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The Systematic Position Of The Scorpion Genera Heteroscorpion Birula, 1903 And Urodacus Peters, 1861 (Scorpiones: Scorpionoidea), Michael E. Soleglad, Victor Fet, František Kovařík Dec 2016

The Systematic Position Of The Scorpion Genera Heteroscorpion Birula, 1903 And Urodacus Peters, 1861 (Scorpiones: Scorpionoidea), Michael E. Soleglad, Victor Fet, František Kovařík

Victor Fet

Stockwell (1989), in an unpublished revision, suggested the elevation of subfamily Urodacinae to the family rank, and considered Heteroscorpioninae a subfamily of Ischnuridae. Heteroscorpioninae was formally elevated to the family rank by Lourenço (1996a), and Urodacinae, by Prendini (2000). As a result of a detailed cladistic analysis, Prendini (2000, 2003b) considered families Heteroscorpionidae and Urodacidae to be sister groups, although an al-ternative topology was available. Soleglad & Fet (2003b) questioned the results of Prendini (2000) but nevertheless retained the monophyly of Heteroscorpion and Urodacus pending more detailed analysis; they recognized two valid monotypic subfamilies, Heteroscorpioninae and Urodacinae, under Urodacidae. Our …


The First Record Of Upper Permian And Lower Triassic Scorpions From Russia (Chelicerata: Scorpiones), Victor Fet, Dmitry E. Shcherbakov, Michael E. Soleglad Dec 2016

The First Record Of Upper Permian And Lower Triassic Scorpions From Russia (Chelicerata: Scorpiones), Victor Fet, Dmitry E. Shcherbakov, Michael E. Soleglad

Victor Fet

Several small fragments of fossil scorpions are reported from two localities in Vologda Province, Russia, representing the Upper Permian (Severodvinian, correlated to Wuchiapingian) (Isady) and Lower Triassic just above the Permian-Triassic boundary (Induan) (Nedubrovo). Most observed structures are not diagnostic at genus or family level. The Isady leg fragment possesses ungues (claws), which are both denticulated and setaceous, and resembles a Carboniferous Eobuthus sp. (Eobuthidae). It is the latest record of this type of ungues, which are known in some Paleozoic scorpions (extinct suborder Mesoscorpiones); all extant scorpions have smooth claws without denticulation or setation.


Taxonomic Position Of The Genus Simonoides Vachon Et Farzanpay, 1987, And Description Of A New Species Of Orthochirus Karsch From Iran (Scorpiones, Buthidae), František Kovařík, Victor Fet Dec 2016

Taxonomic Position Of The Genus Simonoides Vachon Et Farzanpay, 1987, And Description Of A New Species Of Orthochirus Karsch From Iran (Scorpiones, Buthidae), František Kovařík, Victor Fet

Victor Fet

Simonoides Vachon et Farzanpay, 1987 is synonymized with Orthochirus Karsch, 1891. Lectotype is designated for Simonoides farzanpayi Vachon et Farzanpay, 1987. Orthochirus sobotniki Kovařík, 2004 is synonymized with Orthochirus farzanpayi (Vachon et Farzanpay, 1987), comb. n. Orthochirus gruberi, sp. n. (Iran) is described and dis-tinguished from all other Iranian species of the genus on a combination of two characters, the presence of rows of granules with internal and external granules on the movable fingers of pedipalps and hirsuteness of the entire me-tasoma and telson.


