa. Here, two SDM methods were used to test the influence of vertebrate faunas, sedimentary rock type, paleogeography, …

Curation Techniques Of Small-Sized Natural History Specimens: A Collection Of Microfossils., Zachary V Biven-Leslie, Guillermo W. Rougier

Undergraduate Arts and Research Showcase

Small-sized fossil specimens are challenging regarding safety, storage and traceability of information. A recent collection of several hundred minute early mammalian teeth and jaws (most about 1mm) made impossible the standard practices of specimen numbering. A reliable association of the specimen and the collection number is the minimum required, however, a number of other data easily recoverable from the specimen/curating material is desirable. Geological provenance, locality, systematic and anatomical ID, collector, year, etc. is often included if possible. Use of the specimen is expected to be on the order of hundreds of years, manipulated by investigators, transported, and stored in …

A Morphological Phylogeny Of Odonatoptera: Examining Missing Data In A Group With A Lot Of "Naturally" Missing Data, Robert James Erickson

Theses and Dissertations

Odonatopera exhibit a wide diversity of morphologies for an ancient group of winged insects. A morphological matrix of 463 characters is compiled for 347 extant and fossil representatives used in parsimony analyses, implemented in TNT, to document arrangements of taxonomic groups above the family level. Missing data and other challenges approaches implemented and interpretation of the results. We employ a novel approach to testing monophyly relative to quantities of missing data for each taxon. Phylogenetic reconstructions recover patterns of monophyly and trends based on missing data. We discuss the implications of our findings on missing data as well as limitations …

Ashfall Fossil Beds: From Waterhole To Rhino Barn, Sandy Mosel

University of Nebraska State Museum: Programs Information

Twelve million years ago a volcanic super-eruption in what is now southwestern Idaho created an enormous ashfall that blanketed the Great Plains with several inches of volcanic ash, devastating the local wildlife. Evidence of this geologic event lies within the hills of the Niobrara River Valley in northeast Nebraska, at a place aptly named the Ashfall Fossil Beds State Historical Park. Located near the tiny village of Royal, the Ashfall Park has been developed around a waterhole where animals succumbed after breathing in the volcanic dust. Buried deeply by windblown volcanic ash in the weeks following the eruption, the skeletons …

An Assessment Of Convergence In The Feeding Morphology Of Xiphactinus Audax And Megalops Atlanticus Using Landmark-Based Geometric Morphometrics, Edward Chase Shelburne

Master's Theses

Convergence is an evolutionary phenomenon wherein distantly related organisms independently develop features or functional adaptations to overcome similar environmental constraints. Historically, convergence among organisms has been speculated or asserted with little rigorous or quantitative investigation. More recent advancements in systematics has allowed for the detection and study of convergence in a phylogenetic context, but this does little to elucidate convergent anatomical features in extinct taxa with poorly understood evolutionary histories. The purpose of this study is to investigate one potentially convergent system—the feeding structure of Xiphactinus audax (Teleostei: Ichthyodectiformes) and Megalops atlanticus (Teleostei: Elopiformes)—using a comparative anatomical approach to assess …

Exploring The Origins Of Secondary Growth – Two Lower Devonian (Emsian) Euphyllophytes From Gaspé, Canada, And Their Implications For The Evolution Of Secondary Growth, Kelly C. Pfeiler

Cal Poly Humboldt theses and projects

Secondary growth from a vascular cambium has a deep fossil record among euphyllophytes, with the earliest occurrence dated to 407 Ma. To date, Armoricaphyton and Franhueberia represent the only instances of secondary growth formally documented in the Early Devonian. Secondary growth diversified rapidly and was present in all major euphyllophyte lineages by the Middle Devonian. Here, I describe two new Early Devonian euphyllophytes exhibiting secondary growth, from the Emsian (c. 400-395 Ma) Battery Point Formation (Québec, Canada): Gmujij tetraxylopteroides gen. et sp. nov. and Perplexa praestigians gen et sp. nov. Both these plants possess mesarch actinosteles with Psilophyton-type …

Crossed Tracks: Mesolimulus, Archaeopteryx, And The Nature Of Fossils, Leonard Finkelman

Faculty Publications

Organisms leave a variety of traces in the fossil record. Among these traces, vertebrate and invertebrate paleontologists conventionally recognize a distinction between the remains of an organism’s phenotype (body fossils) and the remains of an organism’s life activities (trace fossils). The same convention recognizes body fossils as biological structures and trace fossils as geological objects. This convention explains some curious practices in the classification, as with the distinction between taxa for trace fossils and for tracemakers. I consider the distinction between “parallel taxonomies,” or parataxonomies, which privileges some kinds of fossil taxa as “natural” and others as “artificial.” The motivations …

Extending The Fossil Record Of The Polytrichaceae (Bryophyta): Insights From The Early Cretaceous Of Vancouver Island, Canada, Alexander C. Bippus

