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Quo Vadis Equus Hemionus Hemionus In Mongolia?, Annegret Stubbe, Michael Stubbe, O. Shagdarsuren, Nyamsuren Batsaikhan, Ravčigijn Samjaa

Erforschung biologischer Ressourcen der Mongolei / Exploration into the Biological Resources of Mongolia, ISSN 0440-1298

Inspired by studies on biodiversity in the South Gobi we were confronted with the human impact on animal communities and endangered animal species, especially the Asiatic wild ass. So the idea was born to organize an international conference with the aim to actualize the status of Equus hemionus in Central Asia and to find ways for better conservation of this species and its habitats. A short review is given on historical exploration, decreasing area and population number of the Dschiggetai. Today the range of the nominate form of Equus hemionus is located in the Gobi region of southern Mongolia/northern China …

Remarks On The Social System Of The Mongolian Wild Ass (Equus Hemionus Hemionus), Gertrud Neumann-Denzau, Helmut Denzau

Erforschung biologischer Ressourcen der Mongolei / Exploration into the Biological Resources of Mongolia, ISSN 0440-1298

So far the social system of Mongolian wild asses is non-uniformly interpreted. The authors describe their own observations from different parts of Mongolia and review data from available publications. The arguments support a territorial social system, which agrees with the social system of all other wild asses in Asia and Africa.

The Birth Of A Wild Ass (Equus Hemionus Khur) In India’S Little Rann Of Kutch, Gertrud Neumann-Denzau

Erforschung biologischer Ressourcen der Mongolei / Exploration into the Biological Resources of Mongolia, ISSN 0440-1298

The birth of a wild ass was completely described and photographed in the wild for the first time. It happened in the daytime, in the open desert, and in the vicinity of other wild asses. The mother isolated her foal afterwards for an entire day. After birthing she drove away her male yearling, which did not join her any longer from this moment on.

A Review Of Leg Ornamentation In Male Wolf Spiders, With The Description Of A New Species From Australia, Artoria Schizocoides (Araneae, Lycosidae), Volker W. Framenau, Eileen Hebets

Eileen Hebets Publications

Morphological modifications of the first pair of legs in addition to widespread color variations of these legs among males of closely related species have been reported in a variety of spiders. Here, the evidence for sexual dimorphism in male foreleg morphology within wolf spiders (family Lycosidae) is reviewed and shown to occur in a number of species belonging to at least seven genera in five subfamilies: Alopecosa, Hogna, Schizocosa (all Lycosinae) Pirata (Piratinae), Evippa, (Evippinae), Pardosa (Pardosinae) and Artoria (Artoriinae). These modifications, often in combination with distinct dark pigmentation, can be divided into three major groups: leg elongation, segment …

Capture And Anaesthesia Of The Mongolian Wild Ass (E. Hemionus), Chris Walzer, Petra Kaczensky, Oogii Ganbaatar, Namtar Enkhsaikhan, Davaa Lkhagvasuren

Erforschung biologischer Ressourcen der Mongolei / Exploration into the Biological Resources of Mongolia, ISSN 0440-1298

Science-based conservation efforts in general and wide-ranging equid conservation specifically, often require capture and subsequent handling of the subject animal. Safe and animal-welfare appropriate wild equid capture and anesthesia is a complex operation necessitating a multitude of skills that require appropriate veterinary training. In order to develop management plans for the Mongolian wild ass (Equus hemionus), which range over vast areas the use of radiotelemetry, is an essential tool. Radio-telemetry allows the researchers to better understand the habitat requirements and to delineate the areas of potential wildlife-human conflicts. To date we have successfully captured 16 wild ass in …

Biodiversity In Mongolia, Photos, Annegret Stubbe, Michael Stubbe

Erforschung biologischer Ressourcen der Mongolei / Exploration into the Biological Resources of Mongolia, ISSN 0440-1298

Pictures of wildlife in Mongolia in 2007.

Asiatic Wild Asses In The Literature: What Do We Need To Know Now?, Sarah R.B. King

Erforschung biologischer Ressourcen der Mongolei / Exploration into the Biological Resources of Mongolia, ISSN 0440-1298

Asiatic wild asses were once found across central Asia, but over time their range has been reduced until today all subspecies are under threat. This paper examined the literature about wild asses with an aim to illustrating gaps in our knowledge as a target for future research.

