Digital Commons Network^™

Alfred Russel Wallace Notes 37. What About Wallace On Vaccination . . . ?, Charles H. Smith

Faculty/Staff Personal Papers

Best known for his natural history studies on evolution and biogeography, Alfred Russel Wallace (1823−1913) had a significant second career as a social critic. One of his most controversial stances was his position on (smallpox) vaccination. Anti-vaxxers of the present frequently bring up his name as a resource for their side, but the situation is not so easily reducible to that.

The Influence Of Human Presence And Footprint On Animal Space Use In Us National Parks, Kaitlyn M. Gaynor, Forest P. Hayes, Kezia Manlove, Nathan Galloway, John F. Benson, Michael J. Cherry, Rolf O. Peterson, Sarah Hoy, John Vucetich, Et Al.

Michigan Tech Publications

Given the importance of protected areas for biodiversity, the growth of visitation to many areas has raised concerns about the effects of humans on wildlife. In 2020, the COVID-19 pandemic led to temporary closure of national parks in the United States, offering a pseudonatural experiment to tease apart the effects of permanent infrastructure and transient human presence on animals. We compiled GPS tracking data from 229 individuals of 10 mammal species in 14 parks and used third-order hierarchical resource selection functions to evaluate the influence of the human footprint on animal space use in 2019 and 2020. Averaged across all …

Use Of High-Throughput Phenomics With And Without Fungicide As A Wheat Breeding Tool, Gerardo Ivan Rivera Collazo

Dissertations and Doctoral Documents, University of Nebraska-Lincoln, 2023–

Wheat (Triticum aestivum) is an important food staple for many countries around the world and current consumption and production data demonstrate a production deficit. Breeders are challenged to help close this gap in production by selecting better cultivars with improved yields. Several methods aim to accelerate the breeding process to reduce the time required for releasing an elite variety. One of the main breeding bottlenecks is the lack of fast and reliable phenotypic data acquisition that could dissect physiological and morphological plant data. High-throughput remote sensing could have the potential to reduce this bottleneck by streamlining data acquisition, …

Lianas Reduce Tree Height With Negative Consequences For Carbon Storage And Growth Estimates, Xingyan Cao, Frieke Van Coillie, Sruthi M. Krishna Moorthy, Kasper Coppieters, Barbara D'Hont, Stefan A. Schnitzer, Geertje M. F. Van Der Heijden, Hans Verbeeck, Félicien Meunier

Biological Sciences Faculty Research and Publications

1. Current evidence suggests that liana (woody vine) competition with trees could be threatening the carbon sink by reducing carbon uptake and storage in tropical forests. Previous studies investigating forest demography in liana removal experiments have, however, assumed similar allometries for trees, regardless of the amount of lianas they support. Yet, recent observations suggest that liana load may alter tree shape and structure, including height, which could result in an underestimation of the liana effects on forest carbon stocks and tree growth.
2. In this study, we used terrestrial laser scanning (TLS) in a liana removal experiment in Gigante Peninsula, Panama, collected …

Mammals Of Malheur National Wildlife Refuge And Surrounding Harney County, Oregon, John E. Cornely, Leslie N. Carraway

Monographs of the Western North American Naturalist

Malheur National Wildlife Refuge (NWR), located in southeastern Oregon, is part of the National Wildlife Refuge System (NWRS) of the United States. Currently, NWRS includes more than 568 NWRs, 38 Wetland Management Districts, and 5 Marine National Monuments. Malheur NWR, like a majority of the refuges in the NWRS, was established for the conservation of migratory birds. Many of these NWRs also provide significant habitat for mammals. Malheur NWR, one of the oldest (established 1908) and largest refuges in the system, is one of those with significant habitat for mammals. In addition to the refuge, we have included Harney County, …

Seasonal And Landscape-Driven Variations In Forage Resources Of Apis Mellifera Scutellata: Implications For Pollination Sustainability And Colony Health In Taita Taveta County, Kenya, Mary Chege, Mbatha B. Wambua, Wambua J. Kilonzo, Sevgan Subramanian, Beatrice T. Nganso