Cal Poly Humboldt theses and projects

Diverse in modern ecosystems, mosses are dramatically underrepresented in the fossil record. Furthermore, most pre-Cenozoic mosses are known only from compression fossils, which lack detailed anatomical information. Lower Cretaceous deposits at Apple Bay (Vancouver Island, British Columbia, Canada) contain a diverse anatomically preserved flora that includes numerous bryophytes, many of which have yet to be characterized. Among them is a polytrichaceous moss that is described here as Meantoinea alophosioides gen. et sp. nov. Meantoinea alophosioides represents the first occurrence of gemma cups in a fossil moss and is the oldest unequivocal record of Polytrichaceae, providing a hard minimum age for …

A Scorpion From A Peatbog: The First Arthropod Fossil From The Late Viséan Of The Moscow Coal Basin, Victor Fet, Michael E. Soleglad, Yulia V. Mosseichik, Dmitry E. Shcherbakov

Victor Fet

This note describes a fossil fragment from the Upper Viséan (ca. 330 Ma), of the Moscow Coal Basin (Novgorod Region, Russia). The fossil is identified as the cuticle of a single leg segment (left femur) of a scorpion (Arachnida: Scorpiones), without any further taxonomic placement. The anatomical details of the fragment are given, along with ecological considerations. It is the first fossil scorpion record from Russia.

A Scorpion From A Peatbog: The First Arthropod Fossil From The Late Viséan Of The Moscow Coal Basin, Victor Fet, Michael E. Soleglad, Yulia V. Mosseichik, Dmitry E. Shcherbakov

Euscorpius

This note describes a fossil fragment from the Upper Viséan (ca. 330 Ma), of the Moscow Coal Basin (Novgorod Region, Russia). The fossil is identified as the cuticle of a single leg segment (left femur) of a scorpion (Arachnida: Scorpiones), without any further taxonomic placement. The anatomical details of the fragment are given, along with ecological considerations. It is the first fossil scorpion record from Russia.

Carnivores From The Late Miocene Locality Of Hayranlı (Hayranlı, Sivas, Turkey), Şaki̇r Önder Özkurt, Erksi̇n Güleç, Ahmet Cem Erkman

Turkish Journal of Zoology

The locality of Hayranlı-Sivas is situated in the central Anatolian plateau (Sivas, Turkey) and includes many fossil remains, including carnivores collected from 2 different spots. The aim of this study is to contribute to knowledge of carnivore evolution in Turkey based on the fossil specimens in Hayranlı-Sivas. The studied collection indicates the presence of the following taxa: Hyaenictitherium wongii, Hyaenictitherium intuberculatum, Lycyaena dubia, and Machairodus giganteus. L. Dubia is the first record from Anatolia. The material of each taxon was described and determined by comparison with other materials from various Eurasian localities. During the Early or Middle Turolian 9-7 Ma. …

Establishment Of Larrea Tridentata At The Northern Edge Of The Modern Mojave Desert: Insights From Neotoma Paleomiddens, Clare Steinberg

Biology ETDs

Shifting climates affect the composition of biological communities. If environmental conditions change sufficiently, new species can invade, leading to large-scale community turnover. Understanding how and why such shifts occur is crucial in this era of anthropogenic global change. Paleontological studies provide a valuable long-term perspective of the dynamics of community turnover. Here, we examine changes in the plant community over the past 34 thousand years in what is now the northern Mojave Desert. This time period includes the last glacial maximum as well as numerous smaller climatic fluctuations in the Holocene and the end of the Pleistocene. We quantified plant …

Developing Criteria For Identifying Fossil Raindrop Prints, April Menendez

The Research and Scholarship Symposium (2013-2019)

Raindrop imprints have been identified many times in the rock record as stated in papers by Rindsberg, Shul’ga 2004 and Kahle 2009. However, there is a question of whether those raindrops have been determined accurately or if the spherical depressions are caused by some other occurrence. Studies by Twenhofel have been done that describe raindrop prints in modern sediment, and the research conducted during this project continues to broaden that field of information. During this research project, experiments were carried out to collect the appearance of raindrop prints in sand, both wet and dry. After photographs of the imprints were …

New Lauraceous Species From The Siwalik Forest Of Arunachal Pradesh, Eastern Himalaya, And Their Palaeoclimatic And Palaeogeographic Implications, Mahasin Ali Khan, Subir Bera

Turkish Journal of Botany

Leaf remains of 4 lauraceous taxa collected from the Siwalik sediments (Middle Miocene to Pliocene) of Arunachal Pradesh of eastern Himalaya are described in the present communication. One fossil leaf, Cinnamomum palaeobejolghota sp. nov., was recovered from the lower part of the Siwalik succession (Dafla Formation; Middle to Upper Miocene), while Persea mioparviflora sp. nov., Persea preglaucescens sp. nov., and Lindera neobifaria sp. nov. were collected from the middle part of the Siwalik succession (Subansiri Formation; Pliocene) of Arunachal sub-Himalaya. The geographic distribution of the fossils and their modern counterparts is discussed and on that basis tropical vegetation under warm …

Ichnology And Paleoecology Of The Jurassic Aztec Sandstone, Heather Marie Stoller

UNLV Theses, Dissertations, Professional Papers, and Capstones

In this study I describe and interpret tracks and trackways of the Jurassic Aztec Sandstone of southern Nevada and southern California. This study involved mapping of all known tracks and trackways, including foot length, stride length, and trackway width. Photogrammetric data, collected by Bureau of Land Management scientists, were utilized for several trackways in Red Rock Canyon National Conservation Area.

