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Full-Text Articles in Life Sciences

New York City’S Rising Sea Level And Coastal Erosion: Approaches To Resiliency, Ryder D. Isidro May 2024

New York City’S Rising Sea Level And Coastal Erosion: Approaches To Resiliency, Ryder D. Isidro

Student Theses 2015-Present

No abstract provided.


Automated Tree Mortality Detection Using Ubiquitously Available Public Data, Michael T. Huggins Mar 2024

Automated Tree Mortality Detection Using Ubiquitously Available Public Data, Michael T. Huggins

Master's Theses

Understanding the dynamic interplay between fire severity, topography, and tree mortality, is crucial for predicting future forest dynamics and enhancing resilience against climate change-induced wildfire regimes. This thesis develops a multi-sensor approach for automated estimation of tree mortality, then applies it to examine trends in tree mortality over a six-year period across a fire affected study site in the Trinity River basin in Northern California. The Random Forest model uses publicly available USGS 3D Elevation Program Lidar (3DEP) and NAIP imagery as inputs and is likely to be easily adaptable to other landscapes. The model had a Receiver Operating Characteristic …


Longevity Of Prescribed Fire Effectiveness In Mixed-Evergreen Forests Of The Klamath Mountains, Kaily M. Fineran Jan 2024

Longevity Of Prescribed Fire Effectiveness In Mixed-Evergreen Forests Of The Klamath Mountains, Kaily M. Fineran

Cal Poly Humboldt theses and projects

Prescribed fire is a common management tool used across the western United States to create wildfire-resilient ecosystems and communities. The Klamath Mountains of northern California, USA has experienced numerous, large wildfires in recent years. This is due in part to a combination of highly flammable fuels, uncharacteristic weather events, and high fuel accumulations as a result of fire exclusion and forest management practices. Within the last decade, local land management organizations and the Karuk Tribe have begun re-introducing prescribed fire in low elevation, mixed evergreen forests. While the long-term effects of fuel treatments are not well-known, these efforts provided a …


Investigating Birds As Dispersal Vectors Of Litylenchus Crenatae Subsp. Mccannii (Anguinidae), The Nematode Associated With Beech Leaf Disease, Spencer Rock Parkinson Jan 2024

Investigating Birds As Dispersal Vectors Of Litylenchus Crenatae Subsp. Mccannii (Anguinidae), The Nematode Associated With Beech Leaf Disease, Spencer Rock Parkinson

Graduate Theses, Dissertations, and Problem Reports

Beech leaf disease (BLD) is an emerging forest pathogen primarily affecting American beech (Fagus grandifolia, Ehrh.) in North America and has been attributed to tree mortality of sapling sized trees within five to seven years of infection. Symptoms typically occur in regenerating American beech thickets sprouting from roots of trees killed by beech bark disease. Scientists first observed BLD in Ohio in 2012 and currently has spread to 15 states in the USA and one Canadian province. The nematode Litylenchus crenatae subsp. mccannii (Lcm) is highly associated with BLD symptoms, interveinal chlorosis and defoliation of leaves, and is currently …


Quantifying The Ecological Effects Of Salix Fragilis On Riparian Habitat In Kittitas County, Washington, Landon Shaffer Jan 2024

Quantifying The Ecological Effects Of Salix Fragilis On Riparian Habitat In Kittitas County, Washington, Landon Shaffer

All Master's Theses

Invasive species threaten plant community structure and function globally. Riparian areas, the zone near streams where water influences vegetation, are especially sensitive to invasive species colonization, suffering large-scale shifts in community composition. Salix fragilis (crack willow) is a nonnative riparian species abundant in the lower elevation tributaries of central Washington. Some speculate whether this willow should be listed as invasive in Washington, despite a lack of regional supporting evidence. I studied riparian communities dominated by either S. fragilis or native species in the Kittitas Valley and measured biodiversity, quantified differences in solar attenuation, and compared leaf decomposition rates to learn …


A Conservation Model: Costa Rican Conservation Strategies Effectively Preserve Their Threatened Primates, Ryan Belmont Jan 2024

A Conservation Model: Costa Rican Conservation Strategies Effectively Preserve Their Threatened Primates, Ryan Belmont

Regis University Student Publications (comprehensive collection)

