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Articles 1 - 30 of 63
Full-Text Articles in Physical Sciences and Mathematics
Grammar And Variation: Understanding How Cis-Regulatory Information Is Encoded In Mammalian Genomes, Dana Michele King
Grammar And Variation: Understanding How Cis-Regulatory Information Is Encoded In Mammalian Genomes, Dana Michele King
Arts & Sciences Electronic Theses and Dissertations
Understanding how genotype leads to phenotype is key to understand both the development and dysfunction of complex organisms. In the context of regulating the gene expression patterns that contribute to cell identity and function, the goal of my thesis research is to how changes in genome sequence may impact impact gene expression by determining how sequence features contribute to regulatory potential. To accomplish this goal, I first leveraged the key regulatory role of pluripotency transcription factors (TFs) in mouse embryonic stem cells (mESCs) and tested synthetically generated and genomic identified combinations of binding site for four TFs, OCT4, SOX2, KLF4, …
Climatic Range Filling Of North American Trees, Benjamin Seliger
Climatic Range Filling Of North American Trees, Benjamin Seliger
Electronic Theses and Dissertations
Understanding the degree to which species distributions are controlled by climate is crucial for forecasting biodiversity responses to climate change. Climatic equilibrium, when species are found in all places which are climatically suitable, is a fundamental assumption of species distribution models, but there is evidence in support of climate disequilibria in species ranges. Long-lived, sessile organisms such as trees may be especially vulnerable to being outpaced by climate change, and thus prone to disequilibrium. In this dissertation, I tested the degree to which North American trees are in equilibrium with their potential climatic ranges using the ‘range filling’ metric, which …
River Bank Inducement Influence On A Shallow Groundwater Microbial Community And Its Effects On Aquifer Reactivity, Natalie June Gayner
River Bank Inducement Influence On A Shallow Groundwater Microbial Community And Its Effects On Aquifer Reactivity, Natalie June Gayner
Theses and Dissertations
Placing groundwater wells next to riverbanks to draw in surface water, known as riverbank inducement (RBI), is common and proposed as a promising and sustainable practice for municipal and public water production across the globe. However, these systems require further investigation to determine risks associated with river infiltration especially with rivers containing wastewater treatment plant (WWTP) effluent. Since microbes drive biogeochemical transformations in groundwater and largely affect water quality, it is important to understand how the microbial communities in drinking water wells are affected by river infiltration. This study investigated if, and to what extent, the microbial community in a …
Effect Of Macrograzers (Campostoma Spp. And Faxonius Spp.) On Periphyton In Ozark Streams With Considerations Given To Macrograzer Biomass, Phosphorus, And Season: Mensurative And Manipulative Studies, Kayla R. Sayre
Graduate Theses and Dissertations
Nutrient and benthic algal biomass relationships can guide numeric nutrient criteria development in lotic systems. However, herbivorous macrograzers may confound this relationship by reducing the slope of the positive relationship between nutrients and periphyton biomass in streams. I conducted a mensurative field study to determine if stoneroller and crayfish abundance related to algal biomass at varying nutrient concentrations and manipulated macrograzer presence with electrical exclosures in streams to examine macrograzer effects on algal biomass and understand whether these effects on periphyton varied with total phosphorus (TP) or season. Macrograzer density was quantified across a TP gradient (n=15 streams; range = …
Integrating Towed Underwater Video With Multibeam Acoustics For Mapping Benthic Habitat And Assessing Reef Fish Communities On The West Florida Shelf, Alexander Ross Ilich
Integrating Towed Underwater Video With Multibeam Acoustics For Mapping Benthic Habitat And Assessing Reef Fish Communities On The West Florida Shelf, Alexander Ross Ilich
USF Tampa Graduate Theses and Dissertations
Using a towed underwater video camera system, benthic habitats were classified along transects in a popular offshore fishing area on the West Florida Shelf (WFS) known as “The Elbow.” Additionally, high resolution multibeam bathymetry and co-registered backscatter data were collected for the entire study area. Using these data, full coverage geologic and biotic habitat maps were developed using both unsupervised and supervised statistical classification methodologies. The unsupervised methodology used was k-means clustering, and the supervised methodology used a random forest algorithm. The two methods produced broadly similar results; however, the supervised methodology outperformed the unsupervised methodology. The results of the …