All Peer-Reviewed Publications

Seasonality and land-use change are key factors influencing forage availability for managed honey bee colonies, yet knowledge of forage identity and how these factors influence forage availability remains limited in Africa. To address these gaps, we used DNA metabarcoding to identify nectar and pollen plant species supporting the nutrition of the African savannah honey bee, Apis mellifera scutellata, across different land-use/land cover types and seasons in Taita Taveta County, Kenya. We identified 224 forage plant species from 65 families, with Asteraceae, Fabaceae, and Myrtaceae being the top contributors. Forage availability was significantly influenced by landscape and season, with honey bees …

Cross-Continental Analysis Shows That Disturbance Effects On Reptile Body Condition Do Not Predict Abundance Responses, Kristina J. Macdonald, Don A. Driscoll, Michael D. Craig, Robert A. Davis, Steven J. Hromada, C. M. Gienger, Lee A. Fitzgerald, Daniel J. Leavitt, Danielle K. Walkup, Rickard Abom, Adrian F. Wayne, Gordon R. Friend, Brent Johnson, Danielle Stokeld, Chris R. Dickman, Scott A. Thompson, Graham G. Thompson, Daniel F. Bohórquez Fandiño, Christopher K. Woolley, Annabel L. Smith, Calum Irvine, Tim S. Jessop, Jade E. Keehn, Chris R. Feldman, Xavier Santos, Josabel Belliure

Research outputs 2022 to 2026

Ecological disturbances are discrete events that alter or transform the physical, chemical, or biological characteristics of ecosystems. Disturbance can cause animal populations to decline and, according to the risk-disturbance hypothesis and population collapse framework, these declines can be predicted by declines in animal body condition. However, no research has empirically examined the general relationship between body condition and abundance, nor their relationship in response to disturbance. We used a combined dataset representing 33 studies and > 42,000 observations of 75 species from Australia, New Zealand, Spain and the United States of America to test predictions relating to the relationship between reptile …

First-In-Human Phase 1 Study Of Khk2455 Monotherapy And In Combination With Mogamulizumab In Patients With Advanced Solid Tumors, Timothy A Yap, Olivier Rixe, Capucine Baldini, Ursa Brown-Glaberman, Sergey Efuni, David S Hong, Christophe Massard, Jameel Muzaffar, Andreea Varga, Emrullah Yilmaz, Yuta Ikawa, Lisa H Shiue, Yi Liu, Matthew W Hruska, Henry Zhao, Akihiro Tokunaga, Solmaz Sahebjam

Faculty, Staff and Student Publications

Background: Indoleamine 2,3-dioxygenase 1 (IDO1) is a heme-containing enzyme that degrades tryptophan (Trp) to kynurenine (Kyn), which suppresses effector T cells and reduces antitumor activity. KHK2455 is a long-acting selective IDO1 inhibitor that blocks the heme component of the IDO holoenzyme. Mogamulizumab is a humanized immunoglobulin G1 monoclonal antibody targeting CCR4. KHK2455 + mogamulizumab demonstrated enhanced antitumor activity in preclinical studies, which led to a first-in-human, two-part, multicenter, open-label, phase 1, dose-escalation, cohort-expansion trial (ClinicalTrials.gov identifier NCT02867007) evaluating the safety/tolerability, pharmacokinetics, and IDO1 activity of KHK2455 alone and in combination with mogamulizumab in patients with treatment-refractory advanced solid tumors. …

Exploring The Composition Of Forest Collaboratives In Northeastern Oregon, Lee K. Cervany, Rebecca J. Mclain, Kristen Wright

National Policy Consensus Center Publications

Community-based collaboration has been touted as an effective model for forest governance because it promotes democratized decision-making and stakeholder engagement to address landscape-scale problems. Forest collaboratives are assumed to be heterogeneous, consisting of stakeholders with a diverse range of interests. Few studies have systematically explored variables associated with collaborative composition. We identify six elements of collaborative composition for investigation: size, stakeholder diversity, balance, locality–diversity, core attendance, and cross-participation. This exploratory study examines five forest collaborative groups in eastern Oregon (USA). We analyzed meeting minutes over an 18-month period to track attendance and evaluate who shows up and at what frequency. …

Unfolding The Mystery Of Autoimmunity: The Environmental Determinants Of Diabetes In The Young (Teddy) Study, Marian Rewers, Daniel Agardh, Suzanne Bennett Johnson, Ezio Bonifacio, Helena Elding Larsson, Patricia Gesualdo, William Hagopian, Michael J Haller, Heikki Hyöty, Randi Johnson, Richard Mcindoe, Eoin Mckinney, Jessica Melin, Åke Lernmark, Richard E Lloyd, Kristian F Lynch, Jill M Norris, Stephen S Rich, Roswith Roth, Desmond Schatz, Jorma Toppari, Eric Triplett, Kendra Vehik, Suvi M Virtanen, Anette-G Ziegler, Beena Akolkar, Jeffrey P Krischer, Teddy Study Group

