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- Southern Ocean (3)
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Articles 1 - 26 of 26
Full-Text Articles in Physical Sciences and Mathematics
The Effect Of Submarine Canyon Width And Stratification On Coastal Circulation And Across Shelf Exchange, Kyung-Hoon Hyun
The Effect Of Submarine Canyon Width And Stratification On Coastal Circulation And Across Shelf Exchange, Kyung-Hoon Hyun
OES Theses and Dissertations
A 3-D ocean circulation model is used to investigate the effect of the width of a submarine canyon and stratification on adjacent coastal circulation and across shelf exchange. Upwelling winds for 20 d drive circulation over six canyons of varying widths (8–60 km), two escarpments to represent infinitely wide canyons, and a generic shelf-slope without a canyon. Also six stratifications whose first internal Rossby radii (a) range over 2–18 km were applied to two canyons (20 and 40 km) to see the effect of stratification. All simulations adjust to the canyon after 5 days spinup and reach an …
Bio-Optical Properties Of The Arctic Waters: Empirical And Theoretical Observations, Jian Wang
Bio-Optical Properties Of The Arctic Waters: Empirical And Theoretical Observations, Jian Wang
OES Theses and Dissertations
Bio-optical observations were made during August 2000 in the Beaufort and Chukchi Seas. Chlorophyll a concentration (Chl) ranged from 0.068 to 18.51 mg chl m−3. Both total particulate and phytoplankton absorption at 443 nm were closely correlated with chlorophyll concentration. There is no strong correlation between chlorophyll concentration and absorption by soluble materials or nonpigmented particulates. Absorption, scattering, and attenuation all show strong first-order spectral relationships. Two semianalytical remote sensing reflectance models were evaluated and validated using bio-optical data collected in this region. Both models were proficient at retrieving chlorophyll concentration, phytoplankton absorption coefficients, and particulate backscattering coefficients. …
Relating Water And Otolith Chemistry In Chesapeake Bay, And Their Potential To Identify Essential Seagrass Habitats For Juveniles Of An Estuarine-Dependent Fish, Spotted Seatrout (Cynoscion Nebulosus), Emmanis Dorval
Biological Sciences Theses & Dissertations
A quantitative understanding of habitat use of estuarine-dependent fishes is critical to the conservation of their most essential habitats. Because recruitment and fitness may be influenced by the quality of juvenile habitats, developing methods to quantify habitat-specific survivorship is pivotal to such understanding. An initial step to quantify survivorship is to validate the habitat-specific natural tags contained in otoliths. To this aim I investigated the variability in the chemistry of surface waters and otoliths of juvenile spotted seatrout, Cynoscion nebulosus, in five seagrass habitats of Chesapeake Bay, namely: Potomac, Rappahannock, York, Island, and Eastern Shore. I measured Mg, Ca, …
Numerical Study Of The Diapycnal Flow Through A Tidal Front With Passive Tracers, Chingming Dong, Robert Houghton, Hsien-Wang Ou, Dake Chen, Tal Ezer
Numerical Study Of The Diapycnal Flow Through A Tidal Front With Passive Tracers, Chingming Dong, Robert Houghton, Hsien-Wang Ou, Dake Chen, Tal Ezer
CCPO Publications
A two-dimensional numerical model is used to study the diapycnal flow through a tidal front with passive tracers. In a basic numerical experiment a passive tracer is released into the bottom water at the offshore edge of a tidal front, and it subsequently moves on-bank with a velocity that decreases with time. This qualitatively agrees with a recent field experiment using a dye tracer on Georges Bank. Additional experiments are performed to investigate the sensitivity of the tracer dispersion to the tidal phase and the location of tracer release within the front. As the release point is moved on-bank across …
Satellite Evidence Of Hurricane-Induced Phytoplankton Blooms In An Oceanic Desert, S. M. Babin, J. A. Carton, T. D. Dickey, J. D. Wiggert
Satellite Evidence Of Hurricane-Induced Phytoplankton Blooms In An Oceanic Desert, S. M. Babin, J. A. Carton, T. D. Dickey, J. D. Wiggert
CCPO Publications
