Digital Commons Network^™

Drought Drove Forest Decline And Dune Building In Eastern Upper Michigan, Usa, As The Upper Great Lakes Became Closed Basins, Walter L. Loope, Henry M. Loope, Ronald J. Goble, Timothy G. Fisher, David E. Lytle, Robert J. Legg, Douglas A. Wysocki, Paul R. Hanson, Aaron R. Young

Department of Earth and Atmospheric Sciences: Faculty Publications

Current models of landscape response to Holocene climate change in midcontinent North America largely reconcile Earth orbital and atmospheric climate forcing with pollen-based forest histories on the east and eolian chronologies in Great Plains grasslands on the west. However, thousands of sand dunes spread across 12,000 km² in eastern upper Michigan (EUM), more than 500 km east of the present forest-prairie ecotone, present a challenge to such models. We use 65 optically stimulated luminescence (OSL) ages on quartz sand deposited in silt caps (n = 8) and dunes (n = 57) to document eolian activity in EUM. Dune building …

Concatenation And Concordance In The Reconstruction Of Mouse Lemur Phylogeny: An Empirical Demonstration Of The Effect Of Allele Sampling In Phylogenetics, David W. Weisrock, Stacey Dewitt Smith, Lauren M. Chan, Karla Biebouw, Peter M. Kappeler, Anne D. Yoder

School of Biological Sciences: Faculty Publications

The systematics and speciation literature is rich with discussion relating to the potential for gene tree/species tree discordance. Numerous mechanisms have been proposed to generate discordance, including differential selection, longbranch attraction, gene duplication, genetic introgression, and/or incomplete lineage sorting. For speciose clades in which divergence has occurred recently and rapidly, recovering the true species tree can be particularly problematic due to incomplete lineage sorting. Unfortunately, the availability of multilocus or “phylogenomic” data sets does not simply solve the problem, particularly when the data are analyzed with standard concatenation techniques. In our study, we conduct a phylogenetic study for a nearly …

Global Estimation Of Evapotranspiration Using A Leaf Area Index-Based Surface Energy And Water Balance Model, H. Yan, S.Q. Wang, David P. Billesbach, Walter Oechel, J.H. Zhang, Tilden Meyers, Ta. Martin, Roser Matamala, Dennis D. Baldocchi, G. Bohrer, D. Dragoni, R. Scott

Department of Agricultural and Biological Systems Engineering: Faculty Publications

Studies of global hydrologic cycles, carbon cycles and climate change are greatly facilitated when global estimates of evapotranspiration (E) are available. We have developed an air-relative-humidity-based two-source (ARTS) E model that simulates the surface energy balance, soil water balance, and environmental constraints on E. It uses remotely sensed leaf area index (L_ai) and surface meteorological data to estimate E by: 1) introducing a simple biophysical model for canopy conductance (G_c), defined as a constant maximum stomatal conductance g_smax of 12.2mm s⁻¹multiplied by air relative humidity (R_{h …}

Where Do Adaptive Shifts Occur During Invasion? A Multidisciplinary Approach To Unravelling Cold Adaptation In A Tropical Ant Species Invading The Mediterranean Area, Olivier Rey, Arnaud Estoup, Merav Vonshak, Anne Loiseau, Simon Blanchet, Luis Calcaterra, Lucila Chifflet, Jean-Pierre Rossi, Gael J. Kergoat, Julien Foucaud, Jérôme Orivel, Maurice Leponce, Ted Schultz, Benoit Facon

United States Department of Agriculture-Agricultural Research Service / University of Nebraska-Lincoln: Faculty Publications

Evolution may improve the invasiveness of populations, but it often remains unclear whether key adaptation events occur after introduction into the recipient habitat (i.e. post-introduction adaptation scenario), or before introduction within the native range (i.e. prior-adaptation scenario) or at a primary site of invasion (i.e. bridgehead scenario). We used a multidisciplinary approach to determine which of these three scenarios underlies the invasion of the tropical ant Wasmannia auropunctata in a Mediterranean region (i.e. Israel). Species distribution models (SDM), phylogeographical analyses at a broad geographical scale and laboratory experiments on appropriate native and invasive populations indicated that Israeli populations followed an …

Program And Proceedings: Nebraska Academy Of Sciences 1880–2012, 132nd Anniversary Year, One Hundred-Twenty-Second Annual Meeting