The wildlife of Costa Rica has experienced various anthropogenic threats over the last century including climate change and agricultural expansion. The mantled howler monkey (Alloutta palliata), Central American spider monkey (Ateles geoffroyi), white-faced capuchin (Cebus imitator), and the Central American squirrel monkey (Saimiri oerstedii) are Costa Rica’s native primates that face several anthropogenic threats such as deforestation for agriculture and climate change. In response to increased threats to its four native species of non-human primates, Costa Rica has implemented effective governmental conservation tactics such as the Payments for Environmental Services program, ecotourism …


Environmental Biology Masters Capstone, Antonio Gonzalez-Pita Jan 2024

Environmental Biology Masters Capstone, Antonio Gonzalez-Pita

Regis University Student Publications (comprehensive collection)

Human wildlife interactions (HWI) pose a complex challenge for wildlife managers. Human encroachment into wildlife habitat and the growing number of outdoor recreationists are increasing the frequency of contact and conflict, especially in regions such as the Front Range of Colorado. Geographic information systems (GIS), which use a combination of remote sensing and environmental survey data, allow for predictive spatial analyses of where human wildlife interactions are likely to occur. I used publicly reported observations of moose to create spatial predictive maps in a species distribution model framework. Slope and elevation were shown to be the strongest predictors of HWI, …


Movement Behavior And Metapopulation Connectivity Of Stream Salamanders In Response To Disturbance Events, Kathryn M. Greene Jan 2024

Movement Behavior And Metapopulation Connectivity Of Stream Salamanders In Response To Disturbance Events, Kathryn M. Greene

Theses and Dissertations--Biology

Metapopulations are shaped by the dispersal between populations in a landscape. Disturbance events can disrupt this connectivity resulting in local population extinction. For my dissertation, I used a combination of empirical and theoretical techniques to examine dispersal in response to disturbance and assessed it’s population-level consequences. My research used capture-mark-recapture sampling techniques to evaluate stream salamander movement in response to (1) a supraseasonal drought and (2) mountaintop-removal-mining (MTR) and valley-filling (VF) and (3) agent-based simulation modeling to evaluate population extinction risk in response to varying dispersal and mortality rates.

First, I evaluated the effects of a supraseasonal drought, a severe …


Regeneration Response To Salvage Logging Following Tornado Disturbance, Colby K. Bosley-Smith Dec 2023

Regeneration Response To Salvage Logging Following Tornado Disturbance, Colby K. Bosley-Smith

Electronic Theses and Dissertations

In an era of increasing natural disturbances, successful tree regeneration has grown more difficult to achieve. Salvage logging, a common management response to disturbance, may further impede regeneration success, although published literature currently remains inconclusive. In 2013, a rare tornado in northcentral Maine, USA, and subsequent salvage operation created three clear ‘treatments’ for evaluation of post-disturbance regeneration: blowdown, blowdown followed by salvage logging and an undisturbed control. In the summers of 2022 and 2023, (nine and ten) years post-tornado, we revisited this site to examine regeneration outcomes.

During the summer of 2022, we evaluated stand structure and regeneration success of …


Germination Trends Of American Chaffseed, Schwalbea Americana L., And Factors Affecting First-Year Seedling Development, Trenton Miller Dec 2023

Germination Trends Of American Chaffseed, Schwalbea Americana L., And Factors Affecting First-Year Seedling Development, Trenton Miller

All Theses

Following centuries of exploitation and fire suppression, longleaf pine systems are now the focus of many conservation efforts. Efforts to restore populations of Schwalbea americana L. in longleaf pine savannas have been met with frustratingly low recruitment. While past studies have briefly quantified germination rates for Schwalbea, there have not been any studies yet that truly investigate this plant’s germination requirements. Additionally, there has been little research into characterizing the parasitic relationship between Schwalbea and its various host species. We conducted a germination study in a growth chamber that investigated Schwalbea’s germination rate and time to germinates as …


Phenology Of Asian Longhorned Beetle (Anoplophora Glabripennis, Coleoptera: Cerambycidae) In South Carolina, U.S.A., Lena Schmitt Dec 2023

Phenology Of Asian Longhorned Beetle (Anoplophora Glabripennis, Coleoptera: Cerambycidae) In South Carolina, U.S.A., Lena Schmitt