Habitat Preferences Of Blue Marlin (Makaira Nigricans) And Black Marlin (Istiompax Indica) In The Eastern Pacific Ocean, Nima Farchadi, Michael G. Hinton, Andrew R. Thompson, Zhi-Yong Yin
Habitat Preferences Of Blue Marlin (Makaira Nigricans) And Black Marlin (Istiompax Indica) In The Eastern Pacific Ocean, Nima Farchadi, Michael G. Hinton, Andrew R. Thompson, Zhi-Yong Yin
Theses
Overexploitation and climate change can reduce the abundance and shift the spatial distribution of marine species. Determining the habitat suitability of a mobile pelagic species, such as Makaira nigricans (BUM) and Istiompax indica (BLM), can help describe their spatiotemporal distribution patterns over a broad spatial scale, which is a crucial need for fisheries management. Using 14 years (1997-2010) of Inter-American Tropical Tuna Commission (IATTC) catch data from purse-seine vessels in the eastern Pacific Ocean (EPO), we modeled the dynamic habitat suitability of BUM and BLM in response to environmental variables within the EPO using a species distribution model (MaxEnt) with …
Ecology And Population Dynamics Of Salmonids In The Columbia River: Response Of Fishes To Anthropogenic Change In A Large Riverscape, Timothy Sean Elder
Ecology And Population Dynamics Of Salmonids In The Columbia River: Response Of Fishes To Anthropogenic Change In A Large Riverscape, Timothy Sean Elder
Dissertations and Theses
Freshwater ecosystems and the species that reside therein are disproportionately imperiled compared to terrestrial systems. Over the past 150 years, the Columbia River basin in the western United States has gone from one of the most productive and abundant salmon watersheds in the world, to having just a small fraction of its former salmon abundance. The cause of declines in salmon productivity and abundance are related to overlapping and confounding stressors including changes in large-scale climatic patterns and anthropogenic alterations within and adjacent to the Columbia River. Four main anthropogenic stressors have been identified as the leading causes of salmonid …
Investigation Of Biogeochemical Mechanisms Of Greenhouse Gas Production In The Urban Hudson River Estuary, Brian Alan Brigham
Investigation Of Biogeochemical Mechanisms Of Greenhouse Gas Production In The Urban Hudson River Estuary, Brian Alan Brigham
Dissertations, Theses, and Capstone Projects
Coastal megacities deposit significant amounts of carbon (C), nitrogen (N) and other pollutants into surrounding waters. These inputs, including wastewater and surface water runoff, may affect estuarine and adjacent wetland biogeochemical cycles, microbial production and ultimately greenhouse gas (GHG) efflux. In many megacities pollutant loading is typically greatest after periods of precipitation when the volume of wastewater and storm water runoff exceeds local sanitation capacity, resulting in the discharge of raw sewage into adjacent waters. These combined sewage overflow (CSO) events have received considerable attention primarily due to their potential impact on human health and eutrophication. However, whether these events …
Plant Stimuli-Responsive Biodegradable Polymers For The Use In Timed Release Fertilizer Coatings, Spencer Heuchan
Plant Stimuli-Responsive Biodegradable Polymers For The Use In Timed Release Fertilizer Coatings, Spencer Heuchan
Electronic Thesis and Dissertation Repository
The use of nitrogen-based fertilizers continues to accelerate with human population growth and increases in global food requirements. Enhanced efficiency fertilizers (EEFs) have been developed to improve the synchronization between nutrient supply and crop nutrient demand. However, many of the current controlled release fertilizers are coated with non-degradable polymers that contribute to accumulation of microplastics within ecosystems. This thesis describes research towards the development of a new class of fertilizer coatings using a self-immolative polymer known as poly (ethyl glyoxylate) (PEtG). PEtG itself does not have suitable properties to produce a viable coating but once blended with another degradable polyester …
Nitrogen Cycling During A Period Of Environmental Change, Kaizad Patel
Nitrogen Cycling During A Period Of Environmental Change, Kaizad Patel
Electronic Theses and Dissertations
Natural and anthropogenic disturbances have altered the N cycle on the regional to global scale. Although numerous studies have examined the effects of chronic atmospheric N deposition on ecosystem processes, less is known about intra-annual variability of these processes, or how these responses might be altered under conditions of changing climate. The work described in this dissertation focuses on the effects of a changing chemical and physical climate, particularly terrestrial N processes (a) during Maine’s changing winters, and (b) in response to chronic elevated N additions.
