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2016

Zoology

Vaejovidae

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Full-Text Articles in Life Sciences

Laterobasal Aculear Serrations (Las) In Scorpion Family Vaejovidae (Scorpiones: Chactoidea), Victor Fet, Michael E. Soleglad, Michael S. Brewer Dec 2016

Laterobasal Aculear Serrations (Las) In Scorpion Family Vaejovidae (Scorpiones: Chactoidea), Victor Fet, Michael E. Soleglad, Michael S. Brewer

Victor Fet

The discovery of a new structure on the laterobasal aspect of the telson aculeus is described here for the first time. This structure, termed the laterobasal aculear serrations (LAS), is a row of minute denticles located on each side of the aculeus base, found exclusively in scorpion family Vaejovidae, thus potentially providing a new synapomorphy for this New World family. The LAS structure of major representative vaejovid genera and Vaejovis groups is illustrated with SEM micrographs. Also provided is a comprehensive list of non-vaejovid Recent scorpion genera examined where the LAS structure was found to be absent.


Contributions To Scorpion Systematics. Ii. Stahnkeini, A New Tribe In Scorpion Family Vaejovidae (Scorpiones: Chactoidea), Michael E. Soleglad, Victor Fet Dec 2016

Contributions To Scorpion Systematics. Ii. Stahnkeini, A New Tribe In Scorpion Family Vaejovidae (Scorpiones: Chactoidea), Michael E. Soleglad, Victor Fet

Victor Fet

The diagnostic characters originally established by Herbert L. Stahnke (1940a, 1940b, 1974) in his description of genus Serradigitus are studied in detail from several new perspectives. A new genus, Stahnkeus, gen. nov., is described based on the presence of inner accessory (IAD) denticles on the chelal fingers, unprecedented in family Vaejovidae. Five species of Serradigitus are transferred to Stahnkeus: Stahnkeus harbisoni (Williams, 1970), comb. nov. (=Serradigitus harbisoni); Stahnkeus deserticola (Williams, 1970), comb. nov. (=Serradigitus deserticola); Stahnkeus subtilimanus (Soleglad, 1972), comb. nov. (=Serradigitus subtilimanus); Stahnkeus allredi (Sissom et Stockwell, 1991), comb. nov. (=Serradigitus allredi); and Stahnkeus polisi (Sissom et Stockwell, 1991), …


A New Scorpion Genus (Scorpiones: Vaejovidae) From Mexico, Michael E. Soleglad, Victor Fet Dec 2016

A New Scorpion Genus (Scorpiones: Vaejovidae) From Mexico, Michael E. Soleglad, Victor Fet

Victor Fet

A new vaejovid genus from Mexico, Franckeus, gen. nov., is described based on unique neobothriotaxy. Species placed in this genus are from the Vaejovis “nigrescens” group (previously called the “nitidulus” group). Six species comprise this new genus, distributed throughout Mexico (mainland as well as Baja California Sur): Franckeus nitidulus, F. rubrimanus, F. platnicki, F. minckleyi, F. kochi and F. peninsularis. A new species of the Vaejovis “nigrescens” group, Vaejovis davidi, sp. nov., is also described.


Contributions To Scorpion Systematics. Iii. Subfamilies Smeringurinae And Syntropinae (Scorpiones: Vaejovidae), Michael E. Soleglad, Victor Fet Dec 2016

Contributions To Scorpion Systematics. Iii. Subfamilies Smeringurinae And Syntropinae (Scorpiones: Vaejovidae), Michael E. Soleglad, Victor Fet

Victor Fet

Stockwell (1989), in an unpublished revision, suggested splitting the family Vaejovidae in two subfamilies, introducing also four tribes and four new genera. These results have not been reevaluated for 20 years. Here, we establish a new subfamily Smeringurinae, subfam. nov., with two tribes, four genera, and 40 species and subspecies. The subfamily Smeringurinae includes two tribes: Smeringurini, trib. nov. and Paravaejovini, trib. nov. The tribe Smeringurini includes three genera: Paruroctonus Werner, 1934; Smeringurus Haradon, 1983; and Vejovoidus Stahnke, 1974. The tribe Paravaejovini includes the genus Paravaejovis Williams, 1980. We restore from synonymy the subfamily Syntropinae Kraepelin, 1905, and significantly expand …


Graemeloweus, A New Scorpion Genus From Northern California, Usa (Scorpiones: Vaejovidae), Michael E. Soleglad, Victor Fet, Matthew R. Graham, Richard F. Ayrey Dec 2016

Graemeloweus, A New Scorpion Genus From Northern California, Usa (Scorpiones: Vaejovidae), Michael E. Soleglad, Victor Fet, Matthew R. Graham, Richard F. Ayrey

