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- Mixed-species (3)
- Phylogenetics (3)
- Publications (3)
- B. Biodiversity Informatics and Species Distribution Modeling (2)
- Biodiversity (2)
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- C. Land-use Change Modeling (2)
- Conservation (2)
- D. Biodiversity Conservation (2)
- Deforestation (2)
- GIS (2)
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- Lumbricus rubellus (2)
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- Nuthatches (2)
- Nutrient element (2)
- Peer-reviewed publications (2)
- Vermicomposting (2)
- niche (ecology) (1)
- Windmill Islands. (1)
- 4 - Evolutionary Biology & Paleontology (1)
- A. Climate Change and Sea-level Change (1)
- Academic Works (1)
- Accounting (1)
- Acoustic detection (1)
- Alkali specialist (1)
- Allozyme analysis (1)
- Ant (1)
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- Edgar Lehr (5)
- Professor Jerome K Vanclay (5)
- Shaily Menon (5)
- John E. McCormack (3)
- Maxine G Schmidt (3)
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- Amy L. Russell (2)
- Kari E. Veblen (2)
- Adi Ainurzaman Jamaludin (1)
- Alessandro Tavoni (1)
- Andrei Alyokhin (1)
- Daniel Gonzalez-Socoloske (1)
- Daniel Pondella (1)
- David A Bainbridge (1)
- David B Taylor (1)
- David J Depew (1)
- David W Markman (1)
- Frederick M. Cohan (1)
- Heather P. Griscom (1)
- Jonathan Balcombe, Ph.D. (1)
- Larry Clark (1)
- Max Moseley (1)
- Nicole Andréa Mathys (1)
- Noor Zalina Binti Mahmood (1)
- Paul V. Switzer (1)
- Rebecca W. Dolan (1)
- Ryan Kerney (1)
- Sharon Robinson (1)
- Steven J. Hall (1)
- T. Heath Ogden (1)
- Victoria Dreitz (1)
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Articles 1 - 30 of 50
Full-Text Articles in Life Sciences
Mixed Species Plantations: Prospects And Challenges, J Doland Nichols, Mila Bristow, Jerome K. Vanclay
Mixed Species Plantations: Prospects And Challenges, J Doland Nichols, Mila Bristow, Jerome K. Vanclay
Professor Jerome K Vanclay
About 2% of English-language literature on plantations deals with mixed-species plantations, but only a tiny proportion (<0.1%) of industrial plantations are polycultures. Small landholders are more innovative, with 12% of Australia’s farm forestry plantations under mixed-species plantings, and 80% of Queensland’s farm forestry as polycultures. We examine reasons for this discrepancy, and explore the history, silviculture and economics of polycultures. Financial analyses suggest that a yield stimulus of 10%, depending on product and rotation length, may be sufficient to offset increased costs associated with planting and managing a mixed-species plantation, a stimulus that has been demonstrated in many field …
Genetic Structure Of East Antarctic Populations Of The Moss Ceratodon Purpureus, L. J. Clarke, D. J. Ayre, Sharon A. Robinson
Genetic Structure Of East Antarctic Populations Of The Moss Ceratodon Purpureus, L. J. Clarke, D. J. Ayre, Sharon A. Robinson
Sharon Robinson
The capacity of the polar flora to adapt is of increasing concern given current and predicted environmental change in these regions. Previous genetic studies of Antarctic mosses have been of limited value due to a lack of variation in the markers or non-specificity of the methods used. We examined the power of five microsatellite loci developed for the cosmopolitan moss Ceratodon purpureus to detect genetically distinct clones and infer the distribution of clones within and among populations from the Windmill Islands, East Antarctica. Our microsatellite data suggest extraordinarily high levels of variation reported in RAPD studies were artificially elevated by …
Mixed-Species Plantation Of Eucalyptus With Nitrogen Fixing Trees: A Review, David I. Forrester, Jürgen Bauhus, Annette L. Cowie, Jerome K. Vanclay
Mixed-Species Plantation Of Eucalyptus With Nitrogen Fixing Trees: A Review, David I. Forrester, Jürgen Bauhus, Annette L. Cowie, Jerome K. Vanclay
Professor Jerome K Vanclay
