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An Evaluation Of Deterrent Methods Utilized To Prevent Crop Raiding By African Elephants (Loxodonta Africana) In The Kasigau Wildlife Corridor, Kenya, Rebecca Lynn Von Hagen Jul 2018

An Evaluation Of Deterrent Methods Utilized To Prevent Crop Raiding By African Elephants (Loxodonta Africana) In The Kasigau Wildlife Corridor, Kenya, Rebecca Lynn Von Hagen

Masters Theses & Specialist Projects

Escalating human elephant conflict (HEC) continues to be a contributing factor

towards elephant decline, and crop raiding is the most common form of negative

human-elephant interactions. For communities that cannot reverse or prevent crop

raiding, it is necessary to contain HEC events through deterrent measures. Few

deterrent measures exist that combine practicality and affordability while also

preventing habituation by elephants. This project focused on comparing the efficacy of

deterrent methods to assess which was the most successful at preventing elephants

from entering crops in the farming community of Sasenyi, Kenya. In this paired-control

study, four deterrent methods were evaluated: acacia …


Morphological Variation In The Genus Chlorocebus: Ecogeographic And Anthropogenically Mediated Variation In Body Mass, Postcranial Morphology, And Growth, Trudy R. Turner, Christopher A. Schmitt, Jennifer Danzy Cramer, Joseph Lorenz, J. Paul Grobler, Clifford J. Jolly, Nelson B. Freimer Jul 2018

Morphological Variation In The Genus Chlorocebus: Ecogeographic And Anthropogenically Mediated Variation In Body Mass, Postcranial Morphology, And Growth, Trudy R. Turner, Christopher A. Schmitt, Jennifer Danzy Cramer, Joseph Lorenz, J. Paul Grobler, Clifford J. Jolly, Nelson B. Freimer

All Faculty Scholarship for the College of the Sciences

Objectives

Direct comparative work in morphology and growth on widely dispersed wild primate taxa is rarely accomplished, yet critical to understanding ecogeographic variation, plastic local variation in response to human impacts, and variation in patterns of growth and sexual dimorphism. We investigated population variation in morphology and growth in response to geographic variables (i.e., latitude, altitude), climatic variables (i.e., temperature and rainfall), and human impacts in the vervet monkey (Chlorocebus spp.).

Methods

We trapped over 1,600 wild vervets from across Sub‐Saharan Africa and the Caribbean, and compared measurements of body mass, body length, and relative thigh, leg, and foot …


Small Mammals Of Family Muridae In Protected Areas Of Pakistan, Fakhra Nazir, Andleeb Batool, Inayat Ullah Malik, Safdar Ali Shah, Sabiha Shamim Jul 2018

Small Mammals Of Family Muridae In Protected Areas Of Pakistan, Fakhra Nazir, Andleeb Batool, Inayat Ullah Malik, Safdar Ali Shah, Sabiha Shamim

Journal of Bioresource Management

Murids to have more than 1300 species globally, forming the largest mammal group. Murids are found nearly everywhere in the world, though many subfamilies have narrower ranges. Murids are not found in Antarctica and many oceanic islands. Five National Parks from Northern areas of Pakistan were physically surveyed. The parks were studied at different times. Dhirkot National Park (DNP) in February 2008, Banjosa National Park (BJNP) from May to June 2009, Pir Lasura National Park (PLNP) from June to July 2009 and Pir Chanasi National Park (PCNP) from April to May 2010. A total of 6 species belonging to the …


Status Of Jackals And Foxes (Family: Canidae) From Selected Hill Areas Of Pakistan, Andleeb Batool, Asif Gondal, Inayat Ullah Malik, Safdar Ali Shah Jul 2018

Status Of Jackals And Foxes (Family: Canidae) From Selected Hill Areas Of Pakistan, Andleeb Batool, Asif Gondal, Inayat Ullah Malik, Safdar Ali Shah

Journal of Bioresource Management

The family Canidae consists of coyotes, dogs, foxes, jackals, and wolves. The IUCN red list reports 38 species under this family. This study was conducted in Azad Jammu and Kashmir, Pakistan. Conserved areas of Tolipir National Park, Banjosa Game Reserve, Pir Chanasi National Park, Pir Lasura National Park and Dhirkot Nature Reserve were studied from 2008 to 2010. Vulpes vulpes griffithi (Kashmir or hill fox) is a flexible hunter. It mainly feeds on hares, rodents, reptiles and occasionally small birds. In absence of vertebrate prey, they may subsist on insects and fruit. Canis aureus (Golden Jackal) is a very adaptable …