Nebraska Academy of Sciences: Programs and Proceedings

Program

Applied Science and Technology

Biological and Medical Sciences

Nebraska Wesleyan University Health and Sciences Graduate School Fair

Aeronautics and Space Science, Poster Session

Maiben Memorial Lecture: "The Status of Nebraska's Native Aspen," "Buffalo" Bruce McIntosh, Western Nebraska Resources Council

Aeronautics

Anthropology

Biological and Medical Sciences

Chemistry and Physics

Collegiate Academy: Biology

Collegiate Academy: Chemistry and Physics

Earth Science

History/Philosophy of Science

Junior Academy, Senior High Competition

Teaching of Science and Math

Junior Academy, Junior High Competition

Teaching Culture In The 21st Century Language Classroom, Oxana Dema, Aleidine Kramer Moeller

Department of Teaching, Learning, and Teacher Education: Faculty Publications

The paper presents an overview of the research on teaching culture and describes effective pedagogical practices that can be integrated into the second language curriculum. Particularly, this overview tries to advance an approach for teaching culture and language through the theoretical construct of the 3Ps (Products, Practices, Perspectives), combined with an inquiry-based teaching approach utilizing instructional technology. This approach promotes student motivation and engagement that can help overcome past issues of stereotyping and lack of intercultural awareness. The authors summarize the research articles illustrating how teachers successfully integrate digital media together with inquiry learning into instruction to create a rich …

Weighing The Merits Of Corn Ethanol, Christopher Fulk

Op-Eds from ENSC230 Energy and the Environment: Economics and Policies

Weighing the Merits of Corn Ethanol

Magnitude And Structure Of Lymantria Dispar Asiatica Infestations Of Common Forest Steppe Tree Species In Northern Mongolia, Nina Tiralla, Kai Füldner, Stefan Schütz

Erforschung biologischer Ressourcen der Mongolei / Exploration into the Biological Resources of Mongolia, ISSN 0440-1298

Lymantria dispar L. is one of the most serious forest pests worldwide by infesting large forest areas involving massive tree dieback. Unlike outbreaks of Lymantria dispar L. in Germany that cause massive defoliation, observations of outbreaks of Lymantria dispar asiatica made in the research area Khonin Nuga, West Khentii, Mongolia during the past six years showed only punctual defoliation, and dieback was extremely scarce. However, information on the performance of Lymantria dispar asiatica and the corresponding reaction of host tree species in Khonin Nuga is rare. Therefore, this study concentrates on the examination of primarily affected tree species and the …

Bird Red List And Its Future Development In Mongolia, Sundev Gombobaatar, D. Samiya, Jonathan M. Baillie

Erforschung biologischer Ressourcen der Mongolei / Exploration into the Biological Resources of Mongolia, ISSN 0440-1298

With the involvement of the World Bank, Zoological Society of London, Dutch Government and National University of Mongolia, the volumes of Mongolian Red Lists of Fish, Amphibians and Reptiles, Birds and Mammals were completed, and Mongolia is now among the few nations that have up-to-date conservation assessments for all vertebrates. Of the 476 assessed native bird species of Mongolia, 10% were categorized as regionally threatened including Near Threatened. A further 0.6% were categorized as Critically Endangered (CR), 1.7% as Endangered (EN), 3.3% as Vulnerable (VU), and 4.4% as Near Threatened (NT). Almost 90% of Mongolian birds are categorized as Least …

Morphological Disparity Among Rock Voles Of The Genus Alticola From Mongolia, Kazakhstan And Russia (Rodentia, Cricetidae), V. N. Bolshakov, I. A. Vasilyeva, A. G. Vasilyev

Erforschung biologischer Ressourcen der Mongolei / Exploration into the Biological Resources of Mongolia, ISSN 0440-1298

The Central Asian high-mountain or rock voles of the genus Alticola Blanford, 1881 are one of the least studied groups of rodents. Taxonomic status of various geographic forms is not yet established until our days. This genus is not enough examined by biochemical and molecular genetic methods. We analyzed morphological disparity among rock voles of the genus Alticola from 8 localities. The samples studied represent 7 nominal species: A. semicanus Allen, 1924 (East Mongolia), Alticola argentatus Severtzov, 1879 (East Kazakhstan), A. tuvinicus Ognev, 1950 (Russia: Tuva), A. olchonensis Litvinov, 1960 (Russia: Olchon Island), A. macrotis Radde, 1861 (Russia: Khamar-Daban ridge), …