All Theses

A new population of the Asian longhorned beetle (Anoplophora glabripennis Motschulsky, hereafter ALB), an invasive species in the United States since 1996, was discovered in Charleston County, South Carolina in 2020. This population is the furthest south ALB has been found in North America and provides an opportunity to examine the phenology of this beetle in a novel climate. I collected 1009 eggs, larvae, pupae, and adult beetles from the federal quarantine zone in South Carolina and used larval head capsule widths to determine the rate of development and voltinism of ALB in South Carolina’s coastal climate. With this …


The Discovery Of A Novel Bacteria From A Large Co-Assembly Of Metagenomes, Matthew Finkelberg Nov 2023

The Discovery Of A Novel Bacteria From A Large Co-Assembly Of Metagenomes, Matthew Finkelberg

Masters Theses

In the summer of 2022, a co-assembly of metagenome was created using the microbes found at Barres Woods in Harvard Forest. 14 samples were taken, and sample was split into the organic and mineral layer, which totals 28 Bulk MAGs. Within this Co-assembly, 4 different genomes were found which were designated with the phylum of FCPU426. Three of which were considered medium quality and one being assigned high quality. The novel phyla first appeared in NCBI and GTDB databases in June 2018. The name FCPU426 dates to 2010 and was named based on the 16s amplicon sequencing.

The novel phylum …


Climate Interactions Drive Tree Physiology And Growth In A Northeastern Forest Ecotone, Alexandra M. Barry Aug 2023

Climate Interactions Drive Tree Physiology And Growth In A Northeastern Forest Ecotone, Alexandra M. Barry

Electronic Theses and Dissertations

Climate change is a threat to global forest ecosystems. In the northeastern United States, forest trees are facing rising temperatures and increasingly inconsistent moisture regimes. In addition to long-term changes in climate conditions, there is concern about the potential for more frequent and intense climate extremes, which can have severe and rapid negative effects on tree physiology and growth. Further, climate extremes may co-occur to produce a greater magnitude of effect than the sum of their parts, with a prominent example being hot droughts, which are increasing in occurrence and severity. The impact of these and other extreme climate interactions …


A Four-Pronged Approach To Addressing A Wild Pig Invasion In A Bottomland And Upland Forested Landscape, Tyler Scott Evans Aug 2023

A Four-Pronged Approach To Addressing A Wild Pig Invasion In A Bottomland And Upland Forested Landscape, Tyler Scott Evans

Theses and Dissertations

Among exotic species that are capable of invading, establishing, and reaching pest status, few pose the range of impacts to biotic (e.g., competition with native species, predation, herbivory, introduction of other exotics) and abiotic (e.g., soil, hydrology) ecosystem components that can be attributed to the wild pig (Sus scrofa). Despite the presence of wild pigs throughout the southeastern United States for centuries, new invasions continue to occur in previously uninhabited and often under-investigated landscapes, including bottomland and upland forests. The recent invasion of the Sam D. Hamilton Noxubee National Wildlife Refuge (hereafter, NNWR) in east-central Mississippi represents an …


Modeling The Effect Of Neighborhood Competition On Tree Diameter Growth In The Pacific Northwest Coast Range, Htet Lin Naing Aug 2023

Modeling The Effect Of Neighborhood Competition On Tree Diameter Growth In The Pacific Northwest Coast Range, Htet Lin Naing

Theses and Dissertations

Trees compete for various resources such as sunlight, water, and nutrients, which can be expressed as numerical terms, called competition indices (CI). Competition between individual trees is correlated with their growth and mortality. Therefore, CIs are used as independent variables to develop, improve and modify growth and yield models. This study was conducted to test the effect of neighborhood competition on tree diameter growth among Douglas-fir (Pseudotsuga menziesii (Mirb.) Franco), western hemlock (Tsuga heterophylla (Raf.) Sarg) and red alder (Alnus rubra Bong.), in the Pacific Northwest Coast Range, USA. After testing seven distance-independent CIs and three distance-dependent …


Gene Expression Effects On Productivity And Stress Tolerance In Polyclonal Plantings Of Populus Deltoides, Macy Gosselaar Aug 2023

Gene Expression Effects On Productivity And Stress Tolerance In Polyclonal Plantings Of Populus Deltoides, Macy Gosselaar