The physical effects of hurricanes include deepening of the mixed layer and decreasing of the sea surface temperature in response to entrainment, curl-induced upwelling, and increased upper ocean cooling. However, the biological effects of hurricanes remain relatively unexplored. In this paper, we examine the passages of 13 hurricanes through the Sargasso Sea region of the North Atlantic during the years 1998 through 2001. Remotely sensed ocean color shows increased concentrations of surface chlorophyll within the cool wakes of the hurricanes, apparently in response to the injection of nutrients and/or biogenic pigments into the oligotrophic surface waters. This increase in post-storm …
Lagrangian Modelling Studies Of Antarctic Krill (Euphausia Superba) Swarm Formation, Eileen E. Hofmann, A. G. Edward Haskell, John M. Klinck, Cathy M. Lascara
Lagrangian Modelling Studies Of Antarctic Krill (Euphausia Superba) Swarm Formation, Eileen E. Hofmann, A. G. Edward Haskell, John M. Klinck, Cathy M. Lascara
CCPO Publications
A two-dimensional Lagrangian particle model was developed to examine the spatial distribution of Antarctic krill (Euphausia superba). The time-dependent location of particles, which represent krill individuals, is determined by random diffusion, foraging activity, and movement induced by the presence of neighbours. Foraging activity is based on prescribed food conditions and is such that krill swim slower and turn more frequently in areas of high food concentration. The presence or absence of neighbours either disperses krill, if the local concentrations become too dense, or coalesces krill, if concentrations become too dilute, respectively. Predation on krill is included and affects …
A Modelling Study Of The Influence Of Environment And Food Supply On Survival Of Crassostrea Gigas Larvae, Eileen E. Hofmann, Eric N. Powell, Eleanor A. Bochenek, John M. Klinck
A Modelling Study Of The Influence Of Environment And Food Supply On Survival Of Crassostrea Gigas Larvae, Eileen E. Hofmann, Eric N. Powell, Eleanor A. Bochenek, John M. Klinck
CCPO Publications
A biochemically based model was developed to simulate the growth, development, and metamorphosis of larvae of the Pacific oyster (Crassostrea gigas). The unique characteristics of the model are that it: (1) defines larvae in terms of their protein, neutral lipid, polar lipid, carbohydrate, and ash content; (2) tracks weight separately from length to follow larval condition; and (3) includes genetic variation in growth efficiency and egg quality to better simulate cohort population dynamics. The model includes parameterizations for filtration, ingestion, and respiration, which determine larval growth rate, and processes controlling larval mortality and metamorphosis. Changes in larval tissue …
On The Sensitivity Of The West Caribbean Sea Circulation To Tides, Wind, And Mesoscale Ocean Eddies: A Three-Dimensional Ocean Model Study, Deeptha V. Thattai, Tal Ezer, Bjorn Kjerfve
On The Sensitivity Of The West Caribbean Sea Circulation To Tides, Wind, And Mesoscale Ocean Eddies: A Three-Dimensional Ocean Model Study, Deeptha V. Thattai, Tal Ezer, Bjorn Kjerfve
CCPO Publications
A three-dimensional, primitive equation ocean model is used to study the circulation in the West Caribbean Sea (WCS) region, and to test the sensitivity of the coastal flow to various forcing fields such as tides, climatological wind, and Caribbean eddies. The model domain is bordered by latitudes 15 – 22 degrees N and longitudes 76 – 87 degrees W, with the MesoAmerican Barrier Reef System (MBRS, along the coasts of Mexico, Belize, Guatemala, and Honduras) and the southern coast of Cuba as land boundaries. The WCS is open to the Caribbean Sea in the southeast and the Yucatan Channel in …
Black Carbon In Estuarine And Coastal Ocean Dissolved Organic Matter, Antonio Mannino, H. Rodger Harvey
Black Carbon In Estuarine And Coastal Ocean Dissolved Organic Matter, Antonio Mannino, H. Rodger Harvey
OES Faculty Publications
We measured black carbon (BC) in ultrafiltered, high-molecular weight dissolved organic matter (UDOM) in surface waters of Delaware Bay, Chesapeake Bay, and the adjacent Atlantic Ocean (U.S.A.) to investigate the importance of riverine and estuarine dissolved organic matter (DOM) as a source of BC to the ocean. BC was 5-72% of UDOM-C (27 ± 17%), which corresponds to 8.9 ± 6.5% of dissolved organic carbon (DOC), with higher values in the heavily urbanized midbay region of the Delaware Estuary and lower yields in the river and coastal ocean. The spatial and seasonal distributions of BC along the salinity gradient of …
Dinitrogen Fixation And Release Of Ammonium And Dissolved Organic Nitrogen By Trichodesmium Ims101, Margaret R. Mulholland, Deborah A. Bronk, Douglas G. Capone
Dinitrogen Fixation And Release Of Ammonium And Dissolved Organic Nitrogen By Trichodesmium Ims101, Margaret R. Mulholland, Deborah A. Bronk, Douglas G. Capone
