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2005

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A Comparison Of The Growth And Asexual Reproduction By Cryphonectria Parasitica Isolates Infected With Hypoviruses Via Anastomosis And Transfection, Jenise M. Bauman Aug 2005

A Comparison Of The Growth And Asexual Reproduction By Cryphonectria Parasitica Isolates Infected With Hypoviruses Via Anastomosis And Transfection, Jenise M. Bauman

Graduate Theses, Dissertations, and Problem Reports (ETD)

Transfection is an alternative laboratory method of hypovirus transmission that can be used to create hypovirulent strains. Results from a preliminary study at West Virginia University in 1998-99 indicated that recovery of hypovirulent isolates was similar between CHV1-infected anastomosed and transfected isolates. The objective of this research was to compare the two hypovirus acquisition methods in three experiments. The first study confirmed the performance of anastomosed and transfected isolates tested in a 1998-99 field study. In addition to the isolates previously evaluated, a third set of isolates containing CHV3-County Line was included. The size of cankers and the reisolation of …


Evaluating Participants' Perceptions Of Alfabetizacion/Circulo De Aprendizaje, A Literacy Program In Ciudad Eten, Chiclayo-Peru, Veronikha Salazar Aug 2005

Evaluating Participants' Perceptions Of Alfabetizacion/Circulo De Aprendizaje, A Literacy Program In Ciudad Eten, Chiclayo-Peru, Veronikha Salazar

Graduate Theses and Dissertations

The purpose of this study was to evaluate the perceptions of participants regarding the Alfabetizacion/Circulo de Arendizaje literacy program in Ciudad Eten, Chiclayo, Peru.

The sample population for the study consisted of 118 female participants. These women represented participants of the Alfabetizacion literacy program in 2004 and 1994 - 2003 program graduates. The oral survey questions were related to demographic factors of the participants (i.e. age, marital status, family size, level of income and class attendance), factors affecting participants’ perceptions of the importance of education, and reasons for participation in the literacy program. Participants also answered questions regarding their needs …


Armstrong Academics August 2005, Armstrong Atlantic State University Aug 2005

Armstrong Academics August 2005, Armstrong Atlantic State University

Armstrong State University Collections: Armstrong Academics (Archived)

No abstract provided.


A Hydrogeological Perspective Of The Status Of Ground Water Resources Under The Un Watercourse Convention, Gabriel Eckstein Aug 2005

A Hydrogeological Perspective Of The Status Of Ground Water Resources Under The Un Watercourse Convention, Gabriel Eckstein

Faculty Scholarship

When the U.N. General Assembly adopted the Convention on the Non-Navigational Uses of International Watercourses in 1997, it took a decisive step in recognizing the important role that transboundary ground water resources play in human progress and development. In so doing, it also acknowledged the need to establish principles of law governing this "invisible" but valuable natural resource. Transboundary ground water historically has been neglected in treaties, ignored in projects with international implications, and cursorily misunderstood in much of legal discourse.

While the Convention provides substantial clarification on the status of ground water under international law, it also leaves considerable …


47th Rocky Mountain Conference On Analytical Chemistry Jul 2005

47th Rocky Mountain Conference On Analytical Chemistry

Rocky Mountain Conference on Magnetic Resonance

Final program, abstracts, and information about the 47th annual meeting of the Rocky Mountain Conference on Analytical Chemistry, co-endorsed by the Colorado Section of the American Chemical Society and the Rocky Mountain Section of the Society for Applied Spectroscopy. Held in Denver, Colorado, July 31 - August 4, 2005.


College Of Agricultural Sciences And Natural Resources: 12th Annual Report August 1, 2004-July 31, 2005 Jul 2005

College Of Agricultural Sciences And Natural Resources: 12th Annual Report August 1, 2004-July 31, 2005

College of Agricultural Sciences and Natural Resources (CASNR): Annual Reports

It is our pleasure to provide the Twelfth Annual Report for the College of Agricultural Sciences and Natural Resources (CASNR). We believe in the importance of reflecting on the accomplishments of our faculty, staff and students. The annual report offers the opportunity to review the year, and serves as an archive of accomplishments for our College. This report covers the period August 1, 2004 to July 31, 2005.