Faculty, Staff and Students Publications

In 2025, the National Institute of Diabetes and Digestive and Kidney Diseases (NIDDK) at the National Institutes of Health celebrates 75 years of leadership in diabetes research. The NIDDK serves people of the U.S. affected by or at risk for many chronic diseases, including diabetes and other endocrine, metabolic, and digestive disorders, by funding innovative research to develop better treatment and prevention and a cure for these conditions. Autoimmunity that leads to type 1 diabetes or celiac disease or thyroid autoimmunity affects 1 in 20 children and adolescents in the U.S. While treatments are available, prevention of these common autoimmune …

Identifying And Analysing Coastal Landforms In Ar Rāyis, Saudi Arabia: Insights From Sentinel Data And Field Studies, Omar A. Alharbi

Journal of King Abdulaziz University: Marine Science

This study examines the coastal geomorphology of the Ar Rāyis coastal zone on the eastern Red Sea coastal plain, along with the processes underpinning this geomorphology. Coastal areas feature various landforms, including beaches, intertidal flats, khawrs (lagoons), sand spits, sand dunes, sabkhas (salt marshes), coral reefs, wadis, raised coral reef terraces, and alluvial terraces. These landform features suggest rapid alterations in coastal conditions. Furthermore, the analysis of Sentinel-2B data via digital processing reveals a range of natural land covers, with field investigations confirming the results. The supervised classification methods assisted in distinguishing between regions of deep and shallow water, while …

Bite By Bite: How Ungulate Browsing Shapes North America's Forest Future, Darius Hardalau, Vladut Stefanescu, Mindaugas Bakševičius, Michael Manton, Charles Ruffner, Gediminas Brazaitis, Georgeta Ionescu, Ovidiu Ionescu

Aspen Bibliography

Ungulate browsing represents a contemporary issue for forest development, influencing forest regeneration, composition, and management practices across the world, especially in North America. This review synthesizes findings from 101 studies conducted between 1980 and 2025 to examine patterns of herbivory damage through browsing by moose, elk, white-tailed deer, black-tailed deer, and bison. Despite regional variation, high ungulate density consistently emerges as the primary factor driving browsing intensity and ecological and economic impact, leading to decreased social acceptance of coexistence with ungulates. This review highlights the selective suppression of palatable species such as balsam fir, red oak, and white cedar, leading …

The Millipede Genus Alocodesmus Silvestri, 1896 (Polydesmida: Chelodesmidae: Batodesmini): Description Of A New Species From Colombia And The Identity Of A. Hanseni (Silvestri, 1898), Juan Romero-Rincon, Santiago Alvear

Insecta Mundi

The genus Alocodesmus Silvestri, 1896 (Polydesmida: Chelodesmidae: Batodesmini) currently comprises six valid species, three of which are recorded for Colombia, including A. danielae Romero-Rincon and Alvear, new species, from Yondó, Antioquia Department. Two species are known from Panama and one from Venezuela. The new species differs from its congeners by having a spoon-shaped prefemoral process that is ventrally extended forming a conspicuous concave space with branch Z being shorter than branches X and Y. The identity of A. hanseni Silvestri, 1898, previously of uncertain status, is revised, and the species is herein proposed as a new synonymy …

Living Heritage, Resilience And Ethically Informed Infrastructures In Malawi, Martha B. Lerski, Frank Puluputu Mwale

Publications and Research

Rooted in a rural community’s needs, this paper explores how historically, and ethically informed infrastructures can support community engagement towards identification, documentation, and dissemination of local, traditional, and indigenous knowledge relating to climate change. The paper examines climate adaptation and sustainability issues through the lenses of community organizing and Library & Information Science best practices. It draws on a growing body of literature identifying a role for expanded epistemological and subject discipline perspectives. While the impacts of climate on material culture are increasingly recognized, intangible, and biocultural heritage have not received as much attention. Living heritage traditions and knowledge systems …

Agricultural Extension In An Urban Informal Settlement During Covid-19: The Effectiveness Of The “Tanimang Bayan” Initiative In Sitio San Roque, North Triangle, Quezon City, Patrice Bianca R. Yapjoco