Subnivean winter soil processes were examined at the Dwight B. DeMerritt Forest (DBDF) and …
Preventing Extinction Of At-Risk Plant Species In A Complex World, Holly Lee Bernardo
Preventing Extinction Of At-Risk Plant Species In A Complex World, Holly Lee Bernardo
Arts & Sciences Electronic Theses and Dissertations
Earthճ current biodiversity crisis is now considered a true mass extinction event, with species level extinction rates well above background rates and population level extinction rates orders of magnitude more common that species extinctions. There are many threats driving this loss of biodiversity. How each threat impacts the viability of a species is highly context dependent, but all are anthropogenic in origin and so as the human population continues to increase, so too will the pressure of these threats on our natural systems. Ultimately, how much a threat decreases the viability of a species depends on how that threat influences …
Analysis Of Temperature And Salinity Effects On Growth And Mortality Of Oysters (Crassostrea Virginica) In Louisiana, Troy Sehlinger
Analysis Of Temperature And Salinity Effects On Growth And Mortality Of Oysters (Crassostrea Virginica) In Louisiana, Troy Sehlinger
University of New Orleans Theses and Dissertations
Salinity (S) and temperature (T) control every facet of the eastern oyster (Crassostrea virginica) life cycle, principally reproduction, development, growth, and mortality. Previous studies conducted in in the Breton Sound (BR) and Barataria (BA) estuaries have reported differences in growth and mortality rates between the basins. In the present study, environmental conditions were synchronized to compare growth and mortality rates between basins at similar combinations of T and S. Results indicate that when T and S are the same (synchronized), seasonal oyster growth and mortality rates differ between BR and BA. Seasonal analyses revealed that as salinities increased …
Bubble Stream Production By Belugas (Delphinapterus Leucas), Megan Slack
Bubble Stream Production By Belugas (Delphinapterus Leucas), Megan Slack
Theses
Bubble stream production in belugas has been poorly characterized and its function is not well understood. I examined behavioral states when producing bubble streams (“bubbling”), and when bubbling calls, to determine whether bubbling was significantly associated with a particular call category or behavioral state. Using 19 hours of video and audio recordings collected over a two-day period, I quantified bubble streams of a 4-month old calf and an unrelated adult female housed together. Based on the overall activity budgets and pool of vocalizations for both animals, I calculated the expected counts of bubble streams with and without vocalizations, assuming that …
Coastal Wetland Dynamics Under Sea-Level Rise And Wetland Restoration In The Northern Gulf Of Mexico Using Bayesian Multilevel Models And A Web Tool, Tyler Hardy
Master's Theses
There is currently a lack of modeling framework to predict how relative sea-level rise (SLR), combined with restoration activities, affects landscapes of coastal wetlands with uncertainties accounted for at the entire northern Gulf of Mexico (NGOM). I developed such a modeling framework – Bayesian multi-level models to study the spatial pattern of wetland loss in the NGOM, driven by relative RSLR, vegetation productivity, tidal range, coastal slope, and wave height – all interacting with river-borne sediment availability, indicated by hydrological regimes. These interactions have not been comprehensively investigated before. I further modified this model to assess the efficacy of restoration …
Flow Regime Influences On Stream And Riparian Soil Carbon Dynamics In The Ozark Highlands And Boston Mountains Of Arkansas, Allyn Dodd
Graduate Theses and Dissertations
The natural flow regime exerts primacy over lotic ecosystem patterns and processes. However, little work has examined the influence of flow regime on instream and riparian carbon (C) dynamics in minimally-impacted, temperate forested systems. To date, most research efforts have focused on characterizing C movement and transformations across biomes and land use categories; however, flow regime represents an overlooked, finer level of detail that may drive differences in ecosystem function. My dissertation objective was to measure C fixation and movement within and across multiple environmental spheres (e.g. within stream channels, between stream surfaces and the atmosphere, and from riparian soils …
Hydrodynamic Drivers Of Dissolved Oxygen Variability Within A Highly Developed Tidal Creek In Myrtle Beach, South Carolina, Douglas Matthew Pastore