Long-Term Research On Biodiversity In West Khentey, Northern Mongolia, Michael Mühlenberg

Erforschung biologischer Ressourcen der Mongolei / Exploration into the Biological Resources of Mongolia, ISSN 0440-1298

Biodiversity is studied at ecosystem, community, and species level around Khonin Nuga, where the University Goettingen supports a research station since 15 years (established 1997 in cooperation with the National University of Mongolia). In that period 39 scientists have been involved and 67 students graduated with theses (7 PhD) about field work at that station. Inventories started at 1998 for several taxa and are now compiled in a book, which will be printed 2012 in Ulaanbaatar. It covers fungi, lichens, plants, several insect groups, spiders, crustaceans, and all vertebrate classes. For all taxa adequate experts proved the species lists. Because …

Nestling Food In The Desert Wheatear Oenanthe Deserti In The Dzungarian Gobi, Mongolia, Ueli Rehsteiner

Erforschung biologischer Ressourcen der Mongolei / Exploration into the Biological Resources of Mongolia, ISSN 0440-1298

The quality and size of nestling food of the Desert Wheatear Oenanthe deserti ssp. atrogularis were investigated in the spring of 2001 in the Dzungarian Gobi in Mongolia. Data are based on observations of nine pairs. Nestling food consisted of several taxa caught on the ground and in the air. Diptera, Coleoptera larvae, and Hymenoptera were the most frequent prey, that is, they contributed 17 to 30% of all food items each. The food composition changed with nestling age and season.

Food item size decreased with date. A higher proportion of multiple prey loadings were brought to nestlings in the …

Protection Of The Natural And Cultural Heritage Of The Mongolian Altai, Ulikpan Beket, Hans D. Knapp

Erforschung biologischer Ressourcen der Mongolei / Exploration into the Biological Resources of Mongolia, ISSN 0440-1298

The Altai-Sayan Ecoregion is known as a hotspot of biodiversity with large wilderness landscapes and a high rate of endemism in Central Asia and Siberia. There are many large and important protected areas of different categories. Parts of the Russian Altai, the “Golden Mountains of Altai”, are inscribed as World Natural Heritage. Also the neighbouring countries contain pristine landscapes, which could be a potential for an extension to a transnational serial World Heritage Site.

The Mongolian part of the ecoregion is characterized by very diverse landscapes and vegetation complexes: Deserts, semi-deserts and desert steppes in arid basins, river floodplains, salt …

On The Importance Of Pollen Morphology In Classification Of Chenopodiaceae In Mongolia, Gaadan Punsalpaamuu, Frank Schluetz, Tserendorj Gegeensuvd, Davaadorj Saindovdon

Erforschung biologischer Ressourcen der Mongolei / Exploration into the Biological Resources of Mongolia, ISSN 0440-1298

The pollen morphology of 18 species from 12 genera of Chenopodiaceae native to Mongolia was studied. Plants were collected during field courses and research projects on useful plants and are deposited in the Herbarium of the Mongolian State University of Education. Chemical preparation was carried out in the Palynological Laboratories at the School of Natural Sciences of the Mongolian State University of Education and the Department of Palynology and Climate Dynamics of the University of Göttingen in Germany. The results show, that all investigated species are distinguishable in terms of pollen morphology bases on their pollen grain sizes, exine thickness …

Home Range Characteristics Of Corsac And Red Foxes In Mongolia, Tserendorjiin Munkhzul, J. D. Murdoch, R. P. Reading

Erforschung biologischer Ressourcen der Mongolei / Exploration into the Biological Resources of Mongolia, ISSN 0440-1298

Red (Vulpes vulpes) and corsac foxes (V. corsac) live sympatrically throughout most of Mongolia, but few details of their home range characteristics exist. We captured and radio-tagged 13 red fox (♂ = 5, ♀ = 8) and 15 corsac foxes (♂ = 8, ♀ = 7) between 2004 and 2008. We tracked their movements to estimate home range sizes and examined the effects of four factors on home range size, including sex, age, season, and year. We determined mean home range size for 12 red and 10 corsac foxes that had sufficient data using fixed kernel …