The Genus Auyantepuia González-Sponga, 1978 Is Confirmed As A Synonym Of Broteochactas Pocock, 1893 (Scorpiones: Chactidae), Michael E. Soleglad, Victor Fet Dec 2016

The Genus Auyantepuia González-Sponga, 1978 Is Confirmed As A Synonym Of Broteochactas Pocock, 1893 (Scorpiones: Chactidae), Michael E. Soleglad, Victor Fet

Victor Fet

The genus Auyantepuia González-Sponga, 1978 is confirmed as a synonym of Broteochactas Pocock, 1893 (Scorpiones: Chactidae). It was correctly synonymized with Broteochactas by Francke & Boos (1986) and listed as such by Soleglad & Fet (2003) but resurrected by Lourenço & Araújo (2004). We reconfirm that the generic name Auyantepuia, based on the type species Broteochactas scorzai Dagert, 1957 from Venezuela, is a synonym of Broteochactas, based on the type species Broteochactas nitidus Pocock, 1893 from Trinidad. All other seven “short-fingered” species from French Guiana and Brazil, classified by Lourenço & Araújo (2004) under Auyantepuia, belong to the genus Neochactas …


Serrula In Retrospect: A Historical Look At Scorpion Literature (Scorpiones: Orthosterni), Matthew R. Graham, Victor Fet Dec 2016

Serrula In Retrospect: A Historical Look At Scorpion Literature (Scorpiones: Orthosterni), Matthew R. Graham, Victor Fet

Victor Fet

Serrula, a small comb-like structure on the ventral aspect of cheliceral movable finger in scorpions, has often been overlooked or ignored. Scorpion genera that are currently known to possess serrula belong to six families: Chactidae, Euscorpiidae, Iuridae, Pseudochactidae, Superstitioniidae, and Vaejovidae. All relevant literature on scorpion serrula is reviewed, and serrula of representative scorpion taxa is illustrated by SEM images. We suggest that the inconsistency in reporting serrula is based on several factors (or combination thereof): (a) serrula tines can quite often be broken off partially or completely, especially in adults; (b) serrula can be vestigial and hardly visible under …


Taxonomic Position Of The Genus Sassanidotus Farzanpay, 1987 (Scorpiones: Buthidae), František Kovařík, Victor Fet Dec 2016

Taxonomic Position Of The Genus Sassanidotus Farzanpay, 1987 (Scorpiones: Buthidae), František Kovařík, Victor Fet

Victor Fet

Sassanidotus Farzanpay, 1987 is a valid genus closely related to Compsobuthus Vachon, 1949, with which it shares most characters including the shape of carinae on the carapace and mesosoma. The main difference is that Sassanidotus has only three granules located proximally to the terminal granule on the movable finger of pedipalp, whereas Compsobuthus and also Mesobuthus Vachon, 1950 have four such granules. Sassanidotus Farzanpay, 1987 includes two species, Sassanidotus zarudnyi (Birula, 1900) (type species) from Iran and Sassanidotus gracilis thus zarudnyi sarghadensis Birula, 1903, syn. n.) from Afghanistan, Iran, and Pakistan. The neotype is designated for S. zarudnyi.


Scorpions Of Iran (Arachnida: Scorpiones). Part Vii. Kerman Province, Shahrokh Navidpour, Majeed Ezatkhah, František Kovařík, Michael E. Soleglad, Victor Fet Dec 2016

Scorpions Of Iran (Arachnida: Scorpiones). Part Vii. Kerman Province, Shahrokh Navidpour, Majeed Ezatkhah, František Kovařík, Michael E. Soleglad, Victor Fet

Victor Fet

Thirteen species of scorpions belonging to two families are reported from the Kerman Province of Iran. Of these, the species Compsobuthus kaftani Kovařík, 2003, Mesobuthus macmahoni (Pocock, 1900), Orthochirus farzanpayi (Vachon et Farzanpay, 1987), Polisius persicus Fet, Capes et Sissom, 2001, Sassanidotus gracilis (Birula, 1900), and Hemiscorpius lepturus Peters, 1861 are recorded from the province for the first time. Kerman Province contains type localities of six species of scorpions, of which Kraepelinia palpator (Birula, 1903) and Orthochirus gruberi Kovařík et Fet, 2006 are valid. Prionurus crassicauda orientalis Birula, 1900 is a synonym of Androctonus crassicauda (Olivier, 1807), Buthus eupeus kirmanensis …