CSID Journal of Infrastructure Development

The COVID-19 pandemic exposed deep inequalities in food access, particularly in urban informal settlements. This study examines the effectiveness of the “Tanimang Bayan” initiative, an urban agriculture project implemented in Sitio San Roque, Quezon City, as a form of agricultural extension during the pandemic. The research adopts a qualitative approach, drawing on data from key informant interviews, participant observation, and secondary sources. Findings indicate that the initiative provided a localized response to food insecurity by enhancing access to fresh produce and promoting community-led production. It also supported collective action and self-organization among residents, strengthening their autonomy in navigating structural limitations. …

Emergence Of Caenocoris Neri (Germar, 1847) On Fig Trees In The Oases Of Southeastern Algeria, Nacima Diab, Tesnim Deghiche, Khalila Bengouga, Vasily Vladimirovich Verkhoturov

Journal of Bioresource Management

The article considers materials related to current areas of fundamental research in the field of agricultural entomology in Algeria. The most important issues of agricultural entomology in Algeria include identifying patterns of general trends in the dynamics of geographic ranges of phyto-entomophagous under the influence of climatic and anthropogenic factors, as well as studying the adaptation of insects to new habitats and food supply. In a notable discovery during insect sampling conducted in the oasis ecosystem during the summer of 2019, researchers recorded and identified individuals of the species Caenocoris nerii (Germar, 1847) for the first time. These insects were …

First Insight On The Genesis Of Iron Oxide-Hydroxide Biospeleothems In Rancho Chico Lava Tube, Mexico, Rafael López-Martínez, Elizabeth Solleiro-Rebolledo, Rocio Alcántara-Hernández, Teresa Pi Puig, Sergey Sedov, Daisy Valera Fernández, M. Nayeli Luis-Vargas, Yasmany Lima Vera, Jesús Solé, Ramon Espinasa-Pereña

International Journal of Speleology

Iron is a common element in lava tube systems, typically observed as thin crusts or inferred from the reddish coloration of secondary mineral deposits. In this study, we describe iron oxide-hydroxide biospeleothems discovered in the Rancho Chico Lava Tube, located within the Xalapa Monogenetic Volcanic Field in Mexico. We adopt an integrative approach combining petrography, soil micromorphology, scanning electron microscopy, mineralogy, and 16S rRNA gene metabarcoding. DNA sequencing reveals the presence of microorganisms such as Acidithiobacillus, Gallionella, and members of the Leptospirillaceae family, which may be involved in the oxidation of Fe²⁺ within the lava tube. The source …

The Enigmatic Samuel Aughey: Nebraska’S Pastor Naturalist, Hugh H. Genoways, Margaret R. Bolick, Mary Anne Andrei

Zea E-Books Collection

Samuel Aughey at age 39 became the first Professor of Natural Science at the new University of Nebraska in Lincoln, serving from 1871 to 1883. He was an ordained pastor of the Evangelical Lutheran Church, with an addendum degree from Pennsylvania College (now Gettysburg College) in 1856 and an A.M. degree two years later. His early pastoral work was in Pennsylvania, but in 1864 he became a home missionary to a church in Dakota City, Nebraska. Aughey’s career at the university was marked with successes and honors, but also criticism and questions about his basic knowledge of the sciences. While …

Integrative Taxonomy Confirms The Synonymy Of Three Springsnail Species (Hydrobiidae: Pyrgulopsis), Kathryn E. Perez, Megan Solis, Michelle Flores, Chanté Lundskog

School of Integrative Biological & Chemical Sciences Faculty Publications

The small, spring-dwelling snails, Pyrgulopsis Call & Pilsbry, 1886, form a diverse radiation in Western North America. In this study, we examine the validity of three species of Pyrgulopsis, Pyrgulopsis pilsbryana (Baily & Baily, 1952), P. nonaria Hershler, 1998, and P. transversa Hershler, 1998. These three species of Pyrgulopsis are found primarily in Utah, with ranges extending into Idaho, Wyoming, and Nevada. They were originally diagnosed primarily on the number and position of penial glands, but subsequent examination with mitochondrial DNA data suggested they should be synonymized. Here, we incorporate additional mitochondrial and nuclear DNA data, increased geographic sampling, …