A search was made on BIOSIS ISI for the following strings: Equus hemionus, wild ass, khulan, kulan, kiang, khur, and onager. In total 61 relevant references were found, published between 1900 and 2005. Most of these references were published in the 1990s, with only one published between 1900 and 1970. Khur and …

Mapping Khulan Habitats: A Gis-Based Approach, Henrik Von Wehrden, Karsten Wesche

Erforschung biologischer Ressourcen der Mongolei / Exploration into the Biological Resources of Mongolia, ISSN 0440-1298

This paper focuses on the contribution that a combination of public domain GIS data with extensive habitat data can make to Khulan conservation in southern Mongolia. We describe a dataset taken in the Great Gobi 'A' Strictly Protected Area which represents the driest region within the overall Khulan distribution range. Potential habitats were classified following a phyto-sociological approach based on vegetation samples collected in the field. These data are combined with an open source climate model that allowed assessment of the influences of mean precipitation and temperature regime on habitat conditions. The results are put into a Gobi-wide context providing …

Comparative Demography And Dietary Resource Partitioning Of Two Wild Ranging Asiatic Equid Populations, Ellen Schulz, Thomas Kaiser, Annegret Stubbe, Michael Stubbe, Ravčigijn Samjaa, Nyamsuren Batsaikhan, Joachim Wussow

Erforschung biologischer Ressourcen der Mongolei / Exploration into the Biological Resources of Mongolia, ISSN 0440-1298

Tooth wear signatures allow inference on the dietary traits of herbivorous ungulates. Comparing dietary regimes of taxonomically closely related populations further allows inference on habitat structure and food availability. The mesowear method of tooth wear evaluation has opened up a pathway to reconstruct subtle differences in dietary behavior and resource partitioning based on skeletal material as the only source of information. Eighty cheek dentitions of Asian wild asses (Equus hemionus) from the Southern Gobi (Mongolia) and 61 dentitions of African free ranging donkeys (Equus asinus) from the Emirate Sharjah (United Arab Emirates) were investigated for their …

Population Assessment Of Khulan (Equus Hemionus) In Mongolia, Badamjav Lkhagvasuren

Erforschung biologischer Ressourcen der Mongolei / Exploration into the Biological Resources of Mongolia, ISSN 0440-1298

The data presented here suggest that 18,411 (± 224) khulan inhabit southern Mongolia with a density of 1.4 individuals per 1,000 km² within the 157,525 km² territory of its actual distribution. The Dornogobi province contains the highest number of khulan with a mere 67%, while 20% are found in the Ömnögobi province, 12% are in Djungarian Gobi and only 1% in the Gobi-Altay and Bayankhongor provinces. According to the proportion of foals and yearlings within each aimag, the Dornogobi, Ömnögobi east and Khovd populations have an average reproduction rate.

Kulan (Equus Hemionus Pallas 1775) In Turkmenistan, Victor S. Lukarevskiy, Yu. K. Gorelov

Erforschung biologischer Ressourcen der Mongolei / Exploration into the Biological Resources of Mongolia, ISSN 0440-1298

The kulan (Equus hemionus kulan) is the only odd-toed ungulate left in the wild in northern Eurasia. In the 1930s, the kulan survived in southern Turkmenistan only. In the 1940s, kulans were found only in an area named Badkhyz in Turkmenistan. To conserve the last natural population of this subspecies of kulan the Badkhyz Natural Reserve was established in 1941. The historical and contemporary distribution, fluctuation in numbers, conservation and sci research of kulan populations in the Badkhyz Natural reserve are reviewed in this paper.

Zum Vorkommen Ausgewählter Bembidion-Arten In Der Mongolei (Coleoptera: Carabidae, Bembidiinae): 2. Beitrag Zur Carabidenfauna Der Mongolei, Peer Haro Schnitter

Erforschung biologischer Ressourcen der Mongolei / Exploration into the Biological Resources of Mongolia, ISSN 0440-1298

To the occurrence of some Bembidion-species in Mongolia (Coleoptera: Carabidae, Bembidiinae) - Second contribution to the carabid fauna of Mongolia.

The distribution of four Bembidion-species, Bembidion (Asioperyphus) infuscatum DEJEAN, 1831, B. (Asioperyphus) ovale MOTSCHULSKY, 1844, B. (Bracteon) conicolle MOTSCHULSKY, 1844 und B. (Bracteon) lapponicum ZETTERSTEDT, 1828, in Mongolia are discussed, and new data are given.

