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Articles 31 - 59 of 59
Full-Text Articles in Life Sciences
Faunistic Studies On The Species Pachybrachis Chevrolat And Cryptocephalus Geoffroy (Coleoptera, Chrysomelidae, Cryptocephalinae) Of Dedegöl Mountains (Isparta), Ali̇ Gök
Turkish Journal of Zoology
The material of the subfamily Cryptocephalinae collected from Dedegöl Mountains (Isparta) between 1996 and 1998 consisted of 21 species belonging to the genera Pachybrachis and Cryptocephalus. A diagnostic key and distribution records for the species are presented. The species except for Cryptocephalus bipunctatus (L.), C. turcicus Suffrian, C. connexus Olivier, Pachybrachis limbatus (Menetries), P. fimbriolatus (Suffrian) and P. tessellatus orientalis Weise are new for Isparta.
Range Extension Of Caridina Weberi Sumatrensis De Man, 1892 (Decapoda, Caridea, Atyidae) To Sindh Waters (Karachi, Pakistan), Quddusi̇ Bashi̇r Kazmi̇, Feroz Akhtar Siddiqui, Mohammad Afzal Kazmi̇
Range Extension Of Caridina Weberi Sumatrensis De Man, 1892 (Decapoda, Caridea, Atyidae) To Sindh Waters (Karachi, Pakistan), Quddusi̇ Bashi̇r Kazmi̇, Feroz Akhtar Siddiqui, Mohammad Afzal Kazmi̇
Turkish Journal of Zoology
A freshwater atyid Caridina weberi sumatrensis is being reported for the first time from Sindh waters. This report from Karachi is evidently at the margin of its range.
The Ephemeroptera (Insecta) Fauna Of Lake Ulubat Basin, Mustafa Tanatmiş
The Ephemeroptera (Insecta) Fauna Of Lake Ulubat Basin, Mustafa Tanatmiş
Turkish Journal of Zoology
In this study, 1812 Ephemeroptera nymphs were collected in order to determine the Ephemeroptera fauna of Lake Ulubat Basin between April 1996 and September 1999. The distribution of 38 species belonging to 22 genera and 10 families is presented. Among these, the genus Brachycercus Curtis, 1834 and the species Brachycercus harrisella Curtis, 1834 are recorded in Turkey for the first time. In addition, previous records on the distribution of all 38 species in Turkey are given.
The Species Of Tryphoninae (Hymenoptera: Ichneumonidae) In Turkish Thrace, Murat Yurtcan, Ahmet Beyarslan
The Species Of Tryphoninae (Hymenoptera: Ichneumonidae) In Turkish Thrace, Murat Yurtcan, Ahmet Beyarslan
Turkish Journal of Zoology
This study was carried out to investigate the species of the subfamily Tryphoninae (Hymenoptera: Ichneumonidae) in Turkish Thrace between 1992 and 1994. A total of 89 samples were collected from 42 localities and 26 species belonging to 11 genera were determined. Diagnostic keys to the species, subgenera and genera were prepared. In order to distinguish morphological characters that were mentioned in the keys, 31 original figures are given. Of these species, Phytodietus polyzonias (Foerster), Netelia (Prosthodocis) japonicus (Uchida), N. (Bessobates) cristatus (Thomson), N. (Netelia) ocellaris (Thomson), N. (Netelia) rufescens (Tosquinet), Neleges proditor (Gravenhorst), Dyspetes arrogator Heinrich, Ctenochira angulata (Thomson), Exyston …
Level Of Lactate Dehydrogenase (Ldh) In Resistant And Susceptible Strains Of Culicine Mosquitoes Of The Karachi Region After Treatment With Ddt, Malathion And Cyfluthrin, M. Arshad Azmi, Imtiaz Ahmed, Syed Naim-Ul-Hasan Naqvi, Kahkashan Akhtar
Level Of Lactate Dehydrogenase (Ldh) In Resistant And Susceptible Strains Of Culicine Mosquitoes Of The Karachi Region After Treatment With Ddt, Malathion And Cyfluthrin, M. Arshad Azmi, Imtiaz Ahmed, Syed Naim-Ul-Hasan Naqvi, Kahkashan Akhtar
Turkish Journal of Zoology
The effects of Malathion, Cyfluthrin, and DDT were observed on the activity of lactate dehydrogenase (LDH) in susceptible and resistant strains of culicine mosquitoes. Biochemical estimation (colorimetry) revealed a remarkable increase in the enzyme activity in Culex fatigans (L.Y. strain). In contrast, in Culex fatigans (G.I. strain) and Culex fatigans (K.U strain), inhibition of enzyme activity was found after treatment with pesticides.