Theses and Dissertations

Polyclonal plantings of Populus deltoides are expected to display increased site resource use, productivity, and tolerance to stress through plasticity changes leading to niche differentiation (i.e changes to crown/canopy structures). In the present study, P. deltoides Clones S7C8, 110412, and polyclonal plots were tested for differentially expressed genes and enriched biological pathways between planting schemes. Transcriptomic analysis of leaves revealed upregulation of an active growth gene and gene family members that play important roles in plant stress and stress tolerance in polyclonal plantings. A gene associated with oxidative stress was upregulated in polyclonal plantings across all treatments. Secondary metabolic pathways …


Evidence Of Competitive Release Following Overstory Mortality In A Semi-Arid Piñon-Juniper Woodland, Corrie D. Gonzalez Aug 2023

Evidence Of Competitive Release Following Overstory Mortality In A Semi-Arid Piñon-Juniper Woodland, Corrie D. Gonzalez

Biology ETDs

Extreme temperatures and severe drought events have led to widespread tree mortality worldwide. In semi-arid regions of the Southwest United States, these events pose a significant threat to piñon-juniper (PJ) woodlands. We studied the effects of piñon and juniper mortality on the growth and physiology of existing saplings in PJ woodlands by analyzing water status, photosynthetic activity, and tissue chemistry to gain insights into these impacts. Juniper saplings exhibited improved water status and water use efficiency in response to overstory mortality, whereas piñon saplings did not. Additionally, both piñon and juniper saplings exhibited increased photosynthetic rates, increased photosynthetic capacity, and …


Patterns And Drivers Of Wiregrass Gap Longleaf Pine (Pinus Palustris Mill.) Woodland Succession As Part Of Restoration Efforts, Armin Weise Aug 2023

Patterns And Drivers Of Wiregrass Gap Longleaf Pine (Pinus Palustris Mill.) Woodland Succession As Part Of Restoration Efforts, Armin Weise

All Theses

Longleaf Pine (Pinus palustris) communities are widespread throughout the Southeastern United States with a dominant understory vegetation of wiregrass (Aristida spp.) in most of its range. A small area in central South Carolina that is naturally free of wiregrass is called the “Wiregrass Gap”. Here, the understory vegetation is dominated by bluestems grasses (Andropogon spp. and Schizachyrium spp.) which drive the disturbance regime of frequent low-intensity fire. The successful establishment of these grasses is key for longleaf pine woodland restoration efforts in this region, but few resources detail the ecological drivers at play that enable successful restoration in these longleaf …


Influence Of Abiotic Drivers And Plant Community Interactions On Bald Cypress (Taxodium Distichum) Seedlings: Implications For Restoration, Victoria Ellis Aug 2023

Influence Of Abiotic Drivers And Plant Community Interactions On Bald Cypress (Taxodium Distichum) Seedlings: Implications For Restoration, Victoria Ellis

Biological Sciences Theses & Dissertations

Forested freshwater wetlands are valuable ecosystems that provide habitat for numerous species, sequester carbon, and act as sinks for excess water and nutrients. Historically, these ecosystems have been heavily degraded by anthropogenic activities leading to loss of ecosystem services and a desire to restore freshwater forested wetland habitat. Thus, science-backed approaches for the restoration of freshwater forested wetlands are necessary to ensure restoration goals are met. This body of research employed the Stress Gradient Hypothesis to test whether a multi-species planting approach using Juncus effusus (L.) (soft rush) could facilitate the survival of Taxodium distichum (L.) Rich. (bald cypress) seedlings …


Advancing Natural White Oak (Quercus Alba.) Reproduction Through A Midstory Removal, Canaan Jeffrey Dugger Aug 2023

Advancing Natural White Oak (Quercus Alba.) Reproduction Through A Midstory Removal, Canaan Jeffrey Dugger

Masters Theses

In recent decades, there has been an alarming decline in white oak (Quercus alba.) regeneration and recruitment occurring within eastern forests. Historically, the frequency and the intensity of anthropogenic disturbances was dynamic over the landscape. However, over just the last century a major compositional shift has occurred from this change in disturbance regimes. Eastern forests are now promoting the regeneration of mesic species that obstruct white oaks’ ability to compete successfully and establish in the upper canopy.