Investigation Of The Kiang (Equus Kiang, Equidae) Skull From Ladakh, India, Natalia Spasskaya

Erforschung biologischer Ressourcen der Mongolei / Exploration into the Biological Resources of Mongolia, ISSN 0440-1298

The species-rank of Equus kiang was recently supported by molecular and genetic studies (RYDER & CHEMNICK 1990). Three subspecies are currently recognized in this species: eastern - E. k. holdereri, southern - E. k. polyodon and western - E. k. kiang, differing in some cranial and external features (GROVES & MAZAK 1967). Kiang populations are scarce and widely separated at present; which was a reason for a reassessment of the taxonomic status and distribution of at least one or two of these three races.

An interesting skull of kiang was recovered by N. Paklina and C. van Orden …

Sustainable Natural Resource Use And Livelihood Improvement In South Gobi Protected Areas, J. Jargal

Erforschung biologischer Ressourcen der Mongolei / Exploration into the Biological Resources of Mongolia, ISSN 0440-1298

Many of the species in Gobi are in decline. People and livestock are dependent on scarce water supplies. Consequently, there is a strong competition between wild animals and livestock for water sources, grazing pastures and forage plants. This project assists conservation efforts by working with nomadic herding communities who have critical importance in the future of the Gobi desert ecosystem. Training nomadic herders in ecology, conservation and natural resource use is a unique approach to ecosystem management in Mongolia. Using this approach, the project aims to help herding families establish a sustainable use of natural resources, improve their livelihoods, and …

Feeding Ecology Of Asiatic Wild Ass Equus Hemionus, Jochen Lengger, Frieda Tataruch, Christian Walzer

Erforschung biologischer Ressourcen der Mongolei / Exploration into the Biological Resources of Mongolia, ISSN 0440-1298

The Mongolian wild ass or khulan (Equus hemionus) is a potential competitor for forage to reintroduced Przewalski’s horses. To evaluate the major foraging plants of khulan we chose the alkane method that was first described by Mayes in 1984. Different plant species contain varying amounts and proportions of n-alkanes. This fact allows the determination of diet composition by comparing the plant alkane to the fecal alkane pattern. The major advantage is that the method is non-invasive and has been shown to be a reliable method in captive trials. Fecal and plant samples were collected simultaneously and preserved by …

First Records And First Proven Breeding Of Lesser Grey Shrike, Lanius Minor, In Mongolia, Markus Deutsch, Axel Bräunlich

Erforschung biologischer Ressourcen der Mongolei / Exploration into the Biological Resources of Mongolia, ISSN 0440-1298

During a trip to the Dzungarian Gobi in southwestern Mongolia, a group of German birdwatchers found several territorial lesser grey shrikes, Lanius minor, and documented an active nest. The species had not previously been recorded from Mongolia. Observations were made near Bulgan gol (Bulgan river; nest coordinates: 46°6’N 91°32’E, 1190 m a.s.l.) in Khovd aimag (Khovd province) on June 24, 2006. Besides two territorial pairs, of which a female could be photographically documented on the nest, at least four more territorial adults were found within an area of about 1.5 km². Based on these records, an abundance …

Numbers, Distribution And Social Structure Of Kiang (Equus Kiang Moorcroft 1841) Population In The Southwestern Part Of Tibet, China, Natalia V. Paklina, Chris Van Orden

Erforschung biologischer Ressourcen der Mongolei / Exploration into the Biological Resources of Mongolia, ISSN 0440-1298

In September - October 1998 we made an extensive survey in southwest Tibet to study numbers, distribution and social structure of the kiang population in this remote part of its range. Kiangs were sighted between 29º 40’ and 32º 50’ N, and 81º- 86º E. They preferred broad flat valleys with lakes at an altitude from 4,000 up to 5,000 m. Small kiang groups and solitary animals were sighted in Transhimalaya. There were no kiangs in Tsangpo (Brahmaputra) River valley, in the sand desert near Ali and in the Clay Mountains near Tsada. Seven old kiang corpses were found along …

Some Information On The Ecology Of Khulan (Equus Hemionus Pallas, 1775) In The Western Part Of The South Gobi Province, Mongolia, D. Tsendjav, S. Purevsuren

Erforschung biologischer Ressourcen der Mongolei / Exploration into the Biological Resources of Mongolia, ISSN 0440-1298

This short study provides information on the density and distribution of khulans in September 2003 and speculates about factors that negatively influence the khulan population in the sums of Khurmen, Bayandalai, Noyon, Sevrei, and Gurvantes in the South Gobi province.