Mixed-species plantations of Eucalyptus with a nitrogen (N2) fixing species have the potential to increase productivity while maintaining soil fertility, compared to Eucalyptus monocultures. However, it is difficult to predict combinations of species and sites that will lead to these benefits. We review the processes and interactions occurring in mixed plantations, and the influence of species or site attributes, to aid the selection of successful combinations of species and sites. Successful mixtures, where productivity is increased over that of monocultures, have often developed stratified canopies, such that the less shade-tolerant species overtops the more shade-tolerant species. Successful mixtures also have …
Growth And Species Interactions Of Eucalyptus Pellita In A Mixed And Monoculture Plantation In The Humid Tropics Of North Queensland, Mila Bristow, Jerome K. Vanclay, Lyndon O. Brooks, Mark Hunt
Growth And Species Interactions Of Eucalyptus Pellita In A Mixed And Monoculture Plantation In The Humid Tropics Of North Queensland, Mila Bristow, Jerome K. Vanclay, Lyndon O. Brooks, Mark Hunt
Professor Jerome K Vanclay
This study investigated whether mixed-species designs can increase the growth of a tropical eucalypt when compared to monocultures. Monocultures of Eucalyptus pellita (E) and Acacia peregrina (A) and mixtures in various proportions (75E:25A, 50E:50A, 25E:75A) were planted in a replacement series design on the Atherton Tablelands of north Queensland, Australia. High mortality in the establishment phase due to repeated damage by tropical cyclones altered the trial design. Effects of experimental designs on tree growth were estimated using a linear mixed effects model with restricted maximum likelihood analysis (REML). Volume growth of individual eucalypt trees were positively affected by the presence …
Forest Dynamics In Flood Plain Forests In The Peruvian Amazon: Effects Of Disturbance And Implications For Management, Gustav Nebel, Lars Peter Kvist, Jerome K. Vanclay, Hector Vidaurre
Forest Dynamics In Flood Plain Forests In The Peruvian Amazon: Effects Of Disturbance And Implications For Management, Gustav Nebel, Lars Peter Kvist, Jerome K. Vanclay, Hector Vidaurre
Professor Jerome K Vanclay
Forest dynamics were studied from 1993 to 1997 for individuals > 10 cm DBH in nine 1 ha permanent sample plots. They were established in natural flood plain forests located on the lower Ucayali river in the Peruvian Amazon. After inventories of three plots in each of three forest types, a light and a heavy felling treatment were applied to each of the two plots, while a third plot was kept untreated. Average annual stem mortality and recruitment rates in the untreated plots were among the highest observed in neotropical rain forests: mortality 2.2-3.2% per year, recruitment 3.0-4.6% per year. Dead …
Experiment Designs To Evaluate Inter- And Intra-Specific Interactions In Mixed Plantings Of Forest Trees, Jerome K. Vanclay
Experiment Designs To Evaluate Inter- And Intra-Specific Interactions In Mixed Plantings Of Forest Trees, Jerome K. Vanclay
Professor Jerome K Vanclay
A review of three mixed-species trials reveals the utility of competition indices for evaluating inter- and intra-specific interactions between trees, the desirability of experiments that span a range of tree spacing and composition to inform calibration of these competition indices, the need for extremes of species composition and stand density to calibrate response surfaces, and the far-reaching impact of edge-effects. Experiment layouts commonly used for mixed-species trials in forestry (such as replacement series) rarely provide a strong basis to calibrate competition indices and response surfaces. Alternative designs involving systematic changes in species composition may offer a better basis for calibrating …
Maximum Likelihood Estimates Of Species Trees: How Accuracy Of Phylogenetic Inference Depends Upon The Divergence History And Sampling Design, John Mccormack, Huateng Huang, L. Knowles
Maximum Likelihood Estimates Of Species Trees: How Accuracy Of Phylogenetic Inference Depends Upon The Divergence History And Sampling Design, John Mccormack, Huateng Huang, L. Knowles
John E. McCormack