The college enjoyed another year of accomplishments and change. The students excelled, the faculty’s enthusiasm for the academic mission continued to thrive and campus programming expanded. Clearly, the most visible activity this …


Distribution Patterns Of Lentic-Breeding Amphibians In Relation To Ultraviolet Radiation Exposure In Western North America, Michael J. Adams, Blake R. Hossack, Roland A. Knapp, Paul Stephen Corn, Stephen A. Diamond, Peter C. Trenham, Dan B. Fagre Jul 2005

Distribution Patterns Of Lentic-Breeding Amphibians In Relation To Ultraviolet Radiation Exposure In Western North America, Michael J. Adams, Blake R. Hossack, Roland A. Knapp, Paul Stephen Corn, Stephen A. Diamond, Peter C. Trenham, Dan B. Fagre

United States Geological Survey: Staff Publications

An increase in ultraviolet-B (UV-B) radiation has been posited to be a potential factor in the decline of some amphibian population. This hypothesis has received support from laboratory and field experiments showing that current levels of UV-B can cause embryo mortality in some species, but little research has addressed whether UV-B is influencing the distribution of amphibian populations. We compared patterns of amphibian presence to sitespecific estimates of UV-B dose at 683 ponds and lakes in Glacier, Olympic, and Sequoia–Kings Canyon National Parks. All three parks are located in western North America, a region with a concentration of documented amphibian …


Estimated Ultraviolet Radiation Doses In Wetlands In Six National Parks, Stephen A. Diamond, Peter C. Trenham, Michael J. Adams, Blake R. Hossack, Roland A. Knapp, Stacey L. Stark, David Bradford, P. Stephen Corn, Ken Czarnowski, Paul D. Brooks, Dan Fagre, Bob Breen, Naomi E. Detenbeck, Kathy Tonnessen Jul 2005

Estimated Ultraviolet Radiation Doses In Wetlands In Six National Parks, Stephen A. Diamond, Peter C. Trenham, Michael J. Adams, Blake R. Hossack, Roland A. Knapp, Stacey L. Stark, David Bradford, P. Stephen Corn, Ken Czarnowski, Paul D. Brooks, Dan Fagre, Bob Breen, Naomi E. Detenbeck, Kathy Tonnessen

United States Geological Survey: Staff Publications

Ultraviolet-B radiation (UV-B, 280–320-nm wavelengths) doses were estimated for 1024 wetlands in six national parks: Acadia (Acadia), Glacier (Glacier), Great Smoky Mountains (Smoky), Olympic (Olympic), Rocky Mountain (Rocky), and Sequoia/ Kings Canyon (Sequoia). Estimates were made using ground-based UV-B data (Brewer spectrophotometers), solar radiation models, GIS tools, field characterization of vegetative features, and quantification of DOC concentration and spectral absorbance. UV-B dose estimates were made for the summer solstice, at a depth of 1 cm in each wetland. The mean dose across all wetlands and parks was 19.3 W-h m-2 (range of 3.4–32.1 W-h m-2). The mean …


Species Limits Within The Mexican Garter Snakes Of The Thamnophis Godmani Complex, Douglas A. Rossman, Frank T. Burbrink Jul 2005

Species Limits Within The Mexican Garter Snakes Of The Thamnophis Godmani Complex, Douglas A. Rossman, Frank T. Burbrink

Occasional Papers of the Museum of Natural Science, Louisiana State University

The highly variable Thamniphis godmani complex of southern Mexico, comprised of four apparently allopatric populations, was examined by using 28 morphological characters scored on 214 specimens. Character values were tested with univariate and multivariate statistics to determine if populations are morphologically distinct. The results suggest that the four populations represent independently evolving lineages. Four species, three previously undescribed, are recognized in the T. godmani complex. A lectotype is designated for T. godmani. For each species, a diagnosis description of holotype or lectotype, summary of interspecific variation, and statement of distribution are provided. Comparisons are made with T. errans, alleged to …


2005 Summer Commencement, Harding University Jul 2005

2005 Summer Commencement, Harding University

Commencement Programs

Program for the Commencement on July 30.