OES Faculty Publications
Two methods used to measure dinitrogen (N2) fixation (acetylene reduction and 15N2 uptake) often result in different N2 fixation rates. Part of the discrepancy may arise from the observation that Trichodesmium can release a fraction of their recently fixed N2 as dissolved organic nitrogen (DON) and/or ammonium (NH4 +). To resolve outstanding issues regarding N2 fixation and the production of dissolved combined nitrogen (N) by Trichodesmium, we conducted a comprehensive analysis of N2 fixation and the production of DON and NH4+ in cultures of Trichodesmium IMS101. We performed …
Age And Growth Of Scotia Sea Icefish, Chaenocephalus Aceratus, From The South Shetland Islands, M. La Mesa, J. Ashford, E. Larson, M. Vacchi
Age And Growth Of Scotia Sea Icefish, Chaenocephalus Aceratus, From The South Shetland Islands, M. La Mesa, J. Ashford, E. Larson, M. Vacchi
OES Faculty Publications
Samples of Chaenocephalus aceratus (Lönnberg) were collected during a trawl survey carried out around the South Shetland Islands in January–February 2002. Fish were caught by commercial bottom trawl fishing down to 500 m depth, using a stratified randomized sampling design. As observed in other recent surveys within the same area, C. aceratus represented one of the predominant species. Overall, 357 specimens ranging from 13 and 67 cm (TL) were selected for the present study. Ages were estimated by counting annuli present in the sagittal otoliths, exposed by grinding and polishing along their sagittal plane. To estimate the precision of age …
Sources And Cycling Of Carbonyl Sulfide In The Sargasso Sea, Gregory A. Cutter, Lynda S. Cutter, Katherine C. Filippino
Sources And Cycling Of Carbonyl Sulfide In The Sargasso Sea, Gregory A. Cutter, Lynda S. Cutter, Katherine C. Filippino
OES Faculty Publications
The cycling of the radiatively important gas carbonyl sulfide (OCS) was studied in surface waters of the Sargasso Sea. In August 1999, surface OCS concentrations averaged 8.6 pmol L-1, showed minor diel variations, and varied little with depth. An OCS precursor, total dissolved organic sulfur (DOS), was lowest at the surface (40 nmol L-1) and increased with depth. The photoproduction rate of OCS from in situ incubations averaged 9.6 pmol L-1 h-1, whereas dark production was 7.0 pmol L-1 h-1. Apparent quantum yields were 10-5-10-7 from 313-436 …
Influence Of Short-Term Variations In Food On Survival Of Crassotrea Gigas Larvae: A Modeling Study, Eric N. Powell, Eleanor A, Bochenek, John M. Klinck, Eileen E. Hofmann
Influence Of Short-Term Variations In Food On Survival Of Crassotrea Gigas Larvae: A Modeling Study, Eric N. Powell, Eleanor A, Bochenek, John M. Klinck, Eileen E. Hofmann
OES Faculty Publications
A biochemically-based model was developed to simulate the growth, development, and metamorphosis of larvae of the Pacific oyster, Crassostrea gigas. The model defines larvae in terms of their protein, lipid, carbohydrate, and ash content and includes variation in growth efficiency and egg quality to better simulate cohort population dynamics. Changes in tissue composition occur as the larva grows and in response to the biochemical composition of the food. The premise behind this modeling study was that certain periods of larval life are more critical than others with respect to the availability of food and that food quality is as …
A Holocene Record Of Changing Arctic Ocean Ice Drift Analogous To The Effects Of The Arctic Oscillation, Dennis A. Darby, Jens F. Bischof
A Holocene Record Of Changing Arctic Ocean Ice Drift Analogous To The Effects Of The Arctic Oscillation, Dennis A. Darby, Jens F. Bischof
OES Faculty Publications
The Arctic Oscillation (AO) controls the configuration of the Transpolar Drift (TPD). If thicker ice from the Beaufort Gyre were exported, the volume of fresh water/sea ice in the Nordic seas would significantly increase, decreasing the formation of North Atlantic deep water. This would cool large parts of the Northern Hemisphere and affect global climate. Therefore, in order to understand how the global climate system functions, it is imperative to know how the TPD changed over the last millennium or more. The provenance of grains in a sediment core located near the confluence of the TPD and the Beaufort Gyre …
Physical Forcing Of Phytoplankton Community Structure And Primary Production In Continental Shelf Waters Of The Western Antarctic Peninsula, Barbara B. Prézelin, Eileen E. Hofmann, Mark Moline, John M. Klinck