Effects Of Some Antimicrobial Agents On The Total Protein Content Of The Endoparasitoid Pimpla Turionellae L. (Hymenoptera: Ichneumonidae), Kemal Büyükgüzel
Effects Of Some Antimicrobial Agents On The Total Protein Content Of The Endoparasitoid Pimpla Turionellae L. (Hymenoptera: Ichneumonidae), Kemal Büyükgüzel
Turkish Journal of Zoology
The effects of thirteen antimicrobial agents that have different structures and modes of action on the total protein content of pupae of the hymenopterous endoparasitoid, Pimpla turionellae L., were investigated by rearing the larvae aseptically on chemically defined synthetic diets. These effects varied according to the their kind and dietary levels. The protein content of the pupae was significantly increased by penicillin, streptomycin, rifampicin, tetracycline hydrochloride, lincomycin hydrochloride, methyl p-hydroxybenzoate, cycloheximide and sodium benzoate, while it was decreased by nystatin. The other tested antimicrobial agents had no significant effects on the total protein content of the insect. Depending on dietary …
Effects Of Some Dna Gyrase Inhibitors On The Survival And Development Of Pimpla Turionellae (Hymenoptera: Ichneumonidae) Larvae Reared On An Artificial Diet, Kemal Büyükgüzel
Effects Of Some Dna Gyrase Inhibitors On The Survival And Development Of Pimpla Turionellae (Hymenoptera: Ichneumonidae) Larvae Reared On An Artificial Diet, Kemal Büyükgüzel
Turkish Journal of Zoology
The effects of novobiocin, nalidixic and oxolinic acids, which are potent antibacterial agents, on the survival and development of the hymenopterous endoparasitoid Pimpla turionellae L. were investigated alone by rearing the larvae aseptically on chemically defined synthetic diets. The percentage of fifth instar larvae was not significantly affected by the diets with different levels of the antibiotics used in the study. A diet containing the lowest level of novobiocin significantly increased the yields of pupae and adults. This level also significantly shortened the developmental time for fifth instar, but had no significant effect on the complete development of the larvae …
Evaluation Of Methiocarb 50%-Wp As A Taste Repellent Against The House Sparrow (Passer Domesticus L.), Syed Waqar Ali̇ Ri̇zvi̇, Amjad Pervez, Syed Muzaffar Ahmed
Evaluation Of Methiocarb 50%-Wp As A Taste Repellent Against The House Sparrow (Passer Domesticus L.), Syed Waqar Ali̇ Ri̇zvi̇, Amjad Pervez, Syed Muzaffar Ahmed
Turkish Journal of Zoology
Laboratory studies in cages and outdoor aviaries were conducted, using different concentrations of methiocarb 50%-WP to estimate the ideal dose needed to produce taste aversion in the house sparrow (Passer domesticus). Under a nochoice test, sparrows showed a clear difference in millet bait preference among various concentrations. Under choice tests, in cages and outdoor aviaries a clear repellency trend was recorded in both sexes with increases in poison concentration. However, female birds showed a higher repellency pattern than males. It was concluded that methiocarb grain bait at 0.1% proved to be highly effective in repelling sparrows and may function as …
Behavioural Abnormalities Of Cyprinion Watsoni On Exposure To Copper And Zinc, Syed Lal Shah
Behavioural Abnormalities Of Cyprinion Watsoni On Exposure To Copper And Zinc, Syed Lal Shah
Turkish Journal of Zoology
Cyprinion watsoni were exposed to three treatments of copper (0.03, 0.06, 0.12 mg/L) and three treatments of zinc (0.5, 3.0, 7.0 mg/L) for a period of one week to allow behavioural changes to be observed. None of the treatments caused mortality. Treatment with 0.03 mg Cu/L caused little change in fish behaviour; however, 0.06 mg Cu/L caused increased swimming activity and breathing movements. The highest treatment of copper (0.12 mg Cu/L) caused lethargy and loss of equilibrium in C. watsoni. Similarly, the lowest concentration of zinc (0.5 mg/L) caused no visible change in fish behaviour; however, with 3.0 mg Zn/L, …