Commencement Speaker: Charles W. Bradley


Loss Of Outer Membrane Proteins Without Inhibition Of Lipid Export In An Escherichia Coli Yaet Mutant, William T. Doerrler, Christian R.H. Raetz Jul 2005

Loss Of Outer Membrane Proteins Without Inhibition Of Lipid Export In An Escherichia Coli Yaet Mutant, William T. Doerrler, Christian R.H. Raetz

Faculty Publications

Escherichia coli yaeT encodes an essential, conserved outer membrane (OM) protein that is an ortholog of Neisseria meningitidis Omp85. Conflicting data with TV. meningitidis indicate that Omp85 functions either in assembly of OM proteins or in export of OM lipids. The role of YaeT in E. coli was investigated with a new temperature-sensitive mutant harboring nine amino acid substitutions. The mutant stops growing after 60 min at 44 °C. After 30 min at 44 °C, incorporation of [35S]methionine into newly synthesized OM proteins is selectively inhibited. Synthesis and export of OM phospholipids and lipopolysaccharide are not impaired. OM protein levels …


A Role For Mitogen-Activated Protein Kinase (Erk1/2) Activation And Non-Selective Pore Formation In P2x7 Receptor-Mediated Thymocyte Death, Rodolphe Auger, Iris Motta, Karim Benihoud, David M. Ojcius, Jean M. Kanellopoulos Jul 2005

A Role For Mitogen-Activated Protein Kinase (Erk1/2) Activation And Non-Selective Pore Formation In P2x7 Receptor-Mediated Thymocyte Death, Rodolphe Auger, Iris Motta, Karim Benihoud, David M. Ojcius, Jean M. Kanellopoulos

All Dugoni School of Dentistry Faculty Articles

Extracellular ATP (ATPe) binds to P2X7 receptors (P2X7R) expressed on the surface of cells of hematopoietic lineage, including murine thymocytes. Activation of P2X7R by ATPe results in the opening of cation-specific channels, and prolonged ATPe exposure leads to the formation of non-selective pores enabling transmembrane passage of solutes up to 900 Da. In the presence of ATPe, P2X7R-mediated thymocyte death is due primarily to necrosis/lysis and not apoptosis, as measured by the release of lactate dehydrogenase indicative of a loss of plasma membrane integrity. The present study is focused on the identification of P2X7R signaling mediators in ATP-induced thymocyte necrosis/lysis. …


A Vascular Plant Inventory And Description Of The Twelve Plant Community Types Found In The University Of South Florida Ecological Research Area, Hillsborough County, Florida, Anne Candace Schmidt Jul 2005

A Vascular Plant Inventory And Description Of The Twelve Plant Community Types Found In The University Of South Florida Ecological Research Area, Hillsborough County, Florida, Anne Candace Schmidt

USF Tampa Graduate Theses and Dissertations

The University of South Florida Ecological Research Area (USF Eco Area), located in west central Hillsborough County, is an approximately 306 hectare (756 acre) natural area on the Hillsborough River composed of twelve plant communities. While surrounded on three sides by urbanization, the USF Eco Area makes up the western most section of an extended natural corridor that runs approximately 88 kilometers (55 miles) east and north along the Hillsborough River. An inventory of the vascular flora and the associated ecological communities was developed to better assess the USF Eco Area for educational and research purposes as well as enhance …


Test Statistics Null Distributions In Multiple Testing: Simulation Studies And Applications To Genomics, Katherine S. Pollard, Merrill D. Birkner, Mark J. Van Der Laan, Sandrine Dudoit Jul 2005

Test Statistics Null Distributions In Multiple Testing: Simulation Studies And Applications To Genomics, Katherine S. Pollard, Merrill D. Birkner, Mark J. Van Der Laan, Sandrine Dudoit

U.C. Berkeley Division of Biostatistics Working Paper Series

Multiple hypothesis testing problems arise frequently in biomedical and genomic research, for instance, when identifying differentially expressed or co-expressed genes in microarray experiments. We have developed generally applicable resampling-based single-step and stepwise multiple testing procedures (MTP) for control of a broad class of Type I error rates, defined as tail probabilities and expected values for arbitrary functions of the numbers of false positives and rejected hypotheses (Dudoit and van der Laan, 2005; Dudoit et al., 2004a,b; Pollard and van der Laan, 2004; van der Laan et al., 2005, 2004a,b). As argued in the early article of Pollard and van der …


Electromechanical Imaging Of Biological Systems With Sub-10 Nm Resolution, Sergei V. Kalinin, B. J. Rodriguez, Stephen Jesse, T. Thundat, Alexei Gruverman Jul 2005

Electromechanical Imaging Of Biological Systems With Sub-10 Nm Resolution, Sergei V. Kalinin, B. J. Rodriguez, Stephen Jesse, T. Thundat, Alexei Gruverman

Alexei Gruverman Publications

Electromechanical imaging of tooth dentin and enamel has been performed with sub-10 nm resolution using piezoresponse force microscopy. Characteristic piezoelectric domain size and local protein fiber ordering in dentin have been determined. The shape of a single protein fibril in enamel is visualized in real space and local hysteresis loops are measured. Because of the ubiquitous presence of piezoelectricity in biological systems, this approach is expected to find broad application in high-resolution studies of a wide range of biomaterials.


