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When The Desert Meets The Classroom: A Mixed Methods Study On Experiential Learning And Andragogy For Design-Situated Curricula, Julie K. Lamarra
When The Desert Meets The Classroom: A Mixed Methods Study On Experiential Learning And Andragogy For Design-Situated Curricula, Julie K. Lamarra
All Graduate Theses and Dissertations, Spring 1920 to Summer 2023
This study’s significance examined how outdoor experience in place can influence students’ designs in the context of outdoor recreation. Historically, experiential education and outdoor education have been used widely across curricula but are not so commonly accepted in design education except in the form of internships. This study sought to examine how being in the place that activity occurs and learning about the cultural aspects of place, activity, and community impacted students’ designs.
This study looked at how existing theories such as Landscapes of Practice and the development of boundaries, knowledge, and identity are formed within communities of practice and …
Computational Techniques For Elucidating Plant-Pathogen Interactions: A Case Study On Citrus-Hlb Interactome, Cristian D. Loaiza
Computational Techniques For Elucidating Plant-Pathogen Interactions: A Case Study On Citrus-Hlb Interactome, Cristian D. Loaiza
All Graduate Theses and Dissertations, Spring 1920 to Summer 2023
The Citrus fruit industry in the United States has been affected during the last two decades because of the outbreak of the citrus greening disease, also known as Huanglongbing (HLB). Many people and organizations are working in therapeutics to help mitigate the impact of this disease, unfortunately there is not a cure yet. There are many mechanisms that needs to be understood about this disease, especially at the molecular level. Not only HLB, but many other infectious diseases are controlled by the interaction of proteins from the host (Citrus in this case) and from the pathogen that causes the disease. …
Local And Systemic Effects Of Flavonoid Intake With Submaximal Cycling Exercise: A Randomized Controlled Trial, Stephanie Kung
Local And Systemic Effects Of Flavonoid Intake With Submaximal Cycling Exercise: A Randomized Controlled Trial, Stephanie Kung
All Graduate Theses and Dissertations, Spring 1920 to Summer 2023
Moderate to high intensity endurance exercise is a known cause of exercise-induced gastrointestinal syndrome, a condition often associated with intestinal complaints such as discomfort, cramps, and diarrhea. Oftentimes, elevated intestinal inflammation and permeability (a “leaky gut”) are also observed. Previous research has shown that flavonoids, natural compounds found in many fruits and vegetables, may mitigate these exercise-induced effects. In particular, certain cocoa powders contain high levels of flavonoids, and chocolate milk is a good source of protein and sugars for sustaining intense activity. Thus, the purpose of this study was to evaluate the effects of a dairy pre-workout beverage, with …
Seeding Rate, Nitrogen Fertilzer, And Cutting Timing Effects On Teff Forage Yield And Nutritive Value, Michael C. Laca
Seeding Rate, Nitrogen Fertilzer, And Cutting Timing Effects On Teff Forage Yield And Nutritive Value, Michael C. Laca
All Graduate Theses and Dissertations, Spring 1920 to Summer 2023
Teff [Eragostis tef (Zucc.) Trotter] is a relatively new forage gaining popularity in the United States; however, information regarding agronomic production practices is lacking. This study was conducted to determine the combination of seeding rate, fertilization, and harvest timing to optimize teff dry-matter yield (DMY) and nutritive value. Four seeding rates (2, 5, 8, and 11 lb/acre), four nitrogen (N) fertilizer rates (0, 25, 50, and 100 lb/acre), and two harvest strategies [boot stage (2-cut) and full seed-head stage (1-cut; stockpiled)] were evaluated in 2010 and 2011 in Kaysville, UT and Yerington, NV. The effects of harvest (1- vs …
Alienation, Modernization, And Animal Welfare: Human-Animal Relationships At The Farm, State, And Country Levels, Michael D. Briscoe
Alienation, Modernization, And Animal Welfare: Human-Animal Relationships At The Farm, State, And Country Levels, Michael D. Briscoe
All Graduate Theses and Dissertations, Spring 1920 to Summer 2023
Our relationships with animals are important for us as humans, for the environment, and for the animals themselves. In this dissertation I look at the relationships between humans and animals at three scales: farm, U.S. state, and country. Specifically, I address how factors like economic growth, technological innovation, and globalization affect human relationships with animals. Understanding how these factors influence human-animal relationships is important for improving these relationships and deciding which directions will most contribute to sustainable outcomes. I address the social factors that influence human-animal relationships in three studies.