Detection Of Mutagenic-Carcinogenic Pollutants In Aquatic Systems Using Cytogenetic Methods In Fish, Mehmet Ulupinar, İbrahi̇m Okumuş
Detection Of Mutagenic-Carcinogenic Pollutants In Aquatic Systems Using Cytogenetic Methods In Fish, Mehmet Ulupinar, İbrahi̇m Okumuş
Turkish Journal of Zoology
In this study, the detection of mutagenic-carcinogenic pollutants in water by using cytogenetic methods in fish was examined along with the necessity of sister chromatid exchange (SCE), anaphase aberrations (AA) and micronucleus (MN) tests for chemical analysis in aquatic systems. It has been reported that central mudminnow (Umbra limi) appear to be the most suitable species for such analysis because of its large and fewer chromosomes (2n=22) and high cell division ratio. This species also has a wide distribution, and can be easily captured and held for study. In such analysis, intestines, stomach, kidney and gill tissues stand out as …
A Preliminary Study On The Population Dynamics Parameters Of Whiting (Merlangius Merlangus Euxinus) In Turkish Black Sea Coastal Waters, Ali̇ İşmen
Turkish Journal of Zoology
Some population parameters of the whiting (Merlangius merlangus euxinus) distributed along the Turkish Black Sea coast, were determined by using age-length data. The mean annual growth rate in length was found to be 3.7 cm. The maximum age group was determined as IX for females and VI for males. The seasonalized von Bertalanffy growth parameters in length were estimated to be L_o=39.1 cm, K = 0.15 year^{-1}, t_o = -1.53 year, C=0.23, t_s=0.48. There was a difference in growth rate between male and female fish. The females grow faster than males and reach a greater maximum length. Total mortality and …
The Effects Of Phytoplankton Concentration, Size Of Mussel And Water Temperature On Feed Consumption And Filtration Rate Of The Mediterranean Mussel (Mytilus Galloprovincialis Lmk), İbrahi̇m Okumuş, Nadi̇r Başçinar, Mükerrem Özkan
The Effects Of Phytoplankton Concentration, Size Of Mussel And Water Temperature On Feed Consumption And Filtration Rate Of The Mediterranean Mussel (Mytilus Galloprovincialis Lmk), İbrahi̇m Okumuş, Nadi̇r Başçinar, Mükerrem Özkan
Turkish Journal of Zoology
The effects of phytoplankton concentration, size and water temperature on the feeding rate of the Mediterranean mussel (Mytilus galloprovincialis) from the Black Sea were investigated in the laboratory within a static system. Overall consumption (total cell/h) was significantly higher at the highest phytoplankton concentrations than at the lowest concentration (p
A New Record For The Turkish Spider Fauna: Cyclosa Conica Pallas (Araneae, Araneidae), Abdullah Bayram, Murat Ünal
A New Record For The Turkish Spider Fauna: Cyclosa Conica Pallas (Araneae, Araneidae), Abdullah Bayram, Murat Ünal
Turkish Journal of Zoology
The characteristic features and drawings of Cyclosa conica Pallas, 1772, which is recorded for the first time in Turkey, are given in this study.
Sds-Page Patterns Of Blood Serum Proteins In Some Species Of The Genus Meriones (Mammalia: Rodentia), Reyhan Çolak, Nuri̇ Yi̇ği̇t, Ercüment Çolak, Rolf Gattermann, Karsten Neumann
Sds-Page Patterns Of Blood Serum Proteins In Some Species Of The Genus Meriones (Mammalia: Rodentia), Reyhan Çolak, Nuri̇ Yi̇ği̇t, Ercüment Çolak, Rolf Gattermann, Karsten Neumann
Turkish Journal of Zoology
The blood serum proteins of Meriones meridianus, Meriones crassus, Meriones persicus and Meriones tristrami, all found in Turkey, along with domestic Meriones unguiculatus, which originated in Mongolia, were examined for the first time by SDS-PAGE. In the globulin zone, seven to ten bands were identified, one band in the post-albumin and albumin zones and one or two bands in the pre-albumin zone of five species of the genus Meriones. There was no diagnostic difference in the pattern of serum protein bands between species.