The majority of successful white oak regeneration and recruitment is now occurring on the average to more xeric sites. Limited studies …


Disturbance Regimes And Management Strategies Of Mountain Ash Forest Ecosystems In Victoria, Australia; A Literature Review, Zoe Plumb May 2023

Disturbance Regimes And Management Strategies Of Mountain Ash Forest Ecosystems In Victoria, Australia; A Literature Review, Zoe Plumb

International Development, Community and Environment (IDCE)

This paper discusses the ecology of mountain ash forests, the disturbances regimes that currently exist in these ecosystems, and finally addresses the current management practices and future management practices. Mountain ash forests are subjected to a wide range of research in the Central Highlands of Victoria, an area approximately 14,000 hectares in range. These forests are dominated by montane ash trees (Eucalyptus regnans F. Muell), which are critically endangered and at risk of collapse, attributed to the decline in large hollow-bearing trees throughout the region. Management of these forests are controlled by the Department of Environment, Land, Water, and …


Top Down Effects And Resource Selection By Coyotes In South Carolina, Alex Jensen May 2023

Top Down Effects And Resource Selection By Coyotes In South Carolina, Alex Jensen

All Dissertations

Top predators play important roles in functioning ecosystems, including regulating the populations of prey species and competing with other predators. However, in the face of global change, many top terrestrial predators have declined in both range and abundance, making room for some smaller predators to expand into new niches. Coyotes (Canis latrans) in North America are a prime example of this – they have rapidly expanded their range in the last 120 years, raising concerns about their impacts on both domestic and wild species. In eastern North America, research has centered around their effects on white-tailed deer ( …


Geomorphology Of Tidal Wetlands: Impacts Of Extreme And Annual Flood Events To Salt Marsh And Mangrove Systems, Frances R. Griswold Apr 2023

Geomorphology Of Tidal Wetlands: Impacts Of Extreme And Annual Flood Events To Salt Marsh And Mangrove Systems, Frances R. Griswold

Doctoral Dissertations

Tidal wetlands are vital for buffering coastal settings from the threats of accelerated sea level rise and storms. Understanding the factors that are most influential for the maintenance and recovery of tidal wetlands after extreme events compounded by future accelerated sea level rise is of the utmost importance, yet this knowledge is not well established. Two tidal wetland schemas investigated in this dissertation are mangrove systems in Vieques, Puerto Rico (including robust lagoonal-mangrove forest systems and fringing mangrove forests), and salt marshes in New England. While the climatic forcings, vegetation type, and locations are vastly different for these two tidal …


Competition And Herbivory Influence The Survival, Growth, And Physiology Of Native Tree Seedlings In The Kentucky Inner Bluegrass Savanna-Woodland, James D. Shaffer Jan 2023

Competition And Herbivory Influence The Survival, Growth, And Physiology Of Native Tree Seedlings In The Kentucky Inner Bluegrass Savanna-Woodland, James D. Shaffer

Theses and Dissertations--Biology

Terrestrial plant communities are shaped by competition for resources, herbivory, and abiotic processes. Savanna systems represent a dynamic coexistence of contrasting life forms (grasses and trees) shaped by competition and disturbance. The Kentucky Inner Bluegrass Savanna-Woodland (KIBSW) is described as an open woodland of shade intolerant species; however, climatic, and edaphic conditions can support closed-canopy forest. After European pioneer settlement (c1750-1800), over 99% of “savanna-woodlands” have been lost. KIBSW remnants are experiencing a recruitment failure, leading to a dominance shift in tree communities. I researched how tree-grass competition and mammalian herbivory influence KIBSW regeneration and maintenance. The KIBSW does not …


The Long-Term Effects Of Wildfire Severity On Oak-Pine Communities And Their Microclimates, Scott Glenn Culbert Jan 2023

The Long-Term Effects Of Wildfire Severity On Oak-Pine Communities And Their Microclimates, Scott Glenn Culbert

Theses and Dissertations--Forestry and Natural Resources

In the eastern U.S., fire-dependent tree species have historically dominated upland forest communities, but are now experiencing widespread regeneration challenges as a result of 20th century fire suppression policies, and are being replaced by mesophytic species. Wildfires that contain areas of high burn severity may provide an important means of mitigating these challenges and facilitating fire-dependent species regeneration and recruitment into larger size classes. One mechanism by which high-severity fire can accomplish this is by modifying understory microclimate characteristics to be more conducive to these species’ growth. A wildfire within the Daniel Boone National Forest, Kentucky, USA, provided the …