Hydrodynamic Drivers Of Dissolved Oxygen Variability Within A Highly Developed Tidal Creek In Myrtle Beach, South Carolina, Douglas Matthew Pastore
Electronic Theses and Dissertations
Erosion and water quality degradation have been observed in Singleton Swash in Myrtle Beach, SC, and have been hypothesized to be related to migration of the beach-face channel. Dredging this channel temporarily fixes erosional threats to nearby infrastructure but the effects on water quality are not well understood. It is hypothesized that variations in dissolved oxygen (DO) concentration (used here as a proxy for water quality) within the water column are related to changes in vertical mixing and transport due to oceanic tidal forcing. This study utilizes current meters, pressure sensors, and optical DO probes to measure and study the …
The Importance Of Keeping The Big Ones: Harvest Slot Limits And Marine Protected Areas For The Management Of The Caribbean Spiny Lobster, Gayathiri Gnanalingam
The Importance Of Keeping The Big Ones: Harvest Slot Limits And Marine Protected Areas For The Management Of The Caribbean Spiny Lobster, Gayathiri Gnanalingam
Biological Sciences Theses & Dissertations
Fishing typically removes the oldest and/or largest individuals from populations undermining stability and reproductive success. Traditional fisheries management tools fail to protect these oldest and/or largest individuals, but two less conventional tools: marine protected areas (MPAs), and harvest slot limits have the potential to do so. Here I tested the possible use of these tools for the Caribbean spiny lobster, Panulirus argus, an iconic and economically valued species. After decades of intense fishing, the largest lobsters have largely been wiped out. The loss of the largest lobsters is significant as large lobsters have considerably greater reproductive potential than their …
Potential Interaction Analysis Of Offshore Wind Energy Areas And Breeding Avian Species On The Us Mid-Atlantic Coast, Jeri Lynn Wisman
Potential Interaction Analysis Of Offshore Wind Energy Areas And Breeding Avian Species On The Us Mid-Atlantic Coast, Jeri Lynn Wisman
Biological Sciences Theses & Dissertations
Due to increasing US interest in developing wind energy sites in offshore waters, we synthesized existing data on colonial breeding seabird populations with the potential risk of interacting with lease areas in the mid-Atlantic. Previous efforts by BOEM and NOAA have predicted avian density using at-sea survey data; we seek to complement this work by focusing specifically on birds during the critical and energetically demanding breeding life history stage. We combined colony size and location for each species along the mid-Atlantic coast with buffers around the colonies that correlate with the species’ foraging range. We integrated population size, vulnerability to …
Desalination Concentrate Disposal: Ecological Effects And Sustainable Solutions, Ryan Hanley
Desalination Concentrate Disposal: Ecological Effects And Sustainable Solutions, Ryan Hanley
Global Honors Theses
Freshwater availability is a growing global concern, and desalination is often presented as the solution, but from this important technology comes issues of toxic waste. Ecosystems are delicate areas that contain species adapted to that specific location, and any chemical or physical changes can disrupt the fitness of species. The concentrate byproduct waste from desalination plants is toxic to species if the concentrate is not compatible with the receiving water body. A critical review of scientific articles, industry-leading books, conversations with industry experts, and information from the American Membrane Technology Association conference was used to analyze the current knowledge. Species …
Radical Social Ecology As Deep Pragmatism: A Call To The Abolition Of Systemic Dissonance And The Minimization Of Entropic Chaos, Arielle Brender
Radical Social Ecology As Deep Pragmatism: A Call To The Abolition Of Systemic Dissonance And The Minimization Of Entropic Chaos, Arielle Brender
Student Theses 2015-Present
This paper aims to shed light on the dissonance caused by the superimposition of Dominant Human Systems on Natural Systems. I highlight the synthetic nature of Dominant Human Systems as egoic and linguistic phenomenon manufactured by a mere portion of the human population, which renders them inherently oppressive unto peoples and landscapes whose wisdom were barred from the design process. In pursuing a radical pragmatic approach to mending the simultaneous oppression and destruction of the human being and the earth, I highlight the necessity of minimizing entropic chaos caused by excess energy expenditure, an essential feature of systems that aim …