Pleistocene Faunal Fossils From Bayangol I Site, Bulgan Aimag, Mongolia, Takao Sato, Fedora Khenzykhenova, Toshiaki Tsurumaru, Masao Ambiru, Jun Takakura, Yoshiaki Otsuka, Shigeo Iida, Natalya Schepina, Batmunkh Tsogtbaatar

Erforschung biologischer Ressourcen der Mongolei / Exploration into the Biological Resources of Mongolia, ISSN 0440-1298

During excavation of the Bayan-gol I Palaeolithic site in the Bulgan Aimak, Mongolia, some faunistic remains were recovered attributable to Aves (two species), Anura (two species), and mammals (three species). The fossil remains bear a considerable similarity to the corresponding species of today in Mongolia.

Ochotona Daurica Pallas, 1776: Modern And Past Distribution Area In Mongolia And The Transbaikal Region, M. A. Erbjeva, N. V. Alexeeva, T. V. Kisloschaeva

Erforschung biologischer Ressourcen der Mongolei / Exploration into the Biological Resources of Mongolia, ISSN 0440-1298

Ochotona daurica Pallas, 1776 is one of the ancient species among modern taxa having an origination in Transbaikalia possible at the end of Pleistocene. Review on the Late Pleistocene- Holocene-Recent area of distribution of this species is given on the base of new evidences and detail analysis of the previous data.

Ecology And Behaviour Of The Black-Billed Capercaillie (Tetrao Urogalloides Stegmanni) In The Khentej Mountains, Mongolia, Siegfried Klaus, Karl-Heniz Schindlatz, Alexander V. Andreev, Hans-Heiner Bergmann

Erforschung biologischer Ressourcen der Mongolei / Exploration into the Biological Resources of Mongolia, ISSN 0440-1298

Introduction

A common inhabitant of East Siberian larch forests, the Siberian or black-billed capercaillie (Tetrao urogalloides) ranges south to the limits of the boreal forests in the northern Mongolian mountains and east to the very coast of the Asian continent (KLAUS et al. 1989). Tetrao urogalloides MIDDENDORF (1851) has priority over T. parvirostris BONAPARTE (1856). Therefore, we prefer the T. urogalloides.

The subspecies T.u. stegmanni was first described on the basis of morphological differences by POTAPOV (1985) using specimens collected during Russian expeditions by KOZLOVA (1930). This description was based on 18 males in the collection at …

Recruitment Of Larix Sibirica Ledeb. In Closed Forest Stands, On Clear-Felling Sites And At Fire-Sites In The Forests Of Mongolia, Vasiliy T. Yarmishko, Nikolay N. Slemnev

Erforschung biologischer Ressourcen der Mongolei / Exploration into the Biological Resources of Mongolia, ISSN 0440-1298

The paper deals with recruitment patterns in larch forests of Mongolia following anthropogenic impacts (felling, fires), and describes successional trends in highland forest communities. It is established that mass seed recruitment of Larix sibirica Ledeb. took place during anomalous combinations of hydrothermal conditions supposedly occurring at periods of about 100 years. During the last decades, frequent fires of various intensities put serious constraints on reforestation of the larch, and induced successional trends in disturbed forests.

Osnabrück Botanical Expeditions To Mongolia, Barbara Neuffer, Nikolai Friesen, Batlai Oyuntsetseg, Tseden Jamsran, Herbert Hurka

Erforschung biologischer Ressourcen der Mongolei / Exploration into the Biological Resources of Mongolia, ISSN 0440-1298

Three botanical expeditions to Mongolia have been undertaken by the Botany Department of the University of Osnabrück in cooperation with the Botany Department of the National Mongolian University of Ulaanbaatar. The first expedition in 2000 took us to the Mongolian Altay, the second in 2001 to the Gobi and Gobi Altay, and the third in 2010 to East Mongolia including the Khingan Mts. (Numrug). In 2000, we were the first botanists being allowed to enter the border area between China and Russia in the Mongolian Altay since a long time, because this had been a prohibited area for decades. We …

50 Years Mongolian-German Biological Expeditions And Their Future = Fifty Years Mongolian-German Biological Expeditions And Their Future, Michael Stubbe, Ravčigijn Samjaa, Annegret Stubbe

Erforschung biologischer Ressourcen der Mongolei / Exploration into the Biological Resources of Mongolia, ISSN 0440-1298