Scorpions Of Iran (Arachnida, Scorpiones). Part Ix. Hormozgan Province, With A Description Of Odontobuthus Tavighiae Sp. N. (Buthidae), Shahrokh Navidpour, Michael E. Soleglad, Victor Fet, František Kovařík Dec 2016

Scorpions Of Iran (Arachnida, Scorpiones). Part Ix. Hormozgan Province, With A Description Of Odontobuthus Tavighiae Sp. N. (Buthidae), Shahrokh Navidpour, Michael E. Soleglad, Victor Fet, František Kovařík

Victor Fet

Twenty species of scorpions belonging to three families are reported from the Hormozgan Province of Iran. Of these, eight species and subspecies are recorded from the province for the first time: Buthacus macrocentrus (Ehrenberg, 1828), Compsobuthus persicus Navidpour et al., 2008, Iranobuthus krali Kovařík, 1997, Mesobuthus eupeus persicus (Pocock, 1899), Mesobuthus phillipsii (Pocock, 1889), Odontobuthus bidentatus Lourenço & Pézier, 2002, Odontobuthus doriae (Thorell, 1876), and Razianus zarudnyi (Birula, 1903). Odontobuthus tavighiae sp. n. is described and compared with all species of the genus Odontobuthus Vachon, 1950. Paraorthochirus goyffoni Lourenco et Vachon, 1995 is synonymized with Orthochirus farzanpayi (Vachon et Farzanpay, …


Scorpions Of Iran (Arachnida, Scorpiones). Part V. Chahar Mahal & Bakhtiyari Province, Khodadad Pirali-Kheirabadi, Shahrokh Navidpour, Victor Fet, František Kovařík, Michael E. Soleglad Dec 2016

Scorpions Of Iran (Arachnida, Scorpiones). Part V. Chahar Mahal & Bakhtiyari Province, Khodadad Pirali-Kheirabadi, Shahrokh Navidpour, Victor Fet, František Kovařík, Michael E. Soleglad

Victor Fet

Ten species of scorpions belonging to three families are reported from the montane Chahar Mahal & Bakhtiyari Province of Iran. Of these, only Razianus zarudnyi (Birula, 1903) was formerly reported for this province, while nine species are reported for the first time: Androctonus crassicauda (Olivier, 1807), Compsobuthus matthiesseni (Birula, 1905), Hottentotta saulcyi (Simon, 1880), Hottentotta zagrosensis Kovařík, 1997, Mesobuthus eupeus phillipsii (Pocock, 1889), Odontobuthus doriae (Thorell, 1876), Orthochirus zagrosensis Kovařík, 2004, Hemi- scorpius lepturus Peters, 1861, and Scorpio maurus townsendi (Pocock, 1900). Also presented is a key to all species of scorpions found in the Chahar Mahal & Bakhtiyari Province.


Scorpions Of Iran (Arachnida: Scorpiones). Part Vi. Lorestan Province, Shahrokh Navidpour, Hassan H. Nayebzadeh, Michael E. Soleglad, Victor Fet, František Kovařík, Mohammad Hassan Kayedi Dec 2016

Scorpions Of Iran (Arachnida: Scorpiones). Part Vi. Lorestan Province, Shahrokh Navidpour, Hassan H. Nayebzadeh, Michael E. Soleglad, Victor Fet, František Kovařík, Mohammad Hassan Kayedi

Victor Fet

Ten species of scorpions belonging to three families are reported from the Lorestan Province of Iran. Of these, five species are recorded from the province for the first time: Hottentotta zagrosensis Kovařík, 1997; Mesobuthus eupeus phillipsii (Pocock, 1889); Orthochirus iranus Kovařík, 2004; Razianus zarudnyi (Birula, 1903) ; and Scorpio maurus townsendi (Pocock, 1900). One new species is described, Hottentotta lorestanus sp. n.; it can be easily distinguished from the other four species of the genus known from Iran by its coloration; it is the only Iranian species which has the entire pedipalps yellow and the metasomal segments I to IV …