Analyzing Human - Nonhuman Primate Conflict Mitigation Techniques In Mto Wa Mbu, Northern Tanzania, Lil Adams

Undergraduate Theses, Capstones, and Recitals

Human-wildlife conflict is a widespread challenge faced by those living in regular contact with wildlife that can have profound impacts on livelihood outcomes for humans, wildlife, and their shared environment (Barua et al., 2013; Blackie, 2023). Human – non-human primate conflict is particularly crucial due to primates’ high capacity to live among human populations (Chapman & Chapman, 1990; Alberts & Altmann, 2006; Reader et al., 2011; Sinha & Vijayakrishnan, 2017), and is currently on the rise due to increasing contact between human and non-human primates (Hockings, 2016; Uddin et al., 2020). To characterize and analyze techniques currently being used to …

Global Patterns Of Genetic Diversity And Species Delimitation In The Lichen-Forming Fungus Cladia Aggregata: A Restriction Site Associated Dna Sequencing (Rad-Seq) Analysis Across Neotropical, Australasian, And Asian Populations, John De Abreu

College of Science and Health Theses and Dissertations

Cladia aggregata is a group of lichen-forming fungi comprising multiple species which are difficult to differentiate. C. aggregata has a wide distribution on several different continents but is primarily found in the southern hemisphere. We aimed to further delimit the species complex while also investigating the relatedness of the different species, where they can be found geographically, and what phenotypic traits they possess. We used Restriction Site Associated DNA Sequencing (RAD-seq) to compare thousands of loci across 91 individuals from the Americas, Asia, and Australasia. All Asian samples formed a distinct clade in all phylogenetic trees. Asia, as a distinct …

Asset Or Liability? Farmers’ Perceptions Of Improved Maize Seeds As A Climate Change Adaptation Strategy In Semi-Arid Areas Of Chamwino District, Tanzania, Jackson Raymond Sawe

Journal of Humanities and Social Sciences

This study explores the perceptions of smallholder farmers in semi-arid areas of Chamwino District, Tanzania, regarding the adoption of improved maize seeds as an adaptation strategy to climate change. It aims to explore whether farmers view the use of these seeds as an asset that helps them adapt to climate change, or as a liability that presents challenges resulting from the adoption of improved seeds. The study employs both quantitative and qualitative methods for data collection and analysis. Data were collected from 125 smallholder farmers across six study villages through household surveys, in-depth interviews, field observation, and document review. Participants …

Identifying The Drivers Behind The Decline Of Douglas-Fir And True Fir Species In Sw Oregon, Alexander Bryant Bentley

Dissertations and Theses

Increased levels of Pseudotsuga menziesii (DF) and Abies species (TF) decline throughout dry forests in the Klamath Mountains and Western Cascades ecoregions have been recorded in recent years, with preliminary evidence suggesting that factors influenced by changing climatic conditions and encroachment from fire suppression policies are primarily to blame. Yet, the physiological and environmental mechanisms of drought-driven tree decline are poorly understood across biophysical and spatial gradients. Given the complex relations that drive decline, determining the degree of influence of specific drivers is difficult, creating unique challenges for modern forest management. To elucidate the most important factors driving DF and …

Prescribed Fire Leads A Pine Flatwoods Specialist Tree Frog To Seek Local Refugia, Ian Biazzo, Pedro F. Quintana-Ascencio

Faculty Publications

Pine savannas of the southeastern United States evolved with frequent low-intensity wildfires. Today, land managers use prescribed fires to restore this ecological process while also controlling vegetative fuel accumulation. Given the ubiquitous application of this management tool, studies should consider impacts on target and nontarget taxa in these systems. During fires, animal populations persist in the landscape when individuals emigrate to unburned areas or seek refugia within the burn zone; otherwise, they experience local extirpation. We examined the effect of prescribed fires on a pine flatwoods specialist frog, the pinewoods treefrog Dryophytes femoralis, in a frequently burned pine landscape. …

Arts-Based Sustainability: From New York To Malawi, Martha B. Lerski

Publications and Research

Recognizing that libraries serve multiple constituencies and subject areas, this chapter documents and advocates for development of transdisciplinary arts-based research (ABR) and culture-related projects linked to environmental challenges. Libraries contribute collections and spaces, as well as the research of library and information scientists. Libraries are currently among invisible contributors to sustainability planning and services. The chapter will link this invisibility to the value of what visual arts refer to as negative space elements in subjects ranging from traditional ecological knowledge to environmental science. Library collections, projects, and research contribute to education for sustainable development (ESD) as required to achieve the …