Physical Forcing Of Phytoplankton Community Structure And Primary Production In Continental Shelf Waters Of The Western Antarctic Peninsula, Barbara B. Prézelin, Eileen E. Hofmann, Mark Moline, John M. Klinck
OES Faculty Publications
Analyses of a multidisciplinary data set, collected in continental shelf waters of the Western Antarctic Peninsula (WAP) during austral summer of January 1993, identified a previously unrecognized forcing mechanism that sets up a physical and chemical structure that supports and assures site-specific diatom-dominated communities and enhanced biological production (Prézelin et al., 2000). This forcing is active when the southern boundary of the Antarctic Circumpolar Current (ACC) flows along the shelf edge, thereby facilitating onshelf bottom intrusions of nutrient-rich Upper Circumpolar Deep Water (UCDW), which then is upwelled or mixed into the upper water column. At times or locations where UCDW …
Light Dependence Of Selenium Uptake By Phytoplankton And Implications For Predicting Selenium Incorporation Into Food Webs, Stephen B. Baines, Nicholas S. Fisher, Martina A. Doblin, Gregory A. Cutter, Lynda S. Cutter, Brian Cole
Light Dependence Of Selenium Uptake By Phytoplankton And Implications For Predicting Selenium Incorporation Into Food Webs, Stephen B. Baines, Nicholas S. Fisher, Martina A. Doblin, Gregory A. Cutter, Lynda S. Cutter, Brian Cole
OES Faculty Publications
The potentially toxic element selenium is first concentrated from solution to a large but highly variable degree by algae and bacteria before being passed on to consumers. The large loads of abiotic and detrital suspended particles often present in rivers and estuaries may obscure spatial and temporal patterns in Se concentrations at the base of the food web. We used radiotracers to estimate uptake of both selenite (Se(IV)) and C by intact plankton communities at two sites in the Sacramento/San Joaquin River Delta. Our goals were to determine (1) whether C and Se(IV) uptake were coupled, (2) the role of …
Analytical Intercomparison Between Flow Injection-Chemiluminescence And Flow Injection-Spectrophotometry For The Determination Of Picomolar Concentrations Of Iron In Seawater, Andrew R. Bowie, Peter N. Sedwick, Paul J. Worsfold
Analytical Intercomparison Between Flow Injection-Chemiluminescence And Flow Injection-Spectrophotometry For The Determination Of Picomolar Concentrations Of Iron In Seawater, Andrew R. Bowie, Peter N. Sedwick, Paul J. Worsfold
OES Faculty Publications
A lab- and ship-based analytical intercomparison of two flow injection methods for the determination of iron in seawater was conducted, using three different sets of seawater samples collected from the Southern Ocean and South Atlantic. In one exercise, iron was determined in three different size-fractions (< 0.03 &μm, < 0.4 μm, and unfiltered) in an effort to better characterize the operational nature of each analytical technique with respect to filter size. Measured Fe concentrations were in the range 0.19 to 1.19 nM using flow injection with luminol chemiluminescence detection (FI-CL), and 0.07 to 1.54 nM using flow injection with catalytic spectrophotometric detection with N, N-dimethyl-p-phenylenediamine dihydrochloride (FI-DPD). The arithmetic mean for the FI-CL method was higher (by 0.09 nM) than the FI-DPD method for dissolved (< 0.4 μm) Fe, a difference that is comparable to the analytical blanks, which were as high as 0.13 nM ( CL) and 0.09 nM (DPD). There was generally good agreement between the FI-CL determinations for the < 0.03 μm size fraction and the FI-DPD determinations for the < 0.4 μm size fraction in freshly collected samples. Differences in total-dissolvable ( unfiltered) Fe concentrations determined by the two FI methods were more variable, reflecting the added complexity associated with the analysis of partially digested particulate material in these samples. Overall, however, the FI-CL determinations were significantly (P = 0.05) lower than the FI-DPD determinations for the unfiltered samples. Our results suggest that the observed, systematic inter-method differences reflect measurement of different physicochemical fractions of Fe present in seawater, such that colloidal and/or organic iron species are better determined by the FI-CL method than the FI-DPD method. This idea is supported by our observation that inter-method differences were largest for freshly collected acidified seawater, which suggests extended storage (>6 months) of acidified samples as a possible protocol for the determination of dissolved iron in seawater.