Lack Of Association Of Methylenetetrahydrofolate Reductase 677c>T Mutation With Coronary Artery Disease In A Pakistani Population, M Perwaiz Iqbal, Tasneem Fatima, Siddiqa Parveen, Farzana A. Yousuf, Majid Shafiq, Naseema Mehboobali, Abrar H. Khan, Iqbal Azam, Philippe M. Frossard Jul 2005

Lack Of Association Of Methylenetetrahydrofolate Reductase 677c>T Mutation With Coronary Artery Disease In A Pakistani Population, M Perwaiz Iqbal, Tasneem Fatima, Siddiqa Parveen, Farzana A. Yousuf, Majid Shafiq, Naseema Mehboobali, Abrar H. Khan, Iqbal Azam, Philippe M. Frossard

Department of Biological & Biomedical Sciences

Pakistanis belong to the South Asian population which has the highest known rate of coronary artery disease. Folic acid deficiency also appears to be highly prevalent in this population. Methylenetetrahydrofolate reductase (MTHFR) 677C>T polymorphism decreases the activity of this enzyme and can be associated with mild to moderate hyperhomocysteinemia in homozygotes, particularly when there is folic acid deficiency, as well as with coronary artery disease. To assess the value of genotyping the MTHFR 677C>T dimorphism, we carried out a case-control study of dimorphism 677C>T for putative association with myocardial infarction (MI) among Pakistani nationals. We investigated a …


Unh Fisheries Expert Says Stay The Course With Ocean Fish Recovery Policy In Science Article, David Sims Jul 2005

Unh Fisheries Expert Says Stay The Course With Ocean Fish Recovery Policy In Science Article, David Sims

Media Relations

No abstract provided.


Comparing The Effects Of Granivorous Rodents On Persistence Of Indian Ricegrass (Oryzopsis Hymenoides) Seeds In Mixed And Monospecific Seed Patches, Joseph A. Veech, Stephen H. Jenkins Jul 2005

Comparing The Effects Of Granivorous Rodents On Persistence Of Indian Ricegrass (Oryzopsis Hymenoides) Seeds In Mixed And Monospecific Seed Patches, Joseph A. Veech, Stephen H. Jenkins

Western North American Naturalist

In desert environments seeds are often heterogeneously distributed in small patches that vary in number of seed species and in seed density. Because seed harvest by rodents is often density dependent (a larger proportion of seeds is removed from high-density seed patches than from low-density patches), the proportion of residual or post-harvest seeds should be greater in low-density patches. In addition, seed preference can affect harvest. We tested whether the residual proportion of a highly preferred seed (Indian ricegrass, Oryzopsis hymenoides) was less when in a seed patch with a 2nd species (mixed-species patch) than when in a monospecific …


Pollination Needs Of Arrowleaf Balsamroot, Balsamorhiza Sagittata (Heliantheae: Asteraceae), James H. Cane Jul 2005

Pollination Needs Of Arrowleaf Balsamroot, Balsamorhiza Sagittata (Heliantheae: Asteraceae), James H. Cane

Western North American Naturalist

Arrowleaf balsamroot, Balsamorhiza sagittata (Pursh) Nutt, is a common, sometimes dominant, long-lived forb that flowers early in spring from the foothills to upper-montane areas of the northern Rocky Mountains and Intermountain West. Public land managers desire its seed for rangeland rehabilitation. Through manual pollination field trials, the species was found to have a mixed pollination system. It is primarily xenogamous (46% of ovules yielded plump achenes) but partially self-compatible (31% of achenes were plump). Unvisited flower heads formed virtually no mature achenes; only plump achenes contained seeds with endosperm. Freely visited flower heads in 2 populations produced as many achenes …