Victor Fet

Genus Graemeloweus, gen. nov. (Scorpiones: Vaejovidae) is described from northern California, USA. The genus is composed of three species formerly placed in Pseudouroctonus: Graemeloweus iviei (Gertsch et Soleglad, 1972), comb. nov. (type species), G. glimmei (Hjelle, 1972), comb. nov., and G. maidu (Savary et Bryson, 2016), comb. nov. Major diagnostic characters of Graemeloweus include a non-bifurcated primary lamellar hook, the presence of a secondary lamellar hook, a complex mating plug with a two part base and an asymmetric crescent-shape barb, and the presence of a well-developed ventromedian (V2) carina on the pedipalp chela. Evidence is presented suggesting that Graemeloweus is …


Kovarikia, A New Scorpion Genus From California, Usa (Scorpiones: Vaejovidae), Michael E. Soleglad, Victor Fet, Matthew R. Graham Dec 2016

Kovarikia, A New Scorpion Genus From California, Usa (Scorpiones: Vaejovidae), Michael E. Soleglad, Victor Fet, Matthew R. Graham

Victor Fet

Genus Kovarikia, gen. nov. (Scorpiones: Vaejovidae) is described from southern California, USA. The genus is composed of three species previously placed in Pseudouroctonus: Kovarikia williamsi (Gertsch et Soleglad, 1972), comb. nov. (type species), K. bogerti (Gertsch et Soleglad, 1972), comb. nov., and K. angelena (Gertsch et Soleglad, 1972), comb. nov. Major diagnostic characters of Kovarikia are its unique neobothriotaxy found on the ventral surface of the pedipalp chelae, the occurrence of a secondary lamellar hook on the hemispermatophore, a crescent-shaped mating plug barb, the presence of a secondary exteromedian (EMc) carina on the pedipalp patella, and a swollen telson vesicle …


Graemeloweus, A New Scorpion Genus From Northern California, Usa (Scorpiones: Vaejovidae), Michael E. Soleglad, Victor Fet, Matthew R. Graham, Richard F. Ayrey Jul 2016

Graemeloweus, A New Scorpion Genus From Northern California, Usa (Scorpiones: Vaejovidae), Michael E. Soleglad, Victor Fet, Matthew R. Graham, Richard F. Ayrey

Euscorpius

Genus Graemeloweus, gen. nov. (Scorpiones: Vaejovidae) is described from northern California, USA. The genus is composed of three species formerly placed in Pseudouroctonus: Graemeloweus iviei (Gertsch et Soleglad, 1972), comb. nov. (type species), G. glimmei (Hjelle, 1972), comb. nov., and G. maidu (Savary et Bryson, 2016), comb. nov. Major diagnostic characters of Graemeloweus include a non-bifurcated primary lamellar hook, the presence of a secondary lamellar hook, a complex mating plug with a two part base and an asymmetric crescent-shape barb, and the presence of a well-developed ventromedian (V2) carina on the pedipalp chela. Evidence is presented suggesting that …


Pseudouroctonus Peccatum Tate Et Al., 2013, Rediscovered, Richard F. Ayrey Jan 2016

Pseudouroctonus Peccatum Tate Et Al., 2013, Rediscovered, Richard F. Ayrey

Euscorpius

Shortly after the discovery of Pseudouroctonus peccatum Tate, Riddle, Soleglad et Graham, 2013 (Scorpiones: Vaejovidae) there was a large wildfire in the Spring Mountains of Nevada. At the time the species description was published, it was noted that the only known population of this species may have been extirpated by the fire and subsequent flooding of the type locality habitat (Tate et al. 2013). Today it is certain that the species has survived after the collection of 12 new specimens. Additional data on males are also presented.


New Analysis Of The Genus Pseudouroctonus With The Description Of Two New Species (Scorpiones: Vaejovidae), Richard F. Ayrey, Michael E. Soleglad Jan 2016

New Analysis Of The Genus Pseudouroctonus With The Description Of Two New Species (Scorpiones: Vaejovidae), Richard F. Ayrey, Michael E. Soleglad

Euscorpius

Two new species of Pseudouroctonus are described from southeastern Arizona, USA, Pseudouroctonus santarita, sp. nov. from the Santa Rita Mountains and P. kremani, sp. nov. from the Santa Catalina Mountains. These new species are closely related to P. apacheanus (Gertsch et Soleglad, 1972). A combination of morphological differences in the hemispermatophore, the mating plug, and several morphometric-based characters are identified as diagnostic. New substructures are identified for the mating plug.