Larval-Ant Interactions In The Mojave Desert: Communication Brings Us Together, Alicia Mellor
Larval-Ant Interactions In The Mojave Desert: Communication Brings Us Together, Alicia Mellor
UNLV Theses, Dissertations, Professional Papers, and Capstones
Butterflies are a diverse and essential group of pollinators whose abundance is predominantly determined by growth and survival of their larvae. In the family Lycaenidae, many species participate in a larval-ant mutualism where ants feed on nutrient-rich nectar produced by larvae and, in turn, protect those larvae from predators. Emerging evidence indicates larval scent in the form of cuticular hydrocarbons (CHCs), not nectar production, drives this interaction. This study takes two approaches to investigate CHCs as the driver behind the larval-ant mutualism in three butterfly species native to southern Nevada: Euphilotes bernardino martini (Martin’s blue), Brephidium exilis (western pygmy blue), …
Spatio-Temporal Dynamics Of Atlantic Cod Bycatch In The Maine Lobster Fishery And Its Impacts On Stock Assessment, Robert E. Boenish
Spatio-Temporal Dynamics Of Atlantic Cod Bycatch In The Maine Lobster Fishery And Its Impacts On Stock Assessment, Robert E. Boenish
Electronic Theses and Dissertations
Of the most iconic fish species in the world, the Atlantic cod (Gadus morhua, hereafter, cod) has been a mainstay in the North Atlantic for centuries. While many global fish stocks have received increased pressure with the advent of new, more efficient fishing technology in the mid-20th century, exceptional pressure has been placed on this prized gadoid. Bycatch, or the unintended catch of organisms, is one of the biggest global fisheries issues. Directly resulting from the failed recovery of cod in the GoM, attention has been placed as to possible sources of unaccounted catch. Among the most …
Polyp To Population: A Tale Of Two Corals, Christopher T. Fountain
Polyp To Population: A Tale Of Two Corals, Christopher T. Fountain
Electronic Theses and Dissertations
Deep-sea corals are of conservation concern in the North Atlantic due to prolonged disturbances associated with the exploitation of natural resources and a changing environment. As a result, the recovery rates of deep-sea coral communities are of heightened interest. These recovery rates are suggested to be on the order of decades to millennia, based on slow growth rates and longevity, of various deep-sea coral species. In 2014 and 2017 two research cruises in the Gulf of Maine, collected samples of two locally dominant species, Primnoa resedaeformis and Paramuricea placomus. These specimen collections were coupled with video surveys, conducted by …
Population Dynamics Of The Threatened Staghorn Coral, Acropora Cervicornis, And The Development Of A Species-Specific Monitoring Protocol, Elizabeth Goergen
Population Dynamics Of The Threatened Staghorn Coral, Acropora Cervicornis, And The Development Of A Species-Specific Monitoring Protocol, Elizabeth Goergen
HCNSO Student Theses and Dissertations
Historically, Acropora cervicornis was found in high densities on many Caribbean, Florida, and Gulf of Mexico reefs. A disease outbreak in the late 1970s and 80s caused up to 99% loss of A. cervicornis cover at some sites, leaving populations sparsely distributed throughout its range and typically found as isolated colonies. Even though populations are depauperate causing a decrease in sexual reproduction, its fast growth rate and ability to reproduce through asexual fragmentation affords this species the potential for quick recovery and population growth. However, limited to no natural recovery has been documented. Many of these populations are poorly studied …
The Combined Effects Of Soil Fertility And Soil Amendments On The Growth And Mycorrhizal Associations Of American Ginseng (Panax Quinquefolius), Emily Ann Murray
The Combined Effects Of Soil Fertility And Soil Amendments On The Growth And Mycorrhizal Associations Of American Ginseng (Panax Quinquefolius), Emily Ann Murray
Masters Theses, 2010-2019
Arbuscular mycorrhizal fungi (AMF) are fungi that form symbiotic associations with 70-90% of plant families. They are known to allow for the extension of the root system as well as an increase in plant size by assisting with the uptake of nutrients such as nitrogen and phosphorus. The role that AMF play in plant health and success has led to the development of commercial inoculum, which is used in agricultural settings. However, soil fertility, and soil amendments are known to affect AMF and plant associations. This study intends to look at how cultivated American Ginseng seedlings are affected by commercial …
Trophic Ecology And Parasitism Of A Mesopelagic Fish Assemblage, Matthew Woodstock