New Species Of Mentzelia (Loasaceae) And Phacelia (Hydrophyllaceae) From New Mexico, N. Duane Atwood, Stanley L. Welsh Jul 2005

New Species Of Mentzelia (Loasaceae) And Phacelia (Hydrophyllaceae) From New Mexico, N. Duane Atwood, Stanley L. Welsh

Western North American Naturalist

New species of Mentzelia and Phacelia are described from northwestern New Mexico, USA, as Mentzelia todiltoensis, N. Duane Atwood & Stanley L. Welsh, and Phacelia sivinskii N. D. Atwood, sp. nov. The new Mentzelia is compared to M. humilis based on similarities in seed, fruit, leaf, and flower morphology and to M. memorabalis in stem, leaf, and fruit morphology. Phacelia sivinskii is compared to P. constancei N. D. Atwood on the basis of seed, leaf, and flower morphology.


Monitoring Temporal Change In Riparian Vegetation Of Great Basin National Park, Erik A. Beever, David A. Pyke, Jeanne C. Chambers, Fred Landau, Stanley D. Smith Jul 2005

Monitoring Temporal Change In Riparian Vegetation Of Great Basin National Park, Erik A. Beever, David A. Pyke, Jeanne C. Chambers, Fred Landau, Stanley D. Smith

Western North American Naturalist

Disturbance in riparian areas of semiarid ecosystems involves complex interactions of pulsed hydrologic flows, herbivory, fire, climatic effects, and anthropogenic influences. We resampled riparian vegetation within ten 10-m × 100-m plots that were initially sampled in 1992 in 4 watersheds of the Snake Range, east central Nevada. Our finding of significantly lower coverage of grasses, forbs, and shrubs within plots in 2001 compared with 1992 was not consistent with the management decision to remove livestock grazing from the watersheds in 1999. Change over time in cover of life-forms or bare ground was not predicted by scat counts within plots in …


Blue Tilapia (Oreochromis Aureus) Predation On Fishes In The Muddy River System, Clark County, Nevada, G. Gary Scoppettone, J. Antonio Salgado, M. Bridget Nielsen Jul 2005

Blue Tilapia (Oreochromis Aureus) Predation On Fishes In The Muddy River System, Clark County, Nevada, G. Gary Scoppettone, J. Antonio Salgado, M. Bridget Nielsen

Western North American Naturalist

No abstract provided.


Inland Fishes Of California By Peter B. Moyle, Andrew L. Sheldon Jul 2005

Inland Fishes Of California By Peter B. Moyle, Andrew L. Sheldon

Western North American Naturalist

No abstract provided.


Variation In Lewisia Kelloggii (Portulacaceae) With Description Of A New Species Endemic To Idaho, Barbara L. Wilson, Valerie D. Hipkins, Edna Rey-Vizgirdas, Thomas N. Kaye Jul 2005

Variation In Lewisia Kelloggii (Portulacaceae) With Description Of A New Species Endemic To Idaho, Barbara L. Wilson, Valerie D. Hipkins, Edna Rey-Vizgirdas, Thomas N. Kaye

Western North American Naturalist

Lewisia kelloggii has been understood as a rare plant with a disjunct range in California and Idaho. Examination of herbarium specimens and analysis of isozymes in 6 Idaho and 7 California populations revealed consistent differences between plants of the 2 states. Fixed differences in alleles at 2 loci (AAT2 and PGI1) distinguished Idaho from California plants. Genetic identities based on isozymes between Idaho and California populations averaged 0.58, lower than the average for congeneric plant species. Idaho plants were smaller than most California plants, but California plants were variable. The most consistent morphological difference between Idaho and California specimens was …


Westward Expansion Of The Eastern Pipistrelle (Pipistrellus Subflavus) In The United States, Including New Records From New Mexico, South Dakota, And Texas, Keith Geluso, Tony R. Mollhagen, Joel M. Tigner, Michael A. Bogan Jul 2005

Westward Expansion Of The Eastern Pipistrelle (Pipistrellus Subflavus) In The United States, Including New Records From New Mexico, South Dakota, And Texas, Keith Geluso, Tony R. Mollhagen, Joel M. Tigner, Michael A. Bogan

Western North American Naturalist

No abstract provided.