More than three hundred tracks belonging to five ichnotaxa were documented within the Aztec Sandstone, including about 165 tridactylGrallator, 250 tetradactyl (four-toed)Brasilichniumtracks, and 7 arthropod trackways ofOctopodichnusandPaleohelcura. Four of the five ichnotaxa were not previously reported from the Aztec Sandstone. …

Paleoecology Of Late Pleistocene Megaherbivores: Stable Isotope Reconstruction Of Environment, Climate, And Response, Aubrey Mae Bonde

UNLV Theses, Dissertations, Professional Papers, and Capstones

Late Pleistocene megaherbivore communities of the Pacific and Mountain West states of California and Nevada are under-analyzed in regard to ecological function (diet, mobility, niche partitioning, and range of ecological tolerance). Stable isotope analysis is a powerful tool that is known to recover primary paleodiet and paleoenvironmental information from biogenic materials, such as enamel and dentin. This dissertation explores the use of carbon and oxygen stable isotopes in Late Pleistocene (40-10 Ka) megaherbivore teeth to gain a better understanding of inter- and intra-specific behavior and reconstruct Late Pleistocene ecosystems of California and Nevada. Radiocarbon dates exist for most of the …

Towards Space Based Verification Of Co2 Emissions From Strong Localized Sources: Fossil Fuel Power Plant Emissions As Seen By A Carbonsat Constellation, Voltaire A. Velazco, M Buchwitz, H Bovensmann, M Reuter, O Schneising, J P. Heymann, T Krings, K Gerilowski, John P. Burrows

Faculty of Science - Papers (Archive)

No abstract provided.

The Geological Context Of Middle Pleistocene Crater Lake Deposits And Fossil Birds At Ulupau Head, Oahu, Hawaiian Islands, Paul Hearty, H. F. James, S. L. Olson

Faculty of Science - Papers (Archive)

Pleistocene lake deposits in Ulupau Crater on Oahu contain the oldest fossil bird remains known from the Hawaiian Islands. These fossils are important for documenting evolutionary rates in a variety of lineages of land and water birds from the islands. Detailed stratigraphic analysis reveals distinct cycles of deposition involving interbedded lacustrine deposits, colluvium, and paleosols. The volcanic basin was breached by marine erosion prior to the peak of the last interglaciation (marine isotope stage or ‘MIS’ 5e) over 125 ky ago, constraining the age of the lake sediments to the middle Pleistocene. The three cycles of deposition (Units III, IV, …

Reconstructing Cetacean Brain Evolution Using Computed Tomography, Lori Marino, Mark D. Uhen, Nicholas D. Pyenson, Bruno Frohlich

Veterinary Science and Medicine Collection

Until recently, there have been relatively few studies of brain mass and morphology in fossil cetaceans (dolphins, whales, and porpoises) because of difficulty accessing the matrix that fills the endocranial cavity of fossil cetacean skulls. As a result, our knowledge about cetacean brain evolution has been quite limited. By applying the noninvasive technique of computed tomography (CT) to visualize, measure, and reconstruct the endocranial morphology of fossil cetacean skulls, we can gain vastly more information at an unprecedented rate about cetacean brain evolution. Here, we discuss our method and demonstrate it with several examples from our fossil cetacean database. This …

The Helioporacean Octocoral Epiphaxum, Recent And Fossil: A Monographic Iconography, Frederick M. Bayer

Studies in Tropical Oceanography

No abstract provided.

Back Reef And Fore Reef Analogs In The Pleistocene Of North Jamaica: Implications For Facies Recognition And Sediment Flux In Fossil Reefs, S. K. Boss, W. D. Liddell

W. David Liddell

No abstract provided.

Climatic Implications Of Barbados Coral Growth, Richard E. Dodge, Karl K. Turekian, J. Rimas Vaisnys

Marine & Environmental Sciences Faculty Proceedings, Presentations, Speeches, Lectures

Results from a coral growth band analysis utilizing samples of M. annularis from the Recent Barbados reef and from three fossil raised reefs (Barbados I, II, and III dated at 82,000, 105,000, and 125,000 yrs. B.P.) indicate that in the Barbados II collection both average band width and variability were lower than in the other samples. We suggest the climate during formation of the 105,000 yrs. B.P. reef was significantly different than that of the present.

Two New Species Of The Genus Cycadeoidea From The Lower Cretaceous Of Utah, H. Blaine Furniss

Theses and Dissertations

Cyadeoidea medullara and C. cleavelandii, two new species of the genus Cycadeoidea, have been recovered from the Lower Cretaceous Cedar Mountain Formation of Utah. They have been described in histologic detail and relationships with similar species are set forth. Monoecism and pollination mechanisms have been discussed in connection with C. medullara. This data contributes to our knowledge of the diverse composition of Lower Cretaceous floras in Utah.