Scorpions Of Iran (Arachnida: Scorpiones). Part Vi. Lorestan Province, Shahrokh Navidpour, Hassan H. Nayebzadeh, Michael E. Soleglad, Victor Fet, František Kovařík, Mohammad Hassan Kayedi Dec 2016

Scorpions Of Iran (Arachnida: Scorpiones). Part Vi. Lorestan Province, Shahrokh Navidpour, Hassan H. Nayebzadeh, Michael E. Soleglad, Victor Fet, František Kovařík, Mohammad Hassan Kayedi

Victor Fet

Ten species of scorpions belonging to three families are reported from the Lorestan Province of Iran. Of these, five species are recorded from the province for the first time: Hottentotta zagrosensis Kovařík, 1997; Mesobuthus eupeus phillipsii (Pocock, 1889); Orthochirus iranus Kovařík, 2004; Razianus zarudnyi (Birula, 1903); and Scorpio maurus townsendi (Pocock, 1900). One new species is described, Hottentotta lorestanus sp. n.; it can be easily distinguished from the other four species of the genus known from Iran by its coloration; it is the only Iranian species which has the entire pedipalps yellow and the metasomal segments I to IV greenish …


Permian Scorpions From The Petrified Forest Of Chemnitz, Germany, Jason A. Dunlop, David A. Legg, Paul L. Selden, Victor Fet, Joerg W. Schneider, Ronny Rößler Dec 2016

Permian Scorpions From The Petrified Forest Of Chemnitz, Germany, Jason A. Dunlop, David A. Legg, Paul L. Selden, Victor Fet, Joerg W. Schneider, Ronny Rößler

Victor Fet

Background: Paleozoic scorpions (Arachnida: Scorpiones) have been widely documented from the Carboniferous Period; which hosts a remarkable assemblage of more than sixty species including both putative stem- and crown-group fossils. By contrast the succeeding Permian Period is almost completely devoid of records, which are currently restricted to a trace fossil from the early Permian of New Mexico, USA and some limb fragments from the late Permian of the Vologda Region, Russia. Results: ?Opsieobuthus tungeri sp. nov. from the Petrified Forest of Chemnitz, Germany represents the first complete body fossils of scorpions from the Permian. Explosive volcanism preserved these remarkable specimens …


Scorpions Of Iran (Arachnida, Scorpiones). Part Iv. Kohgilouyeh & Boyer Ahmad Province, Shahrokh Navidpour, František Kovařík, Michael E. Soleglad, Victor Fet Dec 2016

Scorpions Of Iran (Arachnida, Scorpiones). Part Iv. Kohgilouyeh & Boyer Ahmad Province, Shahrokh Navidpour, František Kovařík, Michael E. Soleglad, Victor Fet

Victor Fet

According to previous surveys, only Mesobuthus eupeus phillipsii (Pocock, 1889), Orthochirus zagrosensis Kovařík, 2004, and Hemiscorpius lepturus Peters, 1861 have been recorded from Kohgilouyeh & Boyer Ahmad Province. Collections made by a team under Shahrokh Navidpour (Razi Reference Laboratory of Scorpion Research, Razi Vaccine and Serum Research Institute, Ahvaz, Khoozestan, Iran) reveal eight other species belonging to two families recorded from the province for the first time: Androctonus crassicauda (Olivier, 1807); Compsobuthus matthiesseni (Birula, 1905); Hottentotta saulcyi (Simon, 1880); Hottentotta zagrosensis Kovařík, 1997; Odontobuthus bidentatus Lourenço et Pézier, 2002; Orthochirus iranus Kovařík, 2004; Razianus zarudnyi (Birula, 1903); and Scorpio maurus …


Scorpions Of Iran (Arachnida, Scorpiones). Part Iii. Ilam Province, Shahrokh Navidpour, Victor Fet, František Kovařík, Michael E. Soleglad Dec 2016

Scorpions Of Iran (Arachnida, Scorpiones). Part Iii. Ilam Province, Shahrokh Navidpour, Victor Fet, František Kovařík, Michael E. Soleglad