Interactive Regulation Of Dissolved Copper Toxicity By An Estuarine Microbial Community, Christina L. Dryden, Andrew S. Gordon, John R. Donat
Interactive Regulation Of Dissolved Copper Toxicity By An Estuarine Microbial Community, Christina L. Dryden, Andrew S. Gordon, John R. Donat
OES Faculty Publications
Cultured marine microorganisms under copper stress produce extracellular compounds having a high affinity for copper (copper-complexing ligands). These ligands are similar in binding strength to those found in natural waters, but few studies have examined the relationship between copper, copper-complexing ligand concentrations, and natural microbial populations. A series of in situ experiments in the Elizabeth River, Virginia, revealed that an intact estuarine microbial community responded to copper stress by production of extracellular, high-affinity copper-complexing ligands. The rate of ligand production was dependent on copper concentration and resulted in a reduction of the concentration of free cupric ions, Cu2+, …
Modeled And Observed Empirical Orthogonal Functions Of Currents In The Yucatan Channel, Gulf Of Mexico, Lie-Yauw Oey, Tal Ezer, Wilton Sturges
Modeled And Observed Empirical Orthogonal Functions Of Currents In The Yucatan Channel, Gulf Of Mexico, Lie-Yauw Oey, Tal Ezer, Wilton Sturges
CCPO Publications
Candela et al. [2003] have reported empirical orthogonal function (EOF) analyses based on 23-month current-meter and acoustic Doppler current profiler measurements in the Yucatan Channel. Those authors noted the difference between EOFs obtained from observations and their z-level models and EOFs calculated by Ezer et al. [2003] from the results of a terrain-following model. Here a new analysis is reported that explains this difference, and that also suggests the importance of shelf-edge meander mode of the core Loop Current in the channel. We show that the terrain-following model gives EOFs with characteristics similar to those observed when data from the …
Relevant Scales In Zooplankton Ecology: Distribution, Feeding, And Reproduction Of The Copepod Acartia Hudsonica In Response To Thin Layers Of The Diatom Skeletonema Costatum, Alexander B. Bochdansky, Stephen M. Bollens
Relevant Scales In Zooplankton Ecology: Distribution, Feeding, And Reproduction Of The Copepod Acartia Hudsonica In Response To Thin Layers Of The Diatom Skeletonema Costatum, Alexander B. Bochdansky, Stephen M. Bollens
OES Faculty Publications
We investigated the interaction of the copepod Acartia hudsonica in relation to thin layers of the diatom Skeletonema costatum. Thin layers have recently received much attention, since they are common and persistent features in the water column, often overlooked by traditional sampling methods. Their frequent abundance in coastal oceans and the high biomass associated with them has led to the assumption that they are important grazing sites of calanoid copepods. We employed 2-m tall tower tanks that allowed us to simulate thin layers. Three variables representative of three time scales were considered: the distribution of copepods in the tanks …
Estimation Of Drag Coefficient In James River Estuary Using Tidal Velocity Data From A Vessel-Towed Adcp, Chunyan Li, Arnoldo Valle-Levinson, Larry P. Atkinson, Kuo Chuin Wong, Kamazima M. M. Lwiza
Estimation Of Drag Coefficient In James River Estuary Using Tidal Velocity Data From A Vessel-Towed Adcp, Chunyan Li, Arnoldo Valle-Levinson, Larry P. Atkinson, Kuo Chuin Wong, Kamazima M. M. Lwiza
CCPO Publications
[1] A phase-matching method is introduced to calculate the bottom drag coefficient in tidal channels with significant lateral variation of depth. The method is based on the fact that the bottom friction in a tidal channel causes tidal velocity to have a phase difference across the channel. The calculation involves a few steps. First, the observed horizontal velocity components are analyzed to obtain the amplitude and phase of the velocity at the major tidal frequency. The phase of the longitudinal velocity is then fitted to a relationship derived from the linearized momentum balance. The drag coefficient is then calculated. This …
Advection, Krill, And Antarctic Marine Ecosystems, Eileen E. Hofmann, Eugene J. Murphy
Advection, Krill, And Antarctic Marine Ecosystems, Eileen E. Hofmann, Eugene J. Murphy
CCPO Publications