Review Of Microbuthus With Description Of M. Satyrus Sp. N. (Scorpiones, Buthidae) From Oman And Yemen, Graeme Lowe, František Kovařík, Mark Stockmann, František Šťáhlavský Jun 2018

Review Of Microbuthus With Description Of M. Satyrus Sp. N. (Scorpiones, Buthidae) From Oman And Yemen, Graeme Lowe, František Kovařík, Mark Stockmann, František Šťáhlavský

Euscorpius

The taxonomy of the genus Microbuthus is reviewed, and a new species from Oman and Yemen, M. satyrus sp. n., is described and fully illustrated with color photographs of live and preserved specimens, as well as of its habitat. It is compared to the closely similar species M. litoralis, which we also illustrate. Synonymy of the type species M. pusillus Kraepelin, 1898 with M. litoralis (Pavesi, 1885) is confirmed, and the species is recorded for the first time from Yemen. Hemispermatophores of M. satyrus sp. n., M. gardneri Lowe, 2010, and …


Common Loons Respond Adaptively To A Black Fly That Reduces Nesting Success, Walter H. Piper, Keren B. Tischler, Andrew Reinke Jun 2018

Common Loons Respond Adaptively To A Black Fly That Reduces Nesting Success, Walter H. Piper, Keren B. Tischler, Andrew Reinke

Biology, Chemistry, and Environmental Sciences Faculty Articles and Research

Nesting birds must often cope with harassment from biting insects, but it is difficult to ascertain what effect such pests might have on breeding success and population dynamics. We tested the hypothesis that a black fly (Simulium annulus) that feeds on the blood of nesting Common Loons (Gavia immer) causes nest abandonment in this charismatic diving bird. In addition, we measured effects of fly-induced abandonment on a loon population, and examined potential predictors of fly abundance and nest abandonment. We also tested a second hypothesis, which holds that loon pairs that abandon a nest owing to …


A New Species Of Vaejovis From The Patagonia Mountains, Southern Arizona (Scorpiones: Vaejovidae), Richard F. Ayrey Jun 2018

A New Species Of Vaejovis From The Patagonia Mountains, Southern Arizona (Scorpiones: Vaejovidae), Richard F. Ayrey

Euscorpius

A new scorpion species, Vaejovis patagonia sp. nov. is described and placed in the “vorhiesi” group. This small, dark brown species is found near Patagonia, Arizona. It is geographically closest to V. troupi Ayrey & Soleglad, V. grahami Ayrey & Soleglad and V. vorhiesi Stahnke. Those three species are found in a triangle surrounding the Patagonia Mountains, the locality of Vaejovis patagonia. The pedipalp fixed finger has 5 ID denticles and the movable finger has 6, like most, but not all, of the other southern Arizona Vaejovis. Carapace of female is shorter than metasomal segment V.


The Genera Butheolus Simon, 1882 And Xenobuthus Gen. Nov. (Scorpiones: Buthidae) In Oman, Graeme Lowe Jun 2018

The Genera Butheolus Simon, 1882 And Xenobuthus Gen. Nov. (Scorpiones: Buthidae) In Oman, Graeme Lowe

Euscorpius

The genus Butheolus Simon, 1882 is revised based on new material from Dhofar Province in Oman. B. gallagheri Vachon, 1980 is redescribed, and a related new species, B. harrisoni sp. n., is also described. The species B. anthracinus (Pocock, 1895) is redescribed and moved to a new genus Xenobuthus gen. n., that is differentiated from Butheolus by size, pedipalp finger dentition, setation, granulation and hemispermatophore structure, and a related new species, X. xanthus sp. n., is also described. Revised diagnoses are provided for the genus Butheolus, and for the species B. …


Nou Spring Field Days And Annual Meeting, Ft. Robinson, May 18-20, 2018, Janis Paseka Jun 2018

Nou Spring Field Days And Annual Meeting, Ft. Robinson, May 18-20, 2018, Janis Paseka