Hermaphroditism, Gynandromorphism, And Four Pectines: An Extreme Case Of Developmental Anomaly In Scorpions (Scorpiones: Vaejovidae), Rolando Teruel, José Guadalupe Baldazo-Monsivaiz Jan 2016

Hermaphroditism, Gynandromorphism, And Four Pectines: An Extreme Case Of Developmental Anomaly In Scorpions (Scorpiones: Vaejovidae), Rolando Teruel, José Guadalupe Baldazo-Monsivaiz

Euscorpius

In the present note, we describe in detail an aberrant anomalous specimen of the scorpion Mesomexovis punctatus (Karsch, 1879) (Vaejovidae). This strange individual is an adult and exhibits a combination of hermaphroditism, gynandromorphism, and pectinal duplication (i.e., presence of four pectines); the latter represents the first report ever made of such teratology. It was collected by the authors in northeastern Guerrero State, Mexico.


New Species Of Vaejovis From The Whetstone Mountains, Southern Arizona (Scorpiones: Vaejovidae), Richard F. Ayrey, Michael E. Soleglad Jan 2016

New Species Of Vaejovis From The Whetstone Mountains, Southern Arizona (Scorpiones: Vaejovidae), Richard F. Ayrey, Michael E. Soleglad

Euscorpius

A new scorpion species, Vaejovis troupi sp. n., is described and placed in the “vorhiesi” group of the genus Vaejovis. Based on a recent molecular analysis of Bryson et al. (2013), this species is shown to be related to V. vorhiesi and V. grahami. Two of three diagnostic characters found in this new species are the presence of six inner denticles (ID) on the pedipalpal fixed and movable fingers, and a unique arrangement of trichobothria on the external surface of the pedipalp patella. This species was found in an isolated montane habitat in …


A New Species Of Vaejovis From Chaparral Habitat Near Yarnell, Arizona (Scorpiones: Vaejovidae), Richard F. Ayrey Jan 2016

A New Species Of Vaejovis From Chaparral Habitat Near Yarnell, Arizona (Scorpiones: Vaejovidae), Richard F. Ayrey

Euscorpius

A new scorpion species, Vaejovis grayae sp. nov. is described and placed in the “vorhiesi” group of the genus Vaejovis. This small brown species is found near Yarnell, Arizona, USA. It appears most similar to V. trinityae Ayrey and V. crumpi Ayrey et Soleglad. It can be distinguished from the other members of the “vorhiesi” group by a unique combination of non-overlapping morphological characters and multilocus DNA data (Bryson et al., 2013). The pedipalp fixed finger has 6 ID denticles and the movable finger has 7, like most other northern Arizona “vorhiesi” group species. Another …


Kovarikia, A New Scorpion Genus From California, Usa (Scorpiones: Vaejovidae), Michael E. Soleglad, Victor Fet, Matthew R. Graham Jan 2016

Kovarikia, A New Scorpion Genus From California, Usa (Scorpiones: Vaejovidae), Michael E. Soleglad, Victor Fet, Matthew R. Graham

Euscorpius

Genus Kovarikia, gen. nov. (Scorpiones: Vaejovidae) is described from southern California, USA. The genus is composed of three species previously placed in Pseudouroctonus: Kovarikia williamsi (Gertsch et Soleglad, 1972), comb. nov. (type species), K. bogerti (Gertsch et Soleglad, 1972), comb. nov., and K. angelena (Gertsch et Soleglad, 1972), comb. nov. Major diagnostic characters of Kovarikia are its unique neobothriotaxy found on the ventral surface of the pedipalp chelae, the occurrence of a secondary lamellar hook on the hemispermatophore, a crescent-shaped mating plug barb, the presence of a secondary exteromedian (EMc) carina on the pedipalp …


New Species Of Vaejovis From The Santa Rita Mountains, Southern Arizona (Scorpiones: Vaejovidae), Richard F. Ayrey, Michael E. Soleglad Jan 2016

New Species Of Vaejovis From The Santa Rita Mountains, Southern Arizona (Scorpiones: Vaejovidae), Richard F. Ayrey, Michael E. Soleglad

Euscorpius

A new scorpion species, Vaejovis grahami sp. nov. is described. This small, dark brown species is found in the Santa Rita Mountains, Santa Cruz County, Arizona, USA. It is most similar to V. vorhiesi Stahnke and V. deboerae Ayrey. The pedipalp fixed finger usually has 5 inner denticles (ID) and the movable finger has 6, like in most other southern Arizona Vaejovis.


A New Species Of Vaejovis From The Mogollon Rim Of Northern Arizona (Scorpiones: Vaejovidae), Richard F. Ayrey Jan 2016

A New Species Of Vaejovis From The Mogollon Rim Of Northern Arizona (Scorpiones: Vaejovidae), Richard F. Ayrey

Euscorpius

A new scorpion species, Vaejovis trinityae sp. nov. is described. This small brown species is found along the Mogollon Rim above Strawberry, Arizona. This is the first description of a new species of the “vorhiesi” group scorpions whose DNA phylogenetic analysis was published (Bryson et al., 2013); based on DNA data, the new species is most related to V. lapidicola Stahnke and V. crumpi Ayrey et Soleglad. It represents one of the “twenty-seven geographically cohesive lineages inferred from the mtDNA tree”. A unique characteristic of this species is that it exhibits arboreal behavior, being frequently found on Ponderosa …