Victor Fet

Seven species of scorpions belonging to three families have been previously recorded from the Ilam Province of Iran: Androctonus crassicauda (Olivier, 1807), Compsobuthus matthiesseni (Birula, 1905), Hottentotta saulcyi (Simon, 1880), Mesobuthus eupeus phillipsii (Pocock, 1889), Razianus zarudnyi (Birula, 1903), Scorpio maurus townsendi (Pocock, 1900), and Hemiscorpius lepturus Peters, 1861. Collections made by a team organized by Shahrokh Navidpour (Razi Reference Laboratory of Scorpion Research, Razi Vaccine and Serum Research Institute, Ahvaz, Khoozestan, Iran) reveal seven other species recorded from the province for the first time: Apistobuthus susanae Lourenço, 1998, Buthacus macrocentrus (Ehrenberg, 1828), Compsobuthus jakesi Kovařík, 2003, Odontobuthus bidentatus Lourenço …


Scorpions Of Iran (Arachnida, Scorpiones). Part Ii. Bushehr Province, Shahrokh Navidpour, Michael E. Soleglad, Victor Fet, František Kovařík Dec 2016

Scorpions Of Iran (Arachnida, Scorpiones). Part Ii. Bushehr Province, Shahrokh Navidpour, Michael E. Soleglad, Victor Fet, František Kovařík

Victor Fet

The Bushehr Province of Iran contains the type localities of three scorpion taxa, Mesobuthus eupeus phillipsii (Pocock, 1889), Orthochirus iranus Kovařík, 2004, and Scorpio maurus townsendi (Pocock, 1900). Apart from them, Androctonus crassicauda (Olivier, 1807), Compsobuthus matthiesseni (Birula, 1905), Hottentotta saulcyi (Simon, 1880), Odontobuthus bidentatus Lourenço & Pézier, 2002, Razianus zarudnyi (Birula, 1903), and Hemiscorpius lepturus Peters, 1862 have also been known to occur in the province. Collections made by a team under Shahrokh Navidpour (Razi Reference Laboratory of Scorpion Research, Razi Vaccine and Serum Research Institute, Ahvaz, Khoozestan, Iran) reveal four additional species recorded from the province for the …


Scorpions Of Iran (Arachnida, Scorpiones). Part I. Khoozestan Province, Shahrokh Navidpour, František Kovařík, Michael E. Soleglad, Victor Fet Dec 2016

Scorpions Of Iran (Arachnida, Scorpiones). Part I. Khoozestan Province, Shahrokh Navidpour, František Kovařík, Michael E. Soleglad, Victor Fet

Victor Fet

Collections made by a team of Shahrokh Navidpour (Razi Reference Laboratory of Scorpion Research, Razi Vaccine and Serum Research Institute, Ahvaz, Khoozestan, Iran) include 17 of the 19 species known to inhabit Khoozestan Province, and form the basis of this paper. Among them are two new species (Hottentotta khoozestanus sp. n. and Vachoniolus iranus sp. n.), Compsobuthus jakesi Kovařík, 2003 previously known only from Iraq, and five species representing first records for the province: Buthacus macrocentrus (Ehrenberg, 1828); Odontobuthus bidentatus Lourenço & Pézier, 2002; Orthochirus farzanpayi (Vachon et Farzanpay, 1987); Orthochirus stockwelli (Lourenço et Vachon, 1995) comb. n. …


Etudes On Iurids, I. The Orthobothriotaxic Pattern Of Iuridae, With Observations On Neobothriotaxy In Genus Iurus (Scorpiones: Iuroidea), Michael E. Soleglad, František Kovařík, Victor Fet Dec 2016

Etudes On Iurids, I. The Orthobothriotaxic Pattern Of Iuridae, With Observations On Neobothriotaxy In Genus Iurus (Scorpiones: Iuroidea), Michael E. Soleglad, František Kovařík, Victor Fet

Victor Fet

A unique orthobothriotaxic pattern of family Iuridae is examined and illustrated. Significant positional differences in this pattern between two monotypic iurid genera, Calchas and Iurus, are presented. With the analysis of over 100 specimens of genus Iurus, representing many localities in Greece and Turkey, several unique occurrences of neobothriotaxy involving both the chela and patella are discussed and illustrated, representing the first report of neobothriotaxy in family Iuridae.