The understanding that gene trees are often in discord with each other and with the species trees that contain them has led researchers to methods that incorporate the inherent stochasticity of genetic processes in the phylogenetic estimation procedure. Recently developed methods for species-tree estimation that not only consider the retention and sorting of ancestral polymorphism but also quantify the actual probabilities of incomplete lineage sorting are expected to provide an improvement over earlier summary-statistic based approaches that discard much of the information content of gene trees. However, these new methods have yet to be tested on truly challenging evolutionary histories …
Life History, Sexual Dimorphism And 'Ornamental' Feathers In The Mesozoic Bird Confuciusornis Sanctus., Winfried S. Peters, Dieter Stefan Peters
Life History, Sexual Dimorphism And 'Ornamental' Feathers In The Mesozoic Bird Confuciusornis Sanctus., Winfried S. Peters, Dieter Stefan Peters
Winfried S. Peters
Deforestation In The Tropics: Reconciling Disparities In Estimates For India, Shaily Menon, Kamaljit S. Bawa
Deforestation In The Tropics: Reconciling Disparities In Estimates For India, Shaily Menon, Kamaljit S. Bawa
Shaily Menon
Here we examine recent disparate estimates of deforestation obtained for India. We discuss the sources of disparity and the implications of inaccurate estimates and suggest ways in which future attempts at estimating deforestation might reconcile the disparity. Despite the importance of deforestation and its consequences, no attempt has been made to reconcile the different estimates obtained for India.
Identifying Conservation-Priority Areas In The Tropics: A Land-Use Change Modeling Approach, Shaily Menon, R. Gil Pontius Jr., Joseph Rose, M. Khan, Kamaljit Bawa
Identifying Conservation-Priority Areas In The Tropics: A Land-Use Change Modeling Approach, Shaily Menon, R. Gil Pontius Jr., Joseph Rose, M. Khan, Kamaljit Bawa
Shaily Menon
Most quantitative methods for identifying conservation-priority areas require more detailed knowledge about the extent and distribution of biodiversity than is currently available. Accelerated and irreversible losses of biodiversity call for the development of alternative methods to identify priority sites for biodiversity inventory and protection. We focused on the state of Arunachal Pradesh, a biodiversity-rich region in northeast India. We used a geographic information system and spatially explicit modeling to examine the correlation of land-cover and land-use patterns with biogeophysical characteristics and to project future patterns of land-use change. In 1988, 70% of Arunachal Pradesh was covered by forest. We project …
Projected Climate Change Effects On Nuthatch Distribution And Diversity Across Asia, Shaily Menon, M. Zafar-Ul Islam, A. Townsend Peterson
Projected Climate Change Effects On Nuthatch Distribution And Diversity Across Asia, Shaily Menon, M. Zafar-Ul Islam, A. Townsend Peterson
Shaily Menon
We used ecological niche modeling approaches to explore climate change implications for one family of birds, the Sittidae, in Asia. Quantitative niche models based on present-day distributions for each of 13 species were projected onto future climate change scenarios. Species’ potential distributional areas tended to be predicted to retract along their fringes, and at lower elevations along mountain ranges. As observed in other studies, montane systems were relatively more robust to the horizontal effects of climate change on species’ distributions compared to flatland systems, so range contractions were focused in Southeast Asia and peninsular India.
Preliminary Analysis Of The Ecology And Geography Of The Asian Nuthatches (Aves: Sittidae), Shaily Menon, Zafar-Ul Islam, Jorge Soberon, A. Townsend Peterson
Preliminary Analysis Of The Ecology And Geography Of The Asian Nuthatches (Aves: Sittidae), Shaily Menon, Zafar-Ul Islam, Jorge Soberon, A. Townsend Peterson
Shaily Menon
We explored distributions of Asian nuthatch species in ecological and geographic space using ecological niche modeling based on occurrence