In the first study I surveyed and interviewed dairy farmers …
Characterization And Immobilization Of Thermophilic Lipases For The Interesterification Of Soybean Oil, Teif A. Najm
Characterization And Immobilization Of Thermophilic Lipases For The Interesterification Of Soybean Oil, Teif A. Najm
All Graduate Theses and Dissertations, Spring 1920 to Summer 2023
Lipases are currently used in food technology for the modification of fats and oils. Lipase can synthesize, hydrolyze, and rearrange fatty acids, thus, resulting in their extensive use in industrial applications. The thermal stability of lipase is an essential characteristic for this application. Thermophilic bacteria are an important source for producing thermostable lipases because of their tolerance to high heat. Lipolytic enzymes can be purified using several purification methods, but these purification processes are often expensive and complex. Results from this study showed that lipases can be extract from thermophilic bacteria (Geobacillius stearothermophilus (GS) and Anoxybacillus flavithermus (AF)). Molecular …
A Phenological Model For A Southern Population Of Mountain Pine Beetle, Catherine E. Wangen
A Phenological Model For A Southern Population Of Mountain Pine Beetle, Catherine E. Wangen
All Graduate Theses and Dissertations, Spring 1920 to Summer 2023
The mountain pine beetle (MPB, Dendroctonus ponderosae Hopkins) attacks living Pinus trees across a widespread area of western North America, causing significant ecological and economic damage. The ability to make accurate predictions of how MPB populations across this range will respond to temperatures, which affect MPB progress through life stages, is essential. Northern and southern populations of MPB are genetically different in response to temperature, requiring geographic-specific model parameters. There is not currently a predictive model for the southern MPB life cycle, despite concerns that those populations may be more susceptible to increased numbers of generations per year, which would …
Generation Of Sheep Models Of Cystic Fibrosis By Gene Editing And Cloning, Iuri Viótti Périssé
Generation Of Sheep Models Of Cystic Fibrosis By Gene Editing And Cloning, Iuri Viótti Périssé
All Graduate Theses and Dissertations, Spring 1920 to Summer 2023
Cystic Fibrosis (CF) is a genetic disease that affects over 80,000 people worldwide and is caused by mutations that disrupt the Cystic Fibrosis Transmembrane Conductance Regulator (CFTR) gene. This gene encodes a channel protein that regulates ion transport in cells. The most common mutation in the human CFTR gene is F508del, which is characterized by the deletion of a phenylalanine amino acid in the CFTR protein. The second most common mutation is G542X, a protein synthesis “stop” mutation that prevents the production of the CFTR channel protein. An impediment that prevents the further understanding CF is the limited …
An Empirical And Theoretical Investigation Of Random Reinforced Forests And Shallow Convolutional Neural Networks, Nikhil Ganta
An Empirical And Theoretical Investigation Of Random Reinforced Forests And Shallow Convolutional Neural Networks, Nikhil Ganta
All Graduate Theses and Dissertations, Spring 1920 to Summer 2023
For many years, the global population of honey bees has been decreasing due to inconclusive reasons resulting in the syndrome Colony Collapse Disorder (CCD). This syndrome has been plaguing bees and affecting commercial agriculture pollination since 1998. Many researchers have suggested that pesticides, in-hive chemicals, pathogens, etc., might be the causes of CCD. Researchers also believe that any changes in a beehive can disturb the bees, which may negatively affect their health. Honey bees are the most vital among all the animal pollinators contributing to approximately 30% of the world’s commercial pollination services. As they are of keystone importance to …
The Fate And Cycling Of Nitrogen, Phosphorous, And Trace Heavy Metals In Beaver-Altered Headwater Streams, Desneiges S. Murray
The Fate And Cycling Of Nitrogen, Phosphorous, And Trace Heavy Metals In Beaver-Altered Headwater Streams, Desneiges S. Murray
All Graduate Theses and Dissertations, Spring 1920 to Summer 2023
Human land-use can increase the amount of non-point source (NPS) pollution in a stream, negatively affecting ecosystem health and beneficial services provided by an ecosystem. Unfortunately, NPS pollution remains high in many waterbodies. Beaver dams may be a passive, cost-effective strategy for removing NPS pollution in headwater streams because beaver dams slow stream flow and collect sediments. Impounded sediments can change how nutrients and pollutants are cycled in a stream through multiple pathways. In the first part of our study, we investigated whether beaver activity can reduce nitrogen, phosphorous and heavy metals from otherwise traveling downstream. Results suggest beaver ponds …
Fatty Acid Composition Of Forages And Their Effect On The Fatty Acid Composition In Beef Cattle, Britney G. Allen
Fatty Acid Composition Of Forages And Their Effect On The Fatty Acid Composition In Beef Cattle, Britney G. Allen