First two paragraphs:

In 2012 we celebrate the 50th anniversary of the Mongolian-German Biological Expeditions. The roots can be found in the old German-Russian scientific relationships in the history of the University Halle-Wittenberg in connection with August Hermann Francke (1663–1727) and his Foundations (founded in 1698). It’s a long tradition, dating back to the 18th century when early researchers as Georg Wilhelm Steller (1709 –1746) or Peter Simon Pallas (1741–1811) reached the northern border of Mongolia. Other sources of close connections between Halle and Russia are given by persons as the physician Friedrich Hoffmann (1660–1742), the mathematician Christian Wolff (1679-1754) …

Biodiversity And Evolutionary Development Of Oligocene-Pliocene Lagomorphs (Lagomorpha, Mammalia) Of Mongolia, M. A. Erbajeva

Erforschung biologischer Ressourcen der Mongolei / Exploration into the Biological Resources of Mongolia, ISSN 0440-1298

Lagomorphs (pikas and hares) are an ancient group of small mammals originated in Asia in the Paleocene-Eocene. The earliest evidence of their presence in Mongolia is dated to the Early Oligocene. The taxa flourished during the Late Oligocene, Miocene and Pliocene; at the Late Pliocene they were reduced both in their diversity and in abundance. No data on lagomorph are known from the Pleistocene and Holocene, though there are 7 taxa of lagomorphs present in the modern fauna. Altogether, more than 50 lagomorph species, extinct at present, are known to have existed in Mongolia since the Oligocene through the Late …

Buchrezension – Book Review: Jan Bemmann (Herausgeber) (2012): Steppenkrieger. Reiternomaden Des 7.–14. Jahrhunderts Aus Der Mongolei, Primus Verlag Darmstadt, Format Din A4, 416 Seiten, Zahlreiche Farbfotos Und Zeichnungen, Michael Stubbe

Erforschung biologischer Ressourcen der Mongolei / Exploration into the Biological Resources of Mongolia, ISSN 0440-1298

Book review of Jan Bemmann (Herausgeber) (2012): Steppenkrieger. Reiternomaden des 7.–14. Jahrhunderts aus der Mongolei. Primus Verlag Darmstadt, Format DIN A4, 416 Seiten, zahlreiche Farbfotos und Zeichnungen, Preis 39,90 Euro. ISBN: 978-3-86312-011-5.

Opposing Growth Trends Created By External Disturbances In Larch Forests Of The Mongolian Altai, Choimaa Dulamsuren, Mookhoor Khishigjargal

Erforschung biologischer Ressourcen der Mongolei / Exploration into the Biological Resources of Mongolia, ISSN 0440-1298

If tree-ring series from different trees are merged to a chronology, in order to for example make inferences on climate, these series must be checked for consistency. Statistical parameters, including the Gleichläufigkeit (GL = coefficient of agreement) and standard t value have been applied for this purpose and are usually combined to boost the informative value. We tested the hypothesis that low GL and t values can, in turn, be used as indicators of external disturbance in a case study in forests of Siberian larch (Larix sibirica) in the Mongolian Altai. Since these forests were known to …

Diversity And Community Pattern Of Darkling Beetles (Coleoptera: Tenebrionidae) Along An Ecological Gradient In Arid Mongolia, M. Pfeiffer, E. Bayannasan

Erforschung biologischer Ressourcen der Mongolei / Exploration into the Biological Resources of Mongolia, ISSN 0440-1298

One of the most conspicuous detrito- and phytodetritophagous groups of beetles in the Asian steppes and deserts is the family Tenebrionidae (Coleoptera, Polyphaga) (KONSTANTINOV et al. 2009). Mongolia harbors a rich diversity of these beetles with 215 species and 50 genera of Tenebrionidae listed for the country (MEDVEDEV 1990), many of them restricted to its arid parts, where they obtain high abundances and dominate – together with ants – the insect soil fauna (PFEIFFER et al. 2003). Their dark color may be a means to withstand the high radiation at daytime; however, a large number of species is night active …

Comparative Study Of Constituents Of Essential Oils Of Ocimum Basilicum L. Cultivated In The Mongolian Gobi, Shataryn Altantsetseg, Sandui Shatar, N. Javzmaa

Erforschung biologischer Ressourcen der Mongolei / Exploration into the Biological Resources of Mongolia, ISSN 0440-1298