Reproduction And Birth In The “Vorhiesi” Group Of The Genus Vaejovis (Scorpiones: Vaejovidae). Part I. Clutch Size, Richard F. Ayrey Jan 2016

Reproduction And Birth In The “Vorhiesi” Group Of The Genus Vaejovis (Scorpiones: Vaejovidae). Part I. Clutch Size, Richard F. Ayrey

Euscorpius

A total of 2,287 newborn scorpions were counted and photographed on 100 females of all 12 described species of the “vorhiesi” group of the genus Vaejovis (Scorpiones: Vaejovidae) found in Arizona, USA, and several undescribed species. Average clutch size for the “vorhiesi” group and for each individual species is reported. The data clearly shows that the larger species have more young, per parturition, than the smaller species. Representative photographs are presented for each species discussed.


Sky Island Vaejovis: A New Species (Scorpiones: Vaejovidae), Richard F. Ayrey Jan 2016

Sky Island Vaejovis: A New Species (Scorpiones: Vaejovidae), Richard F. Ayrey

Euscorpius

A new scorpion species, Vaejovis deboerae, sp. nov., is described and placed in the “mexicanus” group of the genus Vaejovis. They are small light yellowish brown scorpions found in the Santa Catalina Mountains, one of the sky islands of southern Arizona. They are closely related to V. vorhiesi Stahnke. The original description of V. vorhiesi Stahnke, 1940 and redescription (Graham, 2007), state that they are found in the Huachuca and Santa Catalina Mountains of southern Arizona. No specimens from the Santa Catalina Mountains were included in the redescription based on the lack of adequate material. The redescription …


Contributions To Scorpion Systematics. Iii. Subfamilies Smeringurinae And Syntropinae (Scorpiones: Vaejovidae), Michael E. Soleglad, Victor Fet Jan 2016

Contributions To Scorpion Systematics. Iii. Subfamilies Smeringurinae And Syntropinae (Scorpiones: Vaejovidae), Michael E. Soleglad, Victor Fet

Euscorpius

Stockwell (1989), in an unpublished revision, suggested splitting the family Vaejovidae in two subfamilies, introducing also four tribes and four new genera. These results have not been reevaluated for 20 years. Here, we establish a new subfamily Smeringurinae, subfam. nov., with two tribes, four genera, and 40 species and subspecies. The subfamily Smeringurinae includes two tribes: Smeringurini, trib. nov. and Paravaejovini, trib. nov. The tribe Smeringurini includes three genera: Paruroctonus Werner, 1934; Smeringurus Haradon, 1983; and Vejovoidus Stahnke, 1974. The tribe Paravaejovini includes the genus Paravaejovis Williams, 1980. We restore from synonymy the subfamily Syntropinae Kraepelin, 1905, and …


A New Scorpion Genus Representing A Primitive Taxon Of Tribe Stahnkeini, With A Description Of A New Species From Sonora, Mexico (Scorpiones: Vaejovidae), Matthew R. Graham, Michael E. Soleglad Jan 2016

A New Scorpion Genus Representing A Primitive Taxon Of Tribe Stahnkeini, With A Description Of A New Species From Sonora, Mexico (Scorpiones: Vaejovidae), Matthew R. Graham, Michael E. Soleglad

Euscorpius

A new scorpion genus, Gertschius gen. nov., and species, Gertschius crassicorpus sp. nov., (Scorpiones: Vaejovidae) from Sonora, Mexico is described. Gertschius exhibits a reduced modification of the basal pectinal teeth in the female holotype and serration of the chelal finger denticles is limited to the median denticles (MD), thus we consider it a relict member of tribe Stahnkeini. Serradigitus agilis Sissom et Stockwell, 1991, is transferred to this new genus; Gertschius agilis, comb. nov.


Sky Island Vaejovis: Two New Species And A Redescription Of V. Vorhiesi Stahnke (Scorpiones: Vaejovidae), Matthew R. Graham Jan 2016

Sky Island Vaejovis: Two New Species And A Redescription Of V. Vorhiesi Stahnke (Scorpiones: Vaejovidae), Matthew R. Graham

Euscorpius

Two new scorpion species, Vaejovis cashi sp. nov. and V. feti sp. nov. are described and placed in the “mexicanus” group of the genus Vaejovis. These small scorpions were collected at high-elevations from the sky island archipelago of southern Arizona and New Mexico. Both new species are similar to V. vorhiesi Stahnke, a small mesic scorpion of the Huachuca and Santa Catalina Mountains. The original description of V. vorhiesi consists of a single uninformative paragraph and a holotype was never designated. This paper provides a redescription and lectotype designation for V. vorhiesi and describes the two new related species. …