Turkmenian Kulan (Equus Onager Kulan) Captive Population, Anna Mekarska

Erforschung biologischer Ressourcen der Mongolei / Exploration into the Biological Resources of Mongolia, ISSN 0440-1298

The Turkmenian kulan (Equus onager kulan) is critically endangered, fewer then 1,500 animals are left in the wild. In 2004, there were additional 570 Turkmenian kulans living in captivity: 156 specimens in Euro-Asian zoos, 83 in Askania Nova Reserve and 331 in Canyon Colorado Equid Sanctuary. Population size in EAZA region is decreasing both in terms of numbers and genetic parameters. Those facts underline the urgent need for coordinating development of Eurasian kulan’s captive population. From 2004 onwards, large efforts to create healthy breeding groups in zoos raised bigger interest in this species in European zoos.

Identification Of Herder-Wild Equid Conflicts In The Great Gobi B Strictly Protected Area In Sw Mongolia, Petra Kaczensky, N. Enkhsaihan, O. Ganbaatar, Christian Walzer

Erforschung biologischer Ressourcen der Mongolei / Exploration into the Biological Resources of Mongolia, ISSN 0440-1298

In protected areas of Mongolian overgrazing, competition with wild ungulates and poaching are important management concerns. The Great Gobi B Strictly Protected Area (SPA) in SW Mongolia is a re-introduction site for the Przewalski’s horse (Equus ferus przewalskii), a stronghold of the Asiatic wild ass (Equus hemionus), and remains an important grazing area for seminomadic herders. We show the power of simple inventory and monitoring methods to assess herder-wildlife conflicts, by combining data of: (1) human and livestock demographic data, (2) migration patterns, of semi-nomadic herders, (3) monthly surveys of wild- and domestic ungulates, and (4) …

Kiangs (Equus Kiang, Moorcroft 1841) In Sikkim, India, Natalia V. Paklina, Chris Van Orden

Erforschung biologischer Ressourcen der Mongolei / Exploration into the Biological Resources of Mongolia, ISSN 0440-1298

Between the 5^th and 10^th of March 2001 we completed a 450 km route by car in Northern Sikkim. During the trip along the southern slopes of the Himalayas and Sikkim Plateau we didn’t see a single kiang. According to the local people and military personnel in Sikkim, kiangs are known only in the outer limits of the Sikkim Plateau, where wild animals have a possibility to migrate free over the Chinese-Indian border. In winter only small groups of kiang incidentally come to the Sikkim Plateau from South Tibet. But in May more of them come there and …

Current Status Of The Khulan (Equus Hemionus) In The Trans-Altai Gobi, Enkhbileg Dulamtseren, Adiya Yadamjav, Dovchindorj Ganbold, Mijidorj Batmunkh

Erforschung biologischer Ressourcen der Mongolei / Exploration into the Biological Resources of Mongolia, ISSN 0440-1298

Since 1975, 25 wildlife surveys have been conducted in the Trans-Altai Gobi. In this paper we summarize results from the last 11 surveys, conducted between October 2000 and April 2005, and discuss our findings together with results from previous surveys.

Assessment Of Non-Metric Skull Characters And Age Determination In The Asiatic Wild Ass Equus Hemionus: A Methodological Approach, Hermann Ansorge, Annegret Stubbe, Nyamsuren Batsaikhan, Ravčigijn Samjaa, Michael Stubbe

Erforschung biologischer Ressourcen der Mongolei / Exploration into the Biological Resources of Mongolia, ISSN 0440-1298

The report presents the first information on the use of non-metric skull characters and age determination by tooth section for Asiatic wild asses. It was initiated by a collection of more than 400 skulls of the Asiatic wild ass from the Bordzongijn-Gobi 2002-2004. The methodological overview gives a first research prospect in Asiatic Wild Ass concerning the use of non-metric skull characters as a morphological tool for assessing genetic variability, determining epigenetic divergences and evaluating fluctuating asymmetry. The time course of tooth eruption and replacement in the Asiatic wild ass is described with regard to its use for age determination. …

Territorial Behaviour Of Kiang (Equus Kiang Moorcroft, 1841) In Ladakh (India), Natalia V. Paklina, Chris Van Orden

Erforschung biologischer Ressourcen der Mongolei / Exploration into the Biological Resources of Mongolia, ISSN 0440-1298