Evaluating Lek Occupancy Of Greater Sage-Grouse In Relation To Landscape Cultivation In The Dakotas, Joe T. Smith, Lester D. Flake, Kenneth F. Higgins, Gerald D. Kobriger, Collin G. Homer Jul 2005

Evaluating Lek Occupancy Of Greater Sage-Grouse In Relation To Landscape Cultivation In The Dakotas, Joe T. Smith, Lester D. Flake, Kenneth F. Higgins, Gerald D. Kobriger, Collin G. Homer

Western North American Naturalist

Greater Sage-Grouse (Centrocercus urophasianus) have been declining in many states and provinces of North America, and North and South Dakota hold no exception to these declines. We studied effects of cultivated land on Greater Sage-Grouse lek abandonment in North and South Dakota. Landscape-level data were assessed using satellite imagery within a geographic information system. Comparisons were made of 1972–1976 and 1999–2000 percent cultivated and noncultivated land. These comparisons were made between land uses surrounding active leks versus inactive leks, active leks versus random locations, and abandoned regions versus active regions. The 1999–2000 imagery illustrated that percent cultivated land …


Archaeological Record Of Native Fishes Of The Lower Colorado River: How To Identify Their Remains, Kenneth W. Gobalet, Thomas A. Wake, Kalie L. Hardin Jul 2005

Archaeological Record Of Native Fishes Of The Lower Colorado River: How To Identify Their Remains, Kenneth W. Gobalet, Thomas A. Wake, Kalie L. Hardin

Western North American Naturalist

Archaeological sites in the Salton Basin of southeastern California and along the lower Colorado River provided opportunities to determine which fish species were present prior to extirpations, environmental degradation, and the recession of Lake Cahuilla. These remains also represent the fishes exploited by Native Americans. Bonytail (Gila elegans), razorback sucker (Xyrauchen texanus), Colorado pikeminnow (Ptychocheilus lucius), striped mullet (Mugil cephalus), and machete (Elops affinis) have been recovered from 117 sites in the Salton Basin, once filled by the Colorado River forming Lake Cahuilla. Bonytail and razorback sucker comprise nearly 99% …


Modeling Global Warming Scenarios In Greenback Cutthroat Trout (Oncorhynchus Clarki Stomias) Streams: Implications For Species Recovery, Scott J. Cooney, Alan P. Covich, Paul M. Lukacs, Amy L. Harig, Kurt D. Fausch Jul 2005

Modeling Global Warming Scenarios In Greenback Cutthroat Trout (Oncorhynchus Clarki Stomias) Streams: Implications For Species Recovery, Scott J. Cooney, Alan P. Covich, Paul M. Lukacs, Amy L. Harig, Kurt D. Fausch

Western North American Naturalist

Changes in global climate may exacerbate other anthropogenic stressors, accelerating the decline in distribution and abundance of rare species throughout the world. We examined the potential effects of a warming climate on the greenback cutthroat trout (Oncorhynchus clarki stomias), a resident salmonid that inhabits headwater streams of the central Rocky Mountains. Greenbacks are outcompeted at lower elevations by nonnative species of trout and currently are restricted to upper-elevation habitats where barriers to upstream migration by nonnatives are or have been established. We used likelihood-based techniques and information theoretics to select models predicting stream temperature changes for 10 streams …


Collection Of Asian Tapeworm (Bothriocephalus Acheilognathi) From The Yampa River, Colorado, David Ward Jul 2005

Collection Of Asian Tapeworm (Bothriocephalus Acheilognathi) From The Yampa River, Colorado, David Ward

Western North American Naturalist

No abstract provided.


Characterizing Sound Production In Nearshore Rockfishes (Sebastes Spp.), Bryan Nichols Jul 2005

Characterizing Sound Production In Nearshore Rockfishes (Sebastes Spp.), Bryan Nichols

USF Tampa Graduate Theses and Dissertations

Rockfishes of the genus Sebastes are trophically important in most nearshore environments of the west coast of North America, and support important commercial and recreational fisheries. Though the 72 northeast Pacific species have drumming muscles along their swim bladders, little is known about sound production in the genus. Sounds produced by nearshore rockfish were recorded using fixed hydrophones and underwater video in aquaria, and acoustical dataloggers in the field. Sounds were analyzed from six species: Sebastes nebulosus, S. atrovirens, S.carnatus, S. chrysomelas, S. caurinus, and S.maliger. These six species are closely related, mostly bottom …