Five New Chironomidae (Diptera) Species For The Turkish Fauna, Nurcan Özkan
Five New Chironomidae (Diptera) Species For The Turkish Fauna, Nurcan Özkan
Turkish Journal of Zoology
Five species were identified as new for the Turkish potamofauna of Edirne and Tekirdağ provinces from Thrace region between August 1995 and August 1996 in this study. They are Corynoneura scutellata Thienemann, 1944 (in Ergene River), Beckidia zabolotzkii (Goetghebuer, 1938) (in Meriç River), Robackia demeijerei (Kruseman, 1933) (in Meriç River and Sazlıdere Stream), Parachironomus arcuatus Goetghebuer 1919 (in Meriç River and Ergene River) and Parachironomus longiforceps Kruseman, 1933 (in Meriç River).
On The Distribution And Taxonomic Status Of Microtus Guentheri (Danford And Alston, 1880) And Microtus Lydius Blackler, 1916 (Mammalia: Rodentia) In Turkey, Nuri̇ Yi̇ği̇t, Ercüment Çolak
On The Distribution And Taxonomic Status Of Microtus Guentheri (Danford And Alston, 1880) And Microtus Lydius Blackler, 1916 (Mammalia: Rodentia) In Turkey, Nuri̇ Yi̇ği̇t, Ercüment Çolak
Turkish Journal of Zoology
Topotypes of Microtus guentheri were compared with specimens of the genus Microtus with 2n=54 chromosomes from western Anatolia. According to morphological and biometrical comparisons, it was determined that M. guentheri is a different species from Microtus lydius. The dorsal colour and the shape of the baculum distinguish M. guentheri from M. lydius morphologically. In addition, ear length, interorbital constriction, mastoid breadth, height of tympanic bulla and weight of M. guentheri were found to be statistically different from those of the nominate subspecies of M. lydius (p
The Effects Of Cimetidine And Ranitidine On Kidney Cortex Of Mus Musculus Albinus, Kadri̇ye Akgün Dar
The Effects Of Cimetidine And Ranitidine On Kidney Cortex Of Mus Musculus Albinus, Kadri̇ye Akgün Dar
Turkish Journal of Zoology
In this study, 4.3 µg/g cimetidine and 17 µg/g ranitidine were orally given to mice for two months. The effects of cimetidine and ranitidine on kidney cortical tissue were examined at light microscopical level. In both groups, it was detected that the tubules were separated by their lumen containing darkly stained remnant material. Vacuolization, destruction of apical membrane and decreased PAS positive reaction were observed in the apical membrane of proximal tubule epithelial cells. The nuclei of proximal and distal tubule cells were cylindrical. More excessively in the ranitidine treated group, widening of the tubule lumens and limited proximal tubule …
Muscle Fiber Types Of The Tunica Muscularis Externa At The Upper Part Of The Sparrow (Passer Domesticus) Esophagus, Fati̇me Geyi̇koğlu, Aysel Temelli̇, Abdullah Özkaral
Muscle Fiber Types Of The Tunica Muscularis Externa At The Upper Part Of The Sparrow (Passer Domesticus) Esophagus, Fati̇me Geyi̇koğlu, Aysel Temelli̇, Abdullah Özkaral
Turkish Journal of Zoology
Muscle fiber types of the tunica muscularis externa of the upper part of the sparrow esophagus were characterised by the method of myofibrillar adenosine triphosphatase (mATPase). Muscle fibers were described as type II and type III. Muscle fiber types were histochemically different from those of the esophagus of broiler ross. It is thought that this difference should be evaluated according to the functional difference between the esophagus of domestic and wild bird species.