Trophic Ecology And Parasitism Of A Mesopelagic Fish Assemblage, Matthew Woodstock
HCNSO Student Theses and Dissertations
Mesopelagic (open ocean, 200-1000 m depth) fishes are important consumers of zooplankton and are prey of oceanic predators. Some mesopelagic fishes (e.g., myctophids and stomiids) undertake a diel vertical migration where they ascend to the near-surface waters during the night to feed and descend into the depths during the day to avoid predators. Other mesopelagic fishes (e.g., Sternoptyx spp.) do not vertically migrate and remain at deep depths throughout the day. While in the epipelagic zone (surface – 200 m depth), vertically migrating fishes become prey to upper-trophic level predators, such as: tunas and billfishes. Benthic fishes (e.g., macrourids) often …
Invertebrate Paleoecology Of High Paleo-Latitude Carboniferous Strata Of The Tepuel-Genoa Basin, Argentina, Kate M. Gigstad
Invertebrate Paleoecology Of High Paleo-Latitude Carboniferous Strata Of The Tepuel-Genoa Basin, Argentina, Kate M. Gigstad
Theses and Dissertations
The Late Paleozoic Ice Age (LPIA), which began in the latest Devonian and ended in the Late Permian, was a time of active environmental changes. During this time, glaciers waxed and waned over the ancient continent of Gondwana that was situated over the southern pole. This time of transition from icehouse to greenhouse in the Permian is our closest analogue to our current environmental conditions and increased knowledge of our planetary past will greatly influence our future. Much of the previous research on marine invertebrate paleoecology during the LPIA occurs in far-field or low paleolatitude localities, but less is known …
Life-History Characteristics And Fishery Dynamics Of Red Drum (Sciaenops Ocellatus), In The North-Central Gulf Of Mexico, Corbin Bennetts
Life-History Characteristics And Fishery Dynamics Of Red Drum (Sciaenops Ocellatus), In The North-Central Gulf Of Mexico, Corbin Bennetts
Master's Theses
Red Drum (Sciaenops ocellatus) are highly targeted recreationally in the Gulf of Mexico, and support a small commercial fishery in Mississippi. Despite their popularity, the stock is assessed using data limited approaches that necessitate accurate life history information. I estimated the year-specific and year-aggregated escapement rates for the Mississippi stock (years 2004 to 2015), which were sensitive to mortality estimation methods; year-aggregated estimates were 6.9 to 28.2 % depending on the method, but temporal patterns were consistent. I then addressed concerns with previously estimated life-history characteristics by describing the sex-specific growth and reproductive dynamics. The three-parameter von Bertalanffy …
Detrital Protein Contributes To Oyster Nutrition And Growth In The Damariscotta Estuary, Maine, Usa, Cheyenne M. Adams
Detrital Protein Contributes To Oyster Nutrition And Growth In The Damariscotta Estuary, Maine, Usa, Cheyenne M. Adams
Electronic Theses and Dissertations
Oyster aquaculture is an expanding industry that relies on identifying and utilizing natural estuarine conditions for the economically viable production of a filter-feeding crop. The eastern oyster, Crassostrea virginica, is the principal species currently cultured in Maine. In addition to preferentially consumed phytoplankton, various detrital complexes (non-algal and/or non-living organic matter) may provide some nutrition to C. virginica between times of phytoplankton abundance. Here I investigated the importance of detrital proteins in supporting the growth of oysters cultured in the upper Damariscotta Estuary. Oyster aquaculture in this area is highly successful and previous reports indicate that labile detrital protein …
Devonian Stromatoporoid Interactions At The Falls Of The Ohio State Park, Clarksville, Indiana, Morgan Sierra Hall
Devonian Stromatoporoid Interactions At The Falls Of The Ohio State Park, Clarksville, Indiana, Morgan Sierra Hall
Undergraduate Theses
Stromatoporoids are calcitic sponges that occurred in the fossil record from the Early Ordovician to Late Devonian period. These sponges interacted with other organisms, especially rugose and tabulate corals. Some corals appear to benefit from the rigidity of stromatoporoids in response to turbulent waters. Stromatoporoids and many corals went extinct during the Frasnian-Famennian crisis when paleoenvironmental parameters were shifting. Studying the relationships between these taxa may provide insight to their vulnerability during the extinction.
This research was performed at the Falls of the Ohio in Clarksville, Indiana. Organisms in the Coral Zone were studied using transect sampling. Each fossil along …