United Seas For Resilient Reefs: A Blueprint For Coral Conservation Through International Cooperation, John J. Bohorquez, Maoz Fine, Dana I. Grieco, David A. Gill, Ali Al-Swalimih, Jacqueline Padilla-Gamino, Isaiah W. Bolden, Margaux Hein, Callum Backstrom, Melanie Mcfield, Raphael Martinez, Benjamin Midberry, Fahd Al-Guthmy, Ellen K. Pikitch, Natalia Benejam, Chen Azulay, Britt Ronen, Oren Levy, Anne Mcelroy, Karine Kleinhaus

School of Marine & Atmospheric Sciences Faculty Publications

Coral reefs are one of Earth’s most vital and threatened ecosystems. They protect coastlines, generate billions of dollars of tourism revenue, support valuable fisheries, and contribute to the development of vital medicines. Yet half of the world's coral reefs have been lost in the last 50 years, and 90% of remaining reefs are projected to die off by the middle of this century, mainly due to warming seas from climate change.

Despite these challenges there is reason for hope. Some coral reefs – such as in the northern Red Sea and Gulf of Aqaba – have demonstrated remarkable resilience to …

Art, Play, And Neuroplasticity: Advancing Dementia Care Through Art-Enriched Environments, Piper Hutson, James Hutson

Faculty Scholarship

Dementia represents a global public health crisis, characterized not only by progressive memory loss but by a pervasive decline in cognitive flexibility, social functioning, and overall quality of life. Traditional therapeutic modalities, often limited to pharmacological management and cognitive training, have yielded only modest gains in symptom mitigation and patient well-being. Recent advances in neuroscience have illuminated the brain‘s capacity for neuroplasticity—its inherent ability to adapt, reorganize, and form new neural connections even in advanced age and the presence of neurodegenerative conditions. This emerging understanding invites a critical re-evaluation of dementia care strategies, highlighting the potential of non-pharmacological interventions to …

Covid-19 Pandemic And Recovery Experiences: A Case Study With Montana Family And Consumer Science Cooperative Extension Agents, Hayley M. Somerfeld, J. Mitchell Vaterlaus, Nicole C. Wanago, Brianna Routh, Alison Brennan

Journal of Human Sciences and Extension

The present study examined the impact of COVID-19 and recovery experiences among Family Consumer Science (FCS) Extension Agents in Montana. All participants (n = 10) completed a survey with fixed-item and open-ended responses, and six also participated in in-depth interviews. A deductive qualitative case study approach was used. Agent experiences were analyzed using The Sendai Framework for Disaster Risk Reduction (SFDRR) four action priorities: (1) understanding risk, (2) strengthening disaster risk governance, (3) investing in disaster risk reduction for resilience, and (4) enhancing disaster preparedness for effective response and to “build back better” in recovery, rehabilitation, and reconstruction. Pandemic risks …

When Trees Disappear: Understory Successional Implications Of Sudden Oak Death And High-Severity Wildfire In Big Sur, California, Sam J. Balthazard

Master's Theses

The rapidly emerging disease sudden oak death, caused by the non-native pathogen Phytophthora ramorum, is transforming coastal California forests by driving extensive mortality of tanoak (Notholithocarpus densiflorus) and increasing fire severity. As a novel disturbance, sudden oak death and its interaction with wildfire may catalyze unexpected shifts in successional trajectories and reconfigure forest understory communities. Furthermore, how tanoak influences the forest understory and how its decline might reshape understory dynamics remain poorly understood. We hypothesize that disease alone and the compounding disturbances of disease and wildfire alter successional dynamics, and that the potential functional extinction of N. …

When Avatars Go To School: The Effect Of Virtual Reality On Learning And Error Correction, Lauren Barack

Dissertations, Theses, and Capstone Projects

The pandemic in 2020 led to an increasing dependency on technology particularly in learning environments. National educational assessments in the proceeding years have indicated steep, measurable learning loss. While a fully virtual learning environment is not the norm today, we chose to investigate how virtual reality (VR) environments influence learning in college students.

Our primary research goal was to look at how students learn and remember information gathered in virtual spaces. In particular, we examined whether sensory inputs in the VR environment, including spatial audio and haptic feedback, such as vibrations felt in a handheld controller, would help to better …