All Graduate Theses and Dissertations, Spring 1920 to Summer 2023
This study focused on how diet changes the rumen microbiome in the cattle and the effects of that on the long chain fatty acids (LCFA) by microbes in the rumen, and on the short chain fatty acids (SCFA) these microbes produce from feed carbohydrates like fiber and starch. The abundance of bacteria belonging to the phyla Tenericutes and Proteobacteria increased in response to high-fiber or high-starch diets, respectively. The production of two SCFA was positively correlated with the presence of increased Tenericutes (acetate) and Proteobacteria (propionate). A greater acetate to propionate ratio is associated with elevated production of enteric methane …
Digital Scripture: An Investigation Of The Design And Use Of A Mobile Application For Reading Sacred Text, Neil Carpenter
Digital Scripture: An Investigation Of The Design And Use Of A Mobile Application For Reading Sacred Text, Neil Carpenter
All Graduate Theses and Dissertations, Spring 1920 to Summer 2023
Digital sacred text reading is rapidly growing as digital devices such as mobile smartphones are becoming more common across the globe. Although sacred text can have strong influence on identify and behavior, the effects of a digital revolution on scripture reading practices are not well understood. In particular, current research literature indicates that more information is needed about the design and use of digital sacred text applications (apps) such as mobile Bibles across different religious groups or cultures. Therefore, this study builds upon and extends previous work to analyze a religious text app, Gospel Library, which is designed and largely …
Characterizing Oligomeric State Changes Of Protein Arginine Methyltransferase 1 As A Mechanism For Regulating Activity, Ariana N. Ortolano
Characterizing Oligomeric State Changes Of Protein Arginine Methyltransferase 1 As A Mechanism For Regulating Activity, Ariana N. Ortolano
All Graduate Theses and Dissertations, Spring 1920 to Summer 2023
If you imagined a living cell as a factory, enzymes would be the workers with each individually performing a very specific and important job. If a job is not done correctly it could be detrimental to the performance of assembly lines and devastate a businesses’ existence. For enzymes, their jobs occur on a biological level by catalyzing reactions inside the cell, which can include altering another protein and the pathway it is involved in. Protein arginine methyltransferases, or PRMTs, is an example of an enzyme that modifies protein substrates. It does this by adding one or two methyl groups onto …
Effects Of Appetitive, Pre-Exposed, And Novel Distracters On Interval Timing Behavior: Role Of Nomifensine Within The Prelimbic Cortex, Alexander R. Matthews
Effects Of Appetitive, Pre-Exposed, And Novel Distracters On Interval Timing Behavior: Role Of Nomifensine Within The Prelimbic Cortex, Alexander R. Matthews
All Graduate Theses and Dissertations, Spring 1920 to Summer 2023
Poor attentional focus harms the estimation of time within the seconds-to-minutes range, i.e. interval timing. Interval timing behavior is related to several cognitive functions including perception of the passage of time, decision making, and to memory processes. Harmful delays in timing are caused by distractions of attention, within the timing task in the form of novel, or familiar distracters, which may or may not have been previously conditioned. The purpose of this dissertation was to explore how attention to novel or familiar distractions within the timing task can be altered using catecholamine reuptake blocker nomifensine. The experiment in Chapter 3, …
Visitor Perceptions And Resource Conditions Of Campsites In Two Coastal Alaskan National Parks, Shannon T. Wesstrom
Visitor Perceptions And Resource Conditions Of Campsites In Two Coastal Alaskan National Parks, Shannon T. Wesstrom
All Graduate Theses and Dissertations, Spring 1920 to Summer 2023
Increasing visitation in parks and protected areas presents managers with the challenge of providing quality visitor experiences while mitigating ecological impacts from recreation. Successful management strategies often suggest determining desired conditions for visitor experiences and ecological conditions to establish thresholds. These thresholds can then be compared to existing conditions in order to determine if changes in management strategies should be made. By integrating visitor survey results with ecological assessments, this research is a unique coastal Alaskan regional analysis of the three components of a management framework: 1. Establishes visitor determined thresholds of acceptability for crowding and coastal resource conditions in …
Evaluating Planning And Management Of Natural Resources Within The Utah State Park System, Erin K. Mann
Evaluating Planning And Management Of Natural Resources Within The Utah State Park System, Erin K. Mann
All Graduate Theses and Dissertations, Spring 1920 to Summer 2023