Fine Characterization Of Leafing Phenology In The Brazilian Atlantic Forest By Optical And Microwave Remote Sensing, James B. Bell Jan 2023

Fine Characterization Of Leafing Phenology In The Brazilian Atlantic Forest By Optical And Microwave Remote Sensing, James B. Bell

Dissertations and Theses

Tropical forests provide important ecosystem functions in the global biosphere, but they remain among the most poorly understood elements of land surface models, especially with regard to their seasonal dynamics. For instance, in seasonally dry forests, the pattern of the annual green-up in their canopies closely follows annual patterns of rainfall. The same, however, does not occur in wet forest canopies which are dominated by evergreen trees. In the latter, water is not scarce enough to limit leaf photosynthetic function. Canopy leafing phenology in these forests is therefore poorly characterized by optical remote sensing methods which are not sensitive to …


Assessing The Recovery Of Forest Understory Vegetation After Clearcut Logging Across A 445-Year Chronosequence, Molly Smith Metok Jan 2023

Assessing The Recovery Of Forest Understory Vegetation After Clearcut Logging Across A 445-Year Chronosequence, Molly Smith Metok

Cal Poly Humboldt theses and projects

The conversion of natural forested lands to managed forests has reduced the amount of older, structurally diverse forests worldwide. In conifer forests of the Pacific Northwest (USA) – where the understory plant communities comprise only 1% of forest biomass but represent 90% of the plant species richness – the long-term impacts of timber harvesting are not fully understood. I used a chronosequence of forests in southwestern Oregon that ranged from 25 to 445 years of age to compare changes in plant communities in logged (i.e., managed) stands with that of stands in late succession and old growth conditions. The chronosequence …


Modeling Growth And Stress Factors For Converted Silvopasture Systems In The Missouri Ozarks, Bailee N. Suedmeyer Jan 2023

Modeling Growth And Stress Factors For Converted Silvopasture Systems In The Missouri Ozarks, Bailee N. Suedmeyer

MSU Graduate Theses

Silvopasture systems are becoming increasingly popular among sustainable agriculture ranchers, due to the increase in knowledge of benefits to the cattle and ability to grow cool season grasses beneath the canopy. This project focuses on the forest crop aspect of silvopasture systems from monitoring of the health of the trees over time to recommendations for thinning management to keep it functioning as viable silvopasture. The study site consists of five acres of upland hardwood forest area in Southern Missouri with 18 monumented fixed area plots. Arial and ground data was collected at each plot throughout the growing season, along with …


Response Of Running Buffalo Clover (Trifolium Stoloniferum Muhl Ex. A. Eaton) To Herbaceous Competition And Transplanting In Monongahela National Forest, Ruben E. Sabella Jan 2023

Response Of Running Buffalo Clover (Trifolium Stoloniferum Muhl Ex. A. Eaton) To Herbaceous Competition And Transplanting In Monongahela National Forest, Ruben E. Sabella

Graduate Theses, Dissertations, and Problem Reports

Running buffalo clover (RBC) is a rare perennial plant that grows throughout the American Midwest and the Appalachian Mountains. It requires disturbed forests to establish and proliferate. It has been suggested that, in the past, these conditions were created by buffalo; now logging operations maintain RBC populations. However, forest managers have been looking for ways to create suitable habitat for RBC that do not involve harvesting practices. This could help create new populations in areas that cannot be logged. Once established, competing vegetation might influence RBC abundance and flowering. This study seeks to quantify this influence by measuring the vegetation …


Knobcone Pine Response To Shading From Competing Chaparral Shrubs Following Stand-Replacing Wildfire, Sean T. Lindley Jan 2023

Knobcone Pine Response To Shading From Competing Chaparral Shrubs Following Stand-Replacing Wildfire, Sean T. Lindley

Cal Poly Humboldt theses and projects

In northern California, fire regimes are shifting towards more frequent and larger severe wildfire. There is growing concern that this shift poses a threat to biodiversity in the form of cover type change at the landscape scale, resulting in the extirpation of some species in favor of +AD617:AD649well-adapted ones. In northern California, mature serotinous conifers, such as knobcone pine (Pinus attenuata), and resprouting shrub species easily regenerate in severe patches of any size. There is no general consensus regarding the effects of shrub competition on conifer recruitment; conifer response varies with shade tolerance and other abiotic factors. Knobcone …