Effect Of Temperature On Development, Sexual Maturation Time, Food Consumption And Body Weight Of Schistocerca Gregaria Forsk. (Orthoptera: Acrididae), N. Eylem Akman Gündüz, Adem Gülel
Effect Of Temperature On Development, Sexual Maturation Time, Food Consumption And Body Weight Of Schistocerca Gregaria Forsk. (Orthoptera: Acrididae), N. Eylem Akman Gündüz, Adem Gülel
Turkish Journal of Zoology
This study was carried out to determine the effect of temperature on the development, sexual maturation time, food consumption and body weight of S. gregaria, Forsk., at 25 and 30ºC. The insects were fed on fresh wheat sprouts (FWS) and bran (B). When the temperature was increased, both developmental time and attainment time to sexual maturation were shortened. Total nymphal developmental time at 25 and 30ºC was 32 and 22 days, respectively. Similarly, the average attainment time to sexual maturity was 33 days at 30ºC, whereas it was 52 days at 25ºC. The food consumption of the insects increased at …
Antimicrobial Agents: Their Combined Effects On Total Protein Content Of The Endoparasitoid Pimpla Turionellae L. (Hymenoptera: Ichneumonidae), Kemal Büyükgüzel
Antimicrobial Agents: Their Combined Effects On Total Protein Content Of The Endoparasitoid Pimpla Turionellae L. (Hymenoptera: Ichneumonidae), Kemal Büyükgüzel
Turkish Journal of Zoology
The effects of some antibiotics and antifungal agents in combination on the total protein content of young pupae of the hymenopterous endoparasitoid, Pimpla turionellae L., were investigated by rearing the larvae aseptically on chemically defined synthetic diets. These effects varied with the kinds and levels of the combination of antimicrobial agents in the diets. The combinations of antibiotics generally increased the total protein content according to their individual levels in the diets. Antifungal agent combinations caused a slight but significant increase in the total protein content. However, some tested combinations of nystatin with methyl p-hydroxybenzoate or cycloheximide significantly decreased the …
The Larval Development Of The Spider Crab Menaethiops Nodulosus (Nobili, 1905) (Crustacea, Decapoda, Majidae) Reared In The Laboratory, Farhana Shafiq Ghory, Feroz Akhtar Siddiqui
The Larval Development Of The Spider Crab Menaethiops Nodulosus (Nobili, 1905) (Crustacea, Decapoda, Majidae) Reared In The Laboratory, Farhana Shafiq Ghory, Feroz Akhtar Siddiqui
Turkish Journal of Zoology
The present paper is based on the larval development of a spider crab Menaethiops nodulosus (Nobili, 1905) reared in the laboratory. The genus Menaethiops is represented by twelve species from Indo-Pacific regions including M. bicornis Alcock, 1895. Later two more species M. gadaniensis Kazmi & Tirmizi and M. nodulosus (Nobili) were added to the marine fauna of Pakistan. The ovigerous female of M. nodulosus was collected from Buleji on October 28, 1996, prezoeae and first zoeae were obtained on November 13, 1996 and the first zoeae were segregated for further experiment. This species passes through two zoeal stages before moulting …
The Effects Of Acute Stress On Rainbow Trout (Oncorhynchus Mykiss), Ayşegül Kubi̇lay, Gülşen Uluköy
The Effects Of Acute Stress On Rainbow Trout (Oncorhynchus Mykiss), Ayşegül Kubi̇lay, Gülşen Uluköy
Turkish Journal of Zoology
The physiological effects of acute stressors (transport, handling, netting and confinement) on rainbow trout in an aquaculture system were investigated. Serum cortisol level, serum glucose and lysozyme activity were determined in rainbow trout stressed by acute stressors, and compared with those of unstressed (control) fish. Serum cortisol, glucose levels and lysozyme activity were significantly higher(P
The Ostracoda (Crustacea) Fauna Of Lakes Erikli, Hamam, Mert, Pedina And Saka (İğneada, Kırklareli,Turkey), Okan Köylüoğlu, Erdoğan Usta, M. Teki̇n Babaç
The Ostracoda (Crustacea) Fauna Of Lakes Erikli, Hamam, Mert, Pedina And Saka (İğneada, Kırklareli,Turkey), Okan Köylüoğlu, Erdoğan Usta, M. Teki̇n Babaç
Turkish Journal of Zoology
No abstract provided.