A change in management style, combined with increased visitation and the advanced age of guiding documents used to describe conditions and goals (known as Resource Management Plans) in Utah’s State Parks, are leading park managers to potentially not fulfill the objectives stated in their Resource Management Plans. Using a research tool known as “content analysis,” we analyzed the Research Management Plans of ten sample parks and found that written plans had strong priorities and goals regarding the care and management of natural resources. Significant findings included strong emphasis on resource protection, data collection, and research. Pre-written questions were asked of …
The Effects Of Cannabinoids On Stress-Coping Behaviors And Neuroendocrinological Measures In Chronic Unpredictable Stress: A Preclinical Systematic Review And Meta-Analysis, Noa Reuveni
All Graduate Theses and Dissertations, Spring 1920 to Summer 2023
It is unclear if cannabis helps individuals cope with life’s stressors or increases the likelihood of developing stress-related disorders such as depression. Chronic unpredictable stress is a validated model of depression that exposes rodents to a series of frequently-occurring, intermittent stressors over the course of a few weeks. This systematic review and meta-analysis summarizes existing research using rodents on the effects of cannabinoids on stress-coping behaviors and neuroendocrinological measurements in chronic unpredictable stress. Databases were systematically searched for articles that met inclusion criteria, and effect sizes between control chronic unpredictable stress groups and cannabinoid-treated chronic unpredictable stress groups were calculated. …
Patterns Of Post-Fire Aspen Seedling Establishment, Growth, And Mortality In The Western United States, Mark Regier Kreider
Patterns Of Post-Fire Aspen Seedling Establishment, Growth, And Mortality In The Western United States, Mark Regier Kreider
All Graduate Theses and Dissertations, Spring 1920 to Summer 2023
Sexual seedling establishment in aspen is increasingly recognized as an important natural regeneration pathway for the species in the western U.S. However, information on seedling abundance as well as factors influencing aspen sexual regeneration is limited and frequently anecdotal, due to historical assumptions of seedling rarity as well as difficulty identifying sexual seedlings from asexual aspen sucker regeneration. This thesis contributes to the field of aspen seedling ecology in three major ways. Chapter 1 utilizes historical aspen seedling occurrences in the western U.S. and a systematic field survey of 2018 fire footprints to explore patterns and test assumptions of aspen …
Mechanisms Of Overyielding And Coexistence In Diverse Tallgrass Prairie Communities, Leslie E. Forero
Mechanisms Of Overyielding And Coexistence In Diverse Tallgrass Prairie Communities, Leslie E. Forero
All Graduate Theses and Dissertations, Spring 1920 to Summer 2023
Plants compete for the same basic nutrient and water resources. According to the competitive exclusion principle, when a substantial overlap in resource pools exists, the best competitor for resources should drive all other species to extinction. The ability for plants to coexist in violation of the competitive exclusion principle is the “biodiversity paradox”. Coexistence is actually beneficial for plants: as species diversity increases, you typically see increases in plant biomass production (known as the biodiversity-productivity relationship). The mechanisms behind coexistence and the biodiversity-productivity relationship remain an ecological mystery. One hypothesis is that plants obtain water and nutrients from different places …
Range-Wide Migratory Connectivity Of Painted Buntings, Andrew J. Sharp
Range-Wide Migratory Connectivity Of Painted Buntings, Andrew J. Sharp
All Graduate Theses and Dissertations, Spring 1920 to Summer 2023
The Painted Bunting (Passerina cirus) is a small (~16g), short/medium-distance migratory songbird that is listed as a species of conservation concern by the US Fish and Wildlife Service. Painted Buntings breed in two distinct populations, one eastern population and one interior population, separated by a 500 km gap that stretches from Mississippi to Georgia. I analyzed tracking data from 112 Painted Buntings from 11 different breeding sites to explore how individuals from different breeding sites differ in wintering location (chapter 2) and migratory timing (chapter 3). Additionally, I examined differences in migratory behavior between male and females in …
Bighorn Sheep Demographics Following Pneumonia Die-Off Events, Kylie Sinclair
Bighorn Sheep Demographics Following Pneumonia Die-Off Events, Kylie Sinclair
All Graduate Theses and Dissertations, Spring 1920 to Summer 2023
Bighorn sheep populations across the Intermountain West are subject to disease pressure from the respiratory bacteria Mycoplasma ovipneumoniae. Although the effects of M. ovipneumoniae-associated disease die-offs are well documented, less is known about the factors driving long-term differences in post-die-off population responses. While many herds experience years to decades in which recruitment is less than 20 lambs per 100 ewes, some herds’ lamb survival rates are able to rebound rapidly following die-off events. The reason why these herds recover quickly while others do not is currently unknown. Here, we assess the roles environmental, demographic, and pathogen-associated factors could …