Nebraska Bird Review

The NOU Annual Meeting and Spring Field Days, organized by Robin Harding, Jan Johnson and Ruth Stearns, was held at Ft. Robinson State Park in Dawes and Sioux Counties on May 18-20, 2018, with 82 in attendance. Our lodgings and meeting room were in reconstructed buildings of the old Fort. The weather was perfect on Friday and Sunday but perfectly awful on Saturday, with rain all day and a high of 44°F. Finding birds and driving on unpaved roads was challenging on Friday, but the total species list for the meeting is 180. Field trips were led by Kathy DeLara, …


Spring Field Report, March - May 2018, W. Ross Silcock Jun 2018

Spring Field Report, March - May 2018, W. Ross Silcock

Nebraska Bird Review

This spring will be remembered for Nebraska’s first unequivocal record of Swainson’s Warbler, unfortunately a “one day wonder”, but well-photographed. Another mega was a Band-tailed Pigeon, Nebraska’s fourth. The old sports saying “wait until next year” was applicable in spades to woodwarblers; last year was noted for its paucity of most species of wood-warblers, but this year, incredible record numbers occurred for several migrant species: Magnolia, Baybreasted, Chestnut-sided, Blackpoll, and Palm. Other amazing numbers were the 598,000 Sandhill Cranes counted at peak migration in the Central Platte Valley and the 598 Great Egrets reported within the amazing 414 reports. Several …


First Record Of Fulvous Whistling-Duck (Dendrocygna Bicolor) For Nebraska, Matthew T. Haugen Jun 2018

First Record Of Fulvous Whistling-Duck (Dendrocygna Bicolor) For Nebraska, Matthew T. Haugen

Nebraska Bird Review

On the afternoon of 5 June 2018, I was scouting local Salt Valley lakes to get an estimate of goose numbers for annual Canada Goose (Branta canadensis) banding operations. At approximately 2:30 p.m. I was tallying my goose counts in the northwest corner of Wagon Train Wildlife Management Area in Lancaster County, when I noticed four ducks on a nearby vegetated mud flat. After closer examination, I recognized the ducks as Fulvous Whistling-Ducks (Dendrocygna bicolor). I spent a few years working in Louisiana in that state’s waterfowl program, and I am familiar with the species and its identification. I was …


The Nebraska Bird Review, Volume 86 June 2018 Number 2 Jun 2018

The Nebraska Bird Review, Volume 86 June 2018 Number 2

Nebraska Bird Review

Spring Field Report, March - May 2018, by W. Ross Silcock …50-89

NOU Spring Field Days and Annual Meeting at Ft. Robinson, May 18-20, 2018, by Janis Paseka … 89-95

First Record of Fulvous Whistling-Duck (Dendrocygna bicolor) for Nebraska, by Matt Haugen … 96-98

Subscription and Organization Information … 99


Feeding Patterns Of Two Commercially Important Fish Species Scomberoides Commersonnianus And S. Tol In The Northern Arabian Sea Coast Of Pakistan, Nazia Qamar, Sher Khan Panhwar, Ralf Riedel Jun 2018

Feeding Patterns Of Two Commercially Important Fish Species Scomberoides Commersonnianus And S. Tol In The Northern Arabian Sea Coast Of Pakistan, Nazia Qamar, Sher Khan Panhwar, Ralf Riedel

Faculty Publications

Port landing of Scomberoides commersonnianus and S. tol were obtained between July 2013 and June 2015 for stomach content analysis. Analysis of prey composition was done using permutational analysis of variance (permanova), with species, life stage (juvenile and adults), gender, and weather (rainy and dry season) as factors. Patterns of empty stomachs were investigated to estimate feeding intensity. Feeding intensity was estimated with logistic regression, using the same independent variables as above. Prey importance was also investigated. Prey importance was assessed using a Wilcox Rank Correlation analysis on the Index of Relative Importance (IRI) by species and life-stage. Permanova analysis …


Digging Biomechanics In Geomyoid Rodents, Alexis Moore Crisp May 2018

Digging Biomechanics In Geomyoid Rodents, Alexis Moore Crisp

UNLV Theses, Dissertations, Professional Papers, and Capstones

Subterranean digging behaviors provide opportunities for protection, access to prey, and predator avoidance for a diverse array of vertebrates, yet studies of the biomechanics of burrowing have been limited by the technical challenges of measuring kinetics and kinematics of animals moving within a substrate. Prior studies of burrowing have recorded a single axis of x-ray video and/or force. However, empirical observations show that burrowing is not restricted to a single axis or plane. I describe a new system called a ‘tunnel-tube’ for measuring 3D reaction force during burrowing. This tunnel-tube has two separate tubes, one ‘entry tube’ that has no …