Advective processes are recognized as being important in structuring and maintaining marine ecosystems. In the Southern Ocean advective effects are perhaps most clearly observed because the Antarctic Circumpolar Current (ACC) provides a connection between most parts of the system, including open ocean and continental shelf regions. The ACC also provides a mechanism for large-scale transport of plankton, such as Antarctic krill (Euphousia superba Dana), which is an important component of the Southern Ocean food web. This overview provides a summary of recent observational and modelling results that consider the importance of advection to the Southern Ocean ecosystem and, in particular, …
Inner Shelf Circulation In Coastal Virginia: A Data Assimilation Approach, Hector Hito Sepulveda
Inner Shelf Circulation In Coastal Virginia: A Data Assimilation Approach, Hector Hito Sepulveda
OES Theses and Dissertations
The primary objective of this dissertation is to describe the tidal and subtidal flow patterns over the inner shelf of the Delmarva Peninsula, located in the Mid-Atlantic Bight of the United States (36.6–38.0 N), north of the Chesapeake Bay. The objective is pursued with a combination of direct measurements and numerical assimilative techniques. The dynamic balance of the study area is little known, and the distribution of tidal properties has not been described for this area since very rough descriptions in the 1950's. Hydrographic and current velocity profiles from four regional cruises in the inner shelf were used to study …
Inter-Annual To Inter-Decadal Variability Of Upwelling And Anchovy Population Off Northern Chile, Jose L. Blanco-Garcia
Inter-Annual To Inter-Decadal Variability Of Upwelling And Anchovy Population Off Northern Chile, Jose L. Blanco-Garcia
OES Theses and Dissertations
The coastal ocean of northern Chile has persistent wind-driven upwelling that produces high nutrient and chlorophyll concentrations in a narrow band along the coast. The objective of this thesis is to study the low frequency temporal variability of the upwelling system, to understand the spatial and temporal changes in the wind field, and how these changes may affect the upwelling and anchovy variability. Data used in this thesis includes time series of wind, sea level, sea surface temperature, and atmospheric pressure at coastal stations, from 1960 to 2003, and oceanographic and acoustic cruises for the period 1993–2003. The time series …
Estimation Of Primary Production And Carbon Flux In Antarctic Coastal Waters: A Modeling Study, Hae-Cheol Kim
Estimation Of Primary Production And Carbon Flux In Antarctic Coastal Waters: A Modeling Study, Hae-Cheol Kim
OES Theses and Dissertations
This study presents results from models that are designed to simulate the underwater light field, to simulate phytoplankton primary production, and to estimate the fate of phytoplankton carbon in continental shelf waters of the west Antarctic Peninsula (WAP) and Ross Sea. Simulation of the underwater light field required derivation of new coefficient sets for power function-type cloud cover correction algorithms, which were found to be influenced by multiple reflections between the bottom of clouds and the surface. The coefficient sets indicate that the spectral effect of clouds on the properties of the surface irradiance was spectrally-neutral for wavelengths greater than …
An Investigation Of Dissolved Organic Matter In A Shallow Coastal Bay Subject To Aureococcus Anophagefferens Blooms, Jean-Paul Simjouw
An Investigation Of Dissolved Organic Matter In A Shallow Coastal Bay Subject To Aureococcus Anophagefferens Blooms, Jean-Paul Simjouw
OES Theses and Dissertations
Aureococcus anophagefferens, the pelagophyte responsible for brown tide blooms, was identified in Chincoteague Bay in 1997 and has “bloomed” there since at least 1998. Aureococcus anophagefferens is capable of using dissolved organic nitrogen (DON) and dissolved organic carbon (DOC) substrates to support growth, and this utilization is hypothesized to give the organism a competitive advantage relative to other phytoplankton when inorganic nutrient concentrations are low or depleted. Because previous studies suggest dissolved organic matter (DOM) is important in initiating and sustaining brown tide blooms, a field study of the variations in DOC concentration and DOM composition was performed at …