The observations of kiang behavior were made in Navokar Valley northeast of Tso Kar Lake (Ladakh, India) between July 30 and November 22, 2001. In the breeding season (end of July until the end of August) adult kiang males kept not overlapping, protected territories (about 10 km²), and marked by single defecation and urination marks. There were adult females with and without offspring on the territories (up to 12 animals, including the male). The distance between male and females on the territories was usually hundred times bigger, then a distance between stallion and his harem in horses. Females …

Semi-Wild Population Of Kulans In The Bukhara Breeding Centre And Their Co-Habitation With Przewalski’S Horses, Karim Bahloul, Olga B. Pereladova, Natalia V. Soldatova, Ekaterina V. Sidorenko, Antoine J. Sempere

Erforschung biologischer Ressourcen der Mongolei / Exploration into the Biological Resources of Mongolia, ISSN 0440-1298

Asiatic wild asses and Przewalski's horses initially inhabited steppe, semi-desert and desert areas, but Przewalski's horses became extinct in the wild, and kulans are under threat of disappearance. The Bukhara Breeding Centre (Uzbekistan) was created in 1976 for conservation and reintroduction of wild ungulate species. In 1977-1978, five kulans (two males and three females) from Barsa-Kelmes Island on the Aral lake were introduced to the reserve. The group increased to 25-30 animals in 1995-1998, when five Przewalski's horses from Moscow and St. Petersburg zoos were introduced to the same territory. We analyzed the home ranges, preferred habitats and social interactions …

Social Networks In Wild Asses: Comparing Patterns And Processes Among Populations, Daniel I. Rubenstein, Siva Sundaresan, Ilya Fischhoff, David Saltz

Erforschung biologischer Ressourcen der Mongolei / Exploration into the Biological Resources of Mongolia, ISSN 0440-1298

Asiatic wild asses inhabit some of the most arid environments in the world. All live in fissionfusion societies, but demography varies and the deserts in which they live often differ in subtle ways. Characterizing details of social structure of wild ass populations has been a challenge and has made it difficult to determine causes and consequences of any differences that might exist. We use network theory to compare the social structures of two populations of Asiatic asses/ onagers inhabiting the Negev desert, Israel and khur of the Little Rann of Kuch, India and show that populations differ in important structural …

An Overview On The State Of Equus Hemionus In Whole China, Weikang Yang

Erforschung biologischer Ressourcen der Mongolei / Exploration into the Biological Resources of Mongolia, ISSN 0440-1298

The wild ass in China is divided into two species: Gobi khulan (Equus hemionus) and kiang (Equus kiang). Maps and charts of the distribution and population sizes in China and Mongolia of each species are shown from the 1980s through the early 2000s.

Review Of Jewish Tradition And The Challenge Of Darwinism, Geoffery Cantor And Marc Swetlitz, Eds., Daniel M. Stoebel

All HMC Faculty Publications and Research

This is a book review.

Distribution And Conservation Genetics Of The Cow Knob Salamander, Plethodon Punctatus Highton (Caudata: Plethodontidae), Matthew R. Graham

Theses, Dissertations and Capstones

Being lungless, plethodontid salamanders respire through their skin and are especially sensitive to environmental disturbances. Habitat fragmentation, low abundance, extreme habitat requirements, and a narrow distribution of less than 70 miles in length, makes one such salamander, Plethodon punctatus, a species of concern (S1) in West Virginia. To better understand this sensitive species, day and night survey hikes were conducted through ideal habitat and coordinate data as well as tail tips (10 to 20 mm in length) were collected. DNA was extracted from the tail tips and polymerase chain reaction (PCR) was used to amplify mitochondrial 16S rRNA gene fragments. …

Serial Reversal Learning And The Evolution Of Behavioral Flexibility In Three Species Of North American Corvids (Gymnorhinus Cyanocephalus, Nucifraga Columbiana, Aphelocoma Californica), Alan B. Bond, Alan Kamil, Russell P. Balda

Papers in Behavior in Biological Sciences

In serial reversal learning, subjects learn to respond differentially to 2 stimuli. When the task is fully acquired, reward contingencies are reversed, requiring the subject to relearn the altered associations. This alternation of acquisition and reversal can be repeated many times, and the ability of a species to adapt to this regimen has been considered as an indication of behavioral flexibility. Serial reversal learning of 2-choice discriminations was contrasted in 3 related species of North American corvids: pinyon jays (Gymnorhinus cyanocephalus), which are highly social; Clark’s nutcrackers (Nucifraga columbiana), which are relatively solitary but specialized for …