Checklist Of Bats In Some Areas Of Azad Jammu And Kashmir, Fakhra Nazir, Inayatullah Malik, Sabiha Shamim May 2018

Checklist Of Bats In Some Areas Of Azad Jammu And Kashmir, Fakhra Nazir, Inayatullah Malik, Sabiha Shamim

Journal of Bioresource Management

Bats form an essential part of the ecosystem by aiding pollination, keeping insect pests in check and overall maybe helpful as indicators of health status of the ecosystem. This study was conducted to formulate a checklist of bats from different regions in Azad Jammu and Kashmir. Five National parks from Azad Jammu and Kashmir were surveyed for the species of bats present (Dhirkot National Park, Pir Lasura National Park, Pir Chanasi National Park, Banjosa National Park and Tolipir National Park). From the study conducted, 10 species of bats belonging to 5 families were found cumulatively in the five national parks.


Morphological Traits As Indicators Of Sexual Dimorphism In Prairie Rattlesnakes (Crotalus Viridis), Colleen Rothe-Groleau, Claudia M. Rauter, James D. Fawcett May 2018

Morphological Traits As Indicators Of Sexual Dimorphism In Prairie Rattlesnakes (Crotalus Viridis), Colleen Rothe-Groleau, Claudia M. Rauter, James D. Fawcett

Transactions of the Nebraska Academy of Sciences and Affiliated Societies

As humans encroach into areas inhabited by predators, the potential of human-predator confrontations increases and the predators become regarded as dangerous pests. Predators exert a measure of population control over pest species such as small rodents, as well as limit the quantity and scope of diseases (e.g. spread of Hantavirus by these prey species). Control of these small rodent pest species can be aided by conserving and managing their predators like rattlesnakes (Crotalus spp). Management of any population requires detailed information on population composition and the ability to determine the key information (especially age and sex) for each individual …


Biological And Abiotic Factors Influencing Cetacean Mass Strandings In The Northeastern United States, Octavia Williams May 2018

Biological And Abiotic Factors Influencing Cetacean Mass Strandings In The Northeastern United States, Octavia Williams

HCNSO Student Capstones

Marine mammal strandings are frequent occurrences along many coastal areas around the world. The significance of stranded cetaceans has been of interest since around 300 B.C. when philosophers, like Aristotle, pondered why marine mammals, including whales, dolphins and porpoise beach themselves; and famous authors, like Henry David Thoreau, whose fascination in mammal strandings lead him to write a book about these mass occurrences off the coast of Cape Cod. Numerous hypothetical theories have been developed to explain these mass stranding phenomena. Though the causes of mass strandings remain unresolved, recent investigations suggest contributing factors could include environmental elements. Less emphasis …


Behavioural Risks In Male Dogs With Minimal Lifetime Exposure To Gonadal Hormones May Complicate Population-Control Benefits Of Desexing, Paul Mcgreevy, Bethany Wilson, Melissa Starling, James A. Serpell May 2018

Behavioural Risks In Male Dogs With Minimal Lifetime Exposure To Gonadal Hormones May Complicate Population-Control Benefits Of Desexing, Paul Mcgreevy, Bethany Wilson, Melissa Starling, James A. Serpell

Endocrinology Collection

Castration of dogs is a widespread practise with clear justification in population control and knock-on benefits for animal welfare. Deleterious behavioural consequences of castration are believed to be negligible. Gonadectomy is widely recommended as part of a multi-factorial approach to prevent problems including aggression in dogs. However, the consequences of early castration on health are still being debated. The current study focused on the reported behaviour of 6,235 male dogs castrated before 520 weeks of life for reasons other than behavioural management, and calculated their percentage lifetime exposure to gonadal hormones (PLGH) as a proportion of their age at the …


Trophic Ecology And Parasitism Of A Mesopelagic Fish Assemblage, Matthew Woodstock May 2018

Trophic Ecology And Parasitism Of A Mesopelagic Fish Assemblage, Matthew Woodstock