The essential oils of the herb Ocimum basilicum L., cultivated in the Mongolian Gobi, have been examined. Oils were isolated by hydro-distillation and analyzed by GC-MS. The principle components of Common Basil were methyl chavicol (52.1 %), linalool (23.9 %). In the oils of the different varieties of Basil were the following compounds found: Sweet Basil: linalool (24.5–27.4 %), methyl chavicol (19.8–20.0 %), bergamotene (10.0 %), 1.8-cineole (8.5 %); Purple Basil: linalool (52.8 %), 1.8-cineole (8.7 %); Cinnamon Basil: methyl chavicol (60.4 %), 1.8-cine-ole (6.3 %), linalool (3.3 %) and cadinol (3.2 %); Italian large leaf Basil …

“Agrionemys Kazachstanica Terbishi” Or The Two-Faced Mongolian Steppe Tortoise, Hermann Ansorge, Uwe Fritz, Khayankhyarvaa Terbish, Setev Shar

Erforschung biologischer Ressourcen der Mongolei / Exploration into the Biological Resources of Mongolia, ISSN 0440-1298

Although no extant native turtle and tortoise species is known to occur in Mongolia, a new subspecies of the Central Asian tortoise was described by Chkhikvadze under the name Agrionemys kazachstanica terbishi in 2009. The description was based on a mummified tortoise kept in a museum collection. Since then the Mongolian steppe tortoise has been considered as an endemic taxon Testudo horsfieldii terbishi (Chkhikvadze, 2009) for Mongolia.

However, there is no evidence for the occurrence of any wild tortoise species in Mongolia, even in the putative area of origin of the type specimen. The closest confirmed occurrence of Central Asian …

New Species Of Eimeria (Apicomplexa: Eimeriidae) From Ochotona Hyperborea And Ochotona Pallasi (Lagomorpha, Ochotonidae) In Mongolia, David S. Tinnin, Ethan T. Jensen, Nyamsuren Batsaikhan, Sumiya Ganzorig, Scott Lyell Gardner

Erforschung biologischer Ressourcen der Mongolei / Exploration into the Biological Resources of Mongolia, ISSN 0440-1298

An examination of the feces from 8 pikas collected in 1999 and from 17 pikas collected in 2009 in Mongolia revealed the presence of 3 new eimerian species. Four of the 5 species of pikas present in Mongolia were studied including: Ochotona alpina, O. dauurica, O. pallasi, and O. hyperborea. Oocysts of Eimeria dunnumi n. sp. from O. hyperborea collected in 1999 are ellipsoid, average length and width of 31.4 x 20.8 μm, with a 1.4 μm thick double layered wall, lacking a micropyle, oocyst residuum, and polar granule. Sporocysts of this species are ellipsoid, 12.8 …

Diploid Allium Ramosum From East Mongolia: A Missing Link For The Origin Of The Crop Species A. Tuberosum?, Batlai Oyuntsetseg, Frank R. Blattner, Nikolai Friesen

Erforschung biologischer Ressourcen der Mongolei / Exploration into the Biological Resources of Mongolia, ISSN 0440-1298

In eastern Mongolia, a diploid close relative of the tetraploid (4x) crop species Allium tuberosum and its closest wild relative A. ramosum (4x) was found and characterized by karyotype analysis and with molecular marker techniques. An earlier analyses revealed A. ramosum to be sister of the crop but excluded it as its progenitor. At that time a putative diploid cytotype of A. ramosum was hypothesized as a potential progenitor taxon of the tetraploids. New phylogenetic analyses of chloroplast and nuclear DNA sequences including the recently found cytotype (A. aff. tuberosum) together with A. tuberosum and A. ramosum accessions …

Biochemical Research On Mongolian Lichens, Bryophytes And Vascular Plants – In Memoriam, Dr. Siegfried Huneck (1928–2011), Hans D. Knapp

Erforschung biologischer Ressourcen der Mongolei / Exploration into the Biological Resources of Mongolia, ISSN 0440-1298

Dr. Siegfried Huneck, biochemist and lichenologist from the Institute for Biochemistry of Plants in Halle/Saale, died on September 9, 2011. He was a world wide leading expert on lichen substances. An obituary with curriculum vitae and a complete list of publications was published by Stordeur et al. (2011)