Contributions To Scorpion Systematics. Ii. Stahnkeini, A New Tribe In Scorpion Family Vaejovidae (Scorpiones: Chactoidea), Michael E. Soleglad, Victor Fet Dec 2016

Contributions To Scorpion Systematics. Ii. Stahnkeini, A New Tribe In Scorpion Family Vaejovidae (Scorpiones: Chactoidea), Michael E. Soleglad, Victor Fet

Victor Fet

The diagnostic characters originally established by Herbert L. Stahnke (1940a, 1940b, 1974) in his description of genus Serradigitus are studied in detail from several new perspectives. A new genus, Stahnkeus, gen. nov., is described based on the presence of inner accessory (IAD) denticles on the chelal fingers, unprecedented in family Vaejovidae. Five species of Serradigitus are transferred to Stahnkeus: Stahnkeus harbisoni (Williams, 1970), comb. nov. (=Serradigitus harbisoni); Stahnkeus deserticola (Williams, 1970), comb. nov. (=Serradigitus deserticola); Stahnkeus subtilimanus (Soleglad, 1972), comb. nov. (=Serradigitus subtilimanus); Stahnkeus allredi (Sissom et Stockwell, 1991), comb. nov. (=Serradigitus allredi); and Stahnkeus polisi (Sissom et Stockwell, 1991), …


Dna Markers Confirm Presence Of Euscorpius Avcii Tropea Et Al., 2012 (Scorpiones: Euscorpiidae) On Samos Island, Greece, Aristeidis Parmakelis, Panayiota Kotsakiozi, Gioele Tropea, Ersen Aydın Yağmur, Iasmi Stathi, Victor Fet, Michael E. Soleglad Dec 2016

Dna Markers Confirm Presence Of Euscorpius Avcii Tropea Et Al., 2012 (Scorpiones: Euscorpiidae) On Samos Island, Greece, Aristeidis Parmakelis, Panayiota Kotsakiozi, Gioele Tropea, Ersen Aydın Yağmur, Iasmi Stathi, Victor Fet, Michael E. Soleglad

Victor Fet

Euscorpius avcii Tropea et al., 2012 has been recently described from Dilek Peninsula in western Anatolia (Turkey, Aydın Province). The population from Samos Island in eastern part of the Aegean Sea is found to match closely the Anatolian E. avcii, making it a new, rare species for the Greek fauna, confirmed by two DNA markers as well as morphology. Samos also shares with western Anatolia two other local recently described scorpion species, Iurus kinzelbachi and Neocalchas gruberi (family Iuridae).


Constellation Array In Scorpion Genera Paruroctonus, Smeringurus, Vejovoidus, And Paravaejovis (Scorpiones: Vaejovidae), Victor Fet, Michael E. Soleglad, Michael Brewer, David P. A. Neff, Michael L. Norton Dec 2016

Constellation Array In Scorpion Genera Paruroctonus, Smeringurus, Vejovoidus, And Paravaejovis (Scorpiones: Vaejovidae), Victor Fet, Michael E. Soleglad, Michael Brewer, David P. A. Neff, Michael L. Norton

Victor Fet

The constellation array (a recently discovered sensory structure on the fixed finger of pedipalp; Fet et al., 2006) is analyzed for a large set of species belonging to four genera in the family Vaejovidae: Paruroctonus, Smeringurus, Vejovoidus, and Paravaejovis. It is shown that this structure is an important taxonomic character. Two distinct configurations are identified, a two-sensilla array for Paruroctonus + Smeringurus + Vejovoidus and a three-sensilla array for genus Paravejovis, both differing from other vaejovid configurations so far investigated. The topology of these two array configurations are analyzed using landmark setae identified in this study.