HCNSO Student Theses and Dissertations

Mesopelagic (open ocean, 200-1000 m depth) fishes are important consumers of zooplankton and are prey of oceanic predators. Some mesopelagic fishes (e.g., myctophids and stomiids) undertake a diel vertical migration where they ascend to the near-surface waters during the night to feed and descend into the depths during the day to avoid predators. Other mesopelagic fishes (e.g., Sternoptyx spp.) do not vertically migrate and remain at deep depths throughout the day. While in the epipelagic zone (surface – 200 m depth), vertically migrating fishes become prey to upper-trophic level predators, such as: tunas and billfishes. Benthic fishes (e.g., macrourids) often …


Effects Of Environmental Enrichment On Behavior In A Domestic Goat Herd, Stephanie Peramas May 2018

Effects Of Environmental Enrichment On Behavior In A Domestic Goat Herd, Stephanie Peramas

Senior Honors Projects

Animal welfare is one of the most important parts of animal management. Apart from other measures of good animal welfare, like being in good health, animal behavior can be indicative of the animal’s internal state and is often one of the first signs of improper welfare. Stereotypies, or stereotypic behavior, are commonly viewed as a symptom of improper animal welfare. Unenriched, or barren, environments associated with farmed livestock have long been associated with inducing stereotypic behavior. The addition of precise enrichment to the animals’ environment is a common remedy for stereotypic behavior. With respect to stereotypic goat behavior, few empirical …


Ichthyology Scientific Collection Spreadsheet, Karisa Boyer May 2018

Ichthyology Scientific Collection Spreadsheet, Karisa Boyer

Ichthyology Collection

No abstract provided.


Winter Bat Activity In A Landscape Without Traditional Hibernacula, Amy M. Hammesfahr, Rene E. Ohms May 2018

Winter Bat Activity In A Landscape Without Traditional Hibernacula, Amy M. Hammesfahr, Rene E. Ohms

United States National Park Service: Publications

Abstract

Prior to 2014, bat research at Devils Tower National Monument (DETO) focused on bats present during the summer months. Biologists at DETO assumed local bats were strictly summer residents due to the presumed lack of typical habitat features associated with bat hibernation, such as caves and mines. This lack of traditional hibernacula features at DETO discouraged staff and research cooperators from studying winter bat populations. Despite the earlier assumption that bats were unlikely to hibernate on the monument, DETO documented significant winter bat activity through passive winter acoustic monitoring. This study is the first study at DETO that documents …


Defending Wild Dogs: Population Dynamics And Disease In Endangered African Wild Dogs, Elizabeth Claire Arredondo May 2018

Defending Wild Dogs: Population Dynamics And Disease In Endangered African Wild Dogs, Elizabeth Claire Arredondo

Graduate Theses and Dissertations

African wild dogs (Lycaon pictus) are endangered carnivores whose population is decreasing from habitat loss and fragmentation, interspecific competition, and disease. Survival rates are especially low in Kruger National Park (KNP), though it is unclear why. I estimated the abundance in KNP and survival rates over different time spans, six years and nine months, using public photographic survey data. In 2015, there were 298 (SE=12.1) individuals. Using a mark-recapture analysis in program R, I found that the survival rate between 2009-2015 was only 3.2%, and within the 9-month survey period (September 2014 – June 2015), monthly survival rates for the …


The Effects Of Habitat And Connectivity On Tropical Ant Ecology And Behavior., Benjamin Jacob Adams May 2018

The Effects Of Habitat And Connectivity On Tropical Ant Ecology And Behavior., Benjamin Jacob Adams

Electronic Theses and Dissertations

This dissertation explores the role that physical connections among and within habitats play in determining the local diversity and behaviors of cursorial organisms. Understanding the processes that regulate local community assembly and that contribute to the maintenance of species diversity is a cornerstone of community ecology. Ants are a useful model for studies of local communities because they are abundant and fulfill a variety of ecological roles. Chapter one of the dissertation examines the how lianas (woody vines) function as connective structures among tree crowns and how these connections affect arboreal ant community structure within a tropical forest canopy. Trees …


Personality In California Sea Lions (Zalophus Californianus) And Harbor Seals (Phoca Vitulina): Methodological Convergence And Species-Specific Emotional Repertoires, Amber De Vere May 2018