The Effects Of High Salinity On The Production Of Capoeta Tinca In A Naturally Contaminated River, Fi̇tnat Güler Ekmekçi̇
The Effects Of High Salinity On The Production Of Capoeta Tinca In A Naturally Contaminated River, Fi̇tnat Güler Ekmekçi̇
Turkish Journal of Zoology
The production properties of Capoeta tinca, the dominant fish population in a naturally contaminated stream in the Kızılırmak Basin, were studied together with the physical and chemical characteristics of the waters and evaluated in order to assess the impact of the high natural salinity on the fish population. The evaluation was based on comparisons of fish specimens caught from Acı stream, which is highly contaminated by a lithology of high salt content, and those from Terme stream, the uncontaminated freshwater branch, between May 1995 and May 1996. The salinity and electrical conductivity of Acı stream varied between 2 and 8.5, …
Relative Importance Of Food Items In Feeding Of Chondrostoma Regium Heckel, 1843, And Its Relation With The Time Of Annulus Formation, Aysun Gümüş, Mahmut Yilmaz, Nazmi̇ Polat
Relative Importance Of Food Items In Feeding Of Chondrostoma Regium Heckel, 1843, And Its Relation With The Time Of Annulus Formation, Aysun Gümüş, Mahmut Yilmaz, Nazmi̇ Polat
Turkish Journal of Zoology
In this study, we analyzed the relative importance of food items of Chondrostoma regium Heckel, 1843. A total of 145 individuals were sampled between April 1993 and March 1994 in Suat Uğurlu Dam Lake. Stomach content analysis was used to determine dietary composition. Relative measures of food item quantity were estimated by two different approaches: the Index of Relative Importance (IRI) and the Geometric Index of Importance (GII). The results from these indices were compared and the most reliable way of estimating dietary composition was discussed. Navicula, Cymbella and Synedra were the most frequently consumed organisms. As C. regium mostly …
Astioglossimetra Karachiensis N. Gen., N. Sp. (Trematoda: Plagiorchiidae: Astiotrematinae) From The Marine Turtle Chelonia Mydas Of Karachi Coast, Fatima Mujib Bilqees, Nasira Khatoon, Aly Khan
Astioglossimetra Karachiensis N. Gen., N. Sp. (Trematoda: Plagiorchiidae: Astiotrematinae) From The Marine Turtle Chelonia Mydas Of Karachi Coast, Fatima Mujib Bilqees, Nasira Khatoon, Aly Khan
Turkish Journal of Zoology
A new trematode genus Astioglossimetra n. gen. has been erected to accommodate an undescribed species A. karachiensis n. gen., n. sp., from Chelonia mydas of the Karachi coast. The genus name refers to its relation to the genera Astiotrema Looss and Glossimetra Mehra of the family Plagiorchiidae and subfamily Astiotrematinae.
Three Unrecorded Helophorus Fabricius, 1775 (Coleoptera, Hydrophilidae) Species For The Turkish Fauna, Ümi̇t İncekara, Abdullah Mart, Orhan Erman
Three Unrecorded Helophorus Fabricius, 1775 (Coleoptera, Hydrophilidae) Species For The Turkish Fauna, Ümi̇t İncekara, Abdullah Mart, Orhan Erman
Turkish Journal of Zoology
Three newly recorded Helophorus Fabricius, 1775 species for Turkish fauna, Helophorus (Rhopalhelophorus) fulgidicollis Motschulsky, 1860, H. (R.) similis Kuwert, 1887 and H. (R.) subarcuatus Rey, 1885, are compared against our specimens. Furthermore, their distribution in Turkey and around the world is presented.
Three New Rhizopulvinaria Species (Homoptera: Coccoidea: Coccidae) For Scale Insect Fauna Of Turkey, Mehmet Bora Kaydan, Ferenc Kozar, Bülent Yaşar
Three New Rhizopulvinaria Species (Homoptera: Coccoidea: Coccidae) For Scale Insect Fauna Of Turkey, Mehmet Bora Kaydan, Ferenc Kozar, Bülent Yaşar
Turkish Journal of Zoology
Three Rhizopulvinaria species, Rhizopulvinaria pyrethri Borchsenius, Rhizopulvinaria turkestanica (Archangelskaya), and Rhizopulvinaria viridis Borchsenius, were identified on wild flora in eastern Anatolia in 1997. All of them are new records for the Turkish scale insect fauna.
Micrommata Virescens (Clerck, 1757), A New Species For The Spider Fauna Of Turkey (Araneae, Sparassidae), Abdullah Bayram, Sevda Özdağ
Micrommata Virescens (Clerck, 1757), A New Species For The Spider Fauna Of Turkey (Araneae, Sparassidae), Abdullah Bayram, Sevda Özdağ
Turkish Journal of Zoology
The morphological characteristics of Micrommata virescens (Clerck, 1757), which is recorded for the first time in Turkey, are given.