A New Scorpion Genus (Scorpiones: Vaejovidae) From Mexico, Michael E. Soleglad, Victor Fet Dec 2016

A New Scorpion Genus (Scorpiones: Vaejovidae) From Mexico, Michael E. Soleglad, Victor Fet

Victor Fet

A new vaejovid genus from Mexico, Franckeus, gen. nov., is described based on unique neobothriotaxy. Species placed in this genus are from the Vaejovis “nigrescens” group (previously called the “nitidulus” group). Six species comprise this new genus, distributed throughout Mexico (mainland as well as Baja California Sur): Franckeus nitidulus, F. rubrimanus, F. platnicki, F. minckleyi, F. kochi and F. peninsularis. A new species of the Vaejovis “nigrescens” group, Vaejovis davidi, sp. nov., is also described.


Contributions To Scorpion Systematics. Iii. Subfamilies Smeringurinae And Syntropinae (Scorpiones: Vaejovidae), Michael E. Soleglad, Victor Fet Dec 2016

Contributions To Scorpion Systematics. Iii. Subfamilies Smeringurinae And Syntropinae (Scorpiones: Vaejovidae), Michael E. Soleglad, Victor Fet

Victor Fet

Stockwell (1989), in an unpublished revision, suggested splitting the family Vaejovidae in two subfamilies, introducing also four tribes and four new genera. These results have not been reevaluated for 20 years. Here, we establish a new subfamily Smeringurinae, subfam. nov., with two tribes, four genera, and 40 species and subspecies. The subfamily Smeringurinae includes two tribes: Smeringurini, trib. nov. and Paravaejovini, trib. nov. The tribe Smeringurini includes three genera: Paruroctonus Werner, 1934; Smeringurus Haradon, 1983; and Vejovoidus Stahnke, 1974. The tribe Paravaejovini includes the genus Paravaejovis Williams, 1980. We restore from synonymy the subfamily Syntropinae Kraepelin, 1905, and significantly expand …


A New Trichobothrial Character For The High-Level Systematics Of Buthoidea (Scorpiones: Buthida), Victor Fet, Soleglad E. Michael, Graeme Lowe Dec 2016

A New Trichobothrial Character For The High-Level Systematics Of Buthoidea (Scorpiones: Buthida), Victor Fet, Soleglad E. Michael, Graeme Lowe

Victor Fet

The high-level systematics and phylogeny of the scorpion superfamily Buthoidea are unresolved. A new, formerly undetected character is reported here: the relative alignment of d3 trichobothrium and DMc carina on the pedipalp patella. This and other characters are discussed in the context of possible high-level divisions of Buthoidea. Based on this analysis, we suggest a breakdown of Buthoidea into six proposed tentative phylogenetic groups: Buthus group, Ananteris group, Isometrus group, Charmus group, Uroplectes group, and Tityus group.


Proceedings Of The 3rd Scorpiology Symposium (American Arachnological Society, 28th Annual Meeting), Victor Fet, Michael E. Soleglad Dec 2016

Proceedings Of The 3rd Scorpiology Symposium (American Arachnological Society, 28th Annual Meeting), Victor Fet, Michael E. Soleglad

Victor Fet

Table of Contents Victor Fet and Douglas D. Gaffin. Preface p. iKaren C. Bost and Douglas D. Gaffin. Sand scorpion home burrow navigation in the laboratory p. 1Douglas D. Gaffin and Mark E. Walvoord. Scorpion peg sensilla: are they the same or are they different? p. 7J. Zachary Porterfield, Douglas D. Gaffin, Caitlin Porterfield and Curtis Johnston. Screening for scorpions: A non-invasive approach to tracking the movements of arachnids in sand p. 17Tsunemi Yamashita. Surface activity, biomass, and phenology of the striped scorpion, Centruroides vittatus (Scorpiones: Buthidae) in Arkansas, USA p. 25C. Neal McReynolds. Temporal patterns in microhabitat use for …