Personality In California Sea Lions (Zalophus Californianus) And Harbor Seals (Phoca Vitulina): Methodological Convergence And Species-Specific Emotional Repertoires, Amber De Vere

Dissertations

Despite the wide acceptance of animal personality as a valid area of study, research on marine mammal personality remains remarkably scarce. What literature does exist predominantly focuses on bottlenose dolphins (Frick, 2016; Highfill & Kuczaj, 2007; Kuczaj, Highfill & Byerly, 2012; Lilley, de Vere, Yeatre & Kuczaj, 2018; Moreno, Highfill & Kuczaj, 2017). There is also strong evidence for individual differences in grey seals (Robinson et al., 2015; Twiss & Franklin, 2010; Twiss, Culloch & Pomeroy, 2011; Twiss, Cairns, Culloch, Richards & Pomeroy, 2012), and preliminary research has provided evidence of broad personality factors in pinniped species using behavioral coding …


Vocalizations Of North American River Otters (Lontra Canadensis) In Two Human Care Populations, Sarah Walkley May 2018

Vocalizations Of North American River Otters (Lontra Canadensis) In Two Human Care Populations, Sarah Walkley

Master's Theses

There is a dearth of information regarding the vocal repertoire of North American river otters (Lontra canadensis). This indicator species is cosmopolitan yet elusive, making recordings methodologically difficult in the wild. Therefore, this exploratory study uses video and audio recordings of two populations of North American river otters in human care to broaden the known vocal repertoire of river otters in various social contexts. The populations consist of a male-female pair at the Stamford Museum and Nature Center and a male-male pair at The Maritime Aquarium at Norwalk. This study is the first to examine the vocalizations produced …


Towards Quantifying Relevant Land Cover Change: A Case Study In The Central Flyway Of The Monarch Butterfly, Hanna Lenee Ford May 2018

Towards Quantifying Relevant Land Cover Change: A Case Study In The Central Flyway Of The Monarch Butterfly, Hanna Lenee Ford

Graduate Theses and Dissertations

Over the past 20 years notable decreases in monarch butterfly populations have led researchers to begin evaluating the landscape for changes and seeking out opportunities for enacting conservation programs to better support their survival. The monarch butterfly has recently come under consideration for listing under the Endangered Species Act which has created a need for a more informed view of the landscape through which the migrate and breed, the central United States. In this research three spatially-explicit models are explored using the most applicable datasets currently available to address pressing policy and land manager decisions regarding monarch butterfly and pollinator …


Effect Of Gestation Length On Litter Size And Piglet Birth Weight, Audrey L. Emery, Dillon Q. Siefert, Courtney K. Moore, Flint W. Harrelson, Patricia L. Harrelson Apr 2018

Effect Of Gestation Length On Litter Size And Piglet Birth Weight, Audrey L. Emery, Dillon Q. Siefert, Courtney K. Moore, Flint W. Harrelson, Patricia L. Harrelson

Celebration of Student Scholarship Poster Sessions Archive

No abstract provided.


Scorpions Of The Horn Of Africa (Arachnida, Scorpiones). Part Xvi. Compsobuthus Maidensis Sp. N. (Buthidae) From Somaliland, František Kovařík Apr 2018

Scorpions Of The Horn Of Africa (Arachnida, Scorpiones). Part Xvi. Compsobuthus Maidensis Sp. N. (Buthidae) From Somaliland, František Kovařík

Euscorpius

Compsobuthus maidensis sp. n. from Somaliland is described and fully complemented with color photos of specimens, as well as its habitat. Data on the occurrence of the genus Compsobuthus Vachon, 1949 in the Horn of Africa is summarized.


Can Diet Affect Coloration In Tiger Salamanders?, Katherine Novak Apr 2018

Can Diet Affect Coloration In Tiger Salamanders?, Katherine Novak

Undergraduate Research Symposium 2018

Amphibians are the fastest known declining taxonomic group in the world with 48% of populations in decline (Stuart et. al. 2004). These declines in amphibian populations are partially due to the lack of basic life history data (Semlitsch 2003). For example, amphibians are known for their bright coloration and ability to rapidly change color. Coloration is key to amphibian survival in terms of defense (as camouflage or a warning sign to predators), thermoregulation (darker skin warms faster), and communication (primarily mating displays) (Rudh and Qvanstrom 2013). One aspect of amphibian coloration that is poorly understood is how much the environment …