Biomethanation And Alkaline Wet Air Oxidation Of Water Hyacinth (Pontederia Crassipes) From Ozama River, Dominican Republic, Yessica A. Castro
Biomethanation And Alkaline Wet Air Oxidation Of Water Hyacinth (Pontederia Crassipes) From Ozama River, Dominican Republic, Yessica A. Castro
All Graduate Theses and Dissertations, Spring 1920 to Summer 2023
Obtaining valuable products from environmental remediation waste is a sustainable approach that contributes to the ecological well-being of developing countries. In the present work, the feasibility of the water hyacinth anaerobic digestion as a post-weed management practice in the Ozama River (Dominican Republic) was demonstrated. The estimated energy required for harvesting was lower than that produced during digestion. The biomethanation of water hyacinth was improved by Alkaline Wet Air Oxidation (AWAO), a thermochemical pretreatment process that almost doubled the methane production rate and increased the yield by 24% when conducted at high temperatures. At lower temperatures, the methane yield of …
Effect Of Dietary Short And Medium Chain Fatty Acids In Murine Lethal Sepsis Model, David Son Dang
Effect Of Dietary Short And Medium Chain Fatty Acids In Murine Lethal Sepsis Model, David Son Dang
All Graduate Theses and Dissertations, Spring 1920 to Summer 2023
Sepsis is defined as the presence of an infection in combination with systemic inflammation in a host. One of the detrimental results of sepsis is multiple organ failure which eventually leads to death. Lipopolysaccharide (LPS), an endotoxin that can be found on the outer surface of gram-negative bacteria, can be used as a model for sepsis. LPS present in the circulatory system of hosts can induce symptoms that are seen in patients during septicemia.
Previous studies have suggested that including milk fat in a diet may protect against a lethal dosage of LPS in rodents. Milk fat has a unique …
"He Beheld The Prince Of Darkness": Joseph Smith And Diabolism In Early Mormonism 1815-1831, Steven R. Hepworth
"He Beheld The Prince Of Darkness": Joseph Smith And Diabolism In Early Mormonism 1815-1831, Steven R. Hepworth
All Graduate Theses and Dissertations, Spring 1920 to Summer 2023
Joseph Smith published his first known recorded history in the preface to the 1830 edition of the Book of Mormon. In it, he recounted the loss of the earliest portions of the book’s manuscript due to theft in a scheme the future prophet believed was concocted by the devil. Smith claimed to receive a revelation that instructed him on how to overcome the diabolic plot meant to discredit and destroy the Book of Mormon. This was not Smith’s first or last recorded encounter with the devil. He believed the devil was real and that he was engaged in a cosmic …
Socioscientific Issues In School-Based Agricultural Education: Describing And Exploring Factors Of Integration, Michelle S. Burrows
Socioscientific Issues In School-Based Agricultural Education: Describing And Exploring Factors Of Integration, Michelle S. Burrows
All Graduate Theses and Dissertations, Spring 1920 to Summer 2023
Socioscientific issues (SSI) are complex issues which are scientific in nature and have societal impacts. Many SSI have connections to agriculture and as such should be included in agricultural education curriculum. A clear understanding of what school-based agricultural education (SBAE) teachers know about SSI is needed. The purpose of this research was to explore the knowledge and integration of SSI among SBAE teachers by describing and explaining the factors that influence integration. This quantitative survey research was guided by the SSI-based instruction framework and the three-component model of agricultural education. The population for this study was all SBAE teachers in …
Monitoring Populations And Movement Of Bobcats (Lynx Rufus) On The Eastern Slope Of The Sierra Nevada Mountains Of California, Michael Brown
Monitoring Populations And Movement Of Bobcats (Lynx Rufus) On The Eastern Slope Of The Sierra Nevada Mountains Of California, Michael Brown
All Graduate Theses and Dissertations, Spring 1920 to Summer 2023
Monitoring the spatial ecology and population densities of carnivores is critical for effective management and conservation of these populations and the ecosystems in which they exist. However, effective monitoring of carnivore populations through estimates of space use, habitat selection and densities can be difficult due to their relatively low densities and wide ranging, elusive behaviors. Bobcats (Lynx rufus) are medium sized, top-level predators which are widely distributed across North America. Quantifying space use, habitat selection and developing effective population monitoring strategies for this species will have important implications for wildlife management.
My first objective was to use telemetry …
Responses Of Pacific Fishers To Habitat Changes As A Result Of Forestry Practices In Southwestern Oregon, Tessa R. Smith
Responses Of Pacific Fishers To Habitat Changes As A Result Of Forestry Practices In Southwestern Oregon, Tessa R. Smith
All Graduate Theses and Dissertations, Spring 1920 to Summer 2023
The fisher (Pekania pennanti) is a medium-sized carnivore found in mature forest stands across much of the northern United States. Although historically abundant in the west, fisher populations declined rapidly after fur trapping, extensive logging, and urban development reduced their numbers. Currently, biologists are concerned about the effects timber harvest practices have on fisher tolerance and adaptability when faced with changes to high-quality habitat stands. Tree removal and thinning of understory vegetation are frequently used to alleviate the spread of wildfires in previously dense forest stands with a potential for large-scale habitat loss; yet, a deficit of large …
A Laboratory Study On The Geometric Effects Of Piano Key Weirs On Scour For Non-Cohesive Substrates And Simple Mitigation Techniques, Wyatt Lantz
All Graduate Theses and Dissertations, Spring 1920 to Summer 2023
Sustainable flood control infrastructure is needed as climate change continues to produce more extreme precipitation events, as our water infrastructure continues to age, and as populations continue to grow and expand. Non-linear weirs, such as labyrinth and piano key weirs, are sustainable passive flood-control structures due to their improved hydraulic performance, and low maintenance and construction costs. However, like other structures, these weirs are susceptible to local scour, which is removal of soil and rock due to hydraulic forces. The scour phenomenon is a complicated process dependent upon multiple variables. There is limited information for non-linear weirs with little directed …
Compost And Cover Crop Effects In Dryland Organic Wheat, Michael D. Deakin
Compost And Cover Crop Effects In Dryland Organic Wheat, Michael D. Deakin
All Graduate Theses and Dissertations, Spring 1920 to Summer 2023
Producers of dryland organic wheat (Triticum aestivum L.) in the western USA struggle to maintain adequate soil fertility due to the high cost of organic fertilizers and concerns over moisture use of cover crops. Low soil fertility results in decreased wheat yield and quality, and increased year-to-year variability in yield and quality based on weather. This study was conducted to measure the effects of, and interactions between, cover crop mixes and a one-time compost application on soil health and winter wheat yield and quality. The study was located on three adjacent certified organic wheat farms near Snowville, UT, each …
An Eulerian Perspective On Spring Migration In Mule Deer, Tatum Del Bosco
An Eulerian Perspective On Spring Migration In Mule Deer, Tatum Del Bosco
All Graduate Theses and Dissertations, Spring 1920 to Summer 2023
Many herbivores travel between low-elevation winter ranges and high-elevation summer sites. These seasonal movements allow them to avoid deep snow cover, ensure access to favorable habitat, and maximize food intake throughout the year. During the spring season, plants at lower elevations green up earlier at lower elevations than at higher elevations. It has been shown that individual animals will track this vegetation growth during their spring migration, which allows them to maximize forage intake coming out of the nutrient scarce winter. This phenomena has previously been studied by monitoring individual movement trajectories, but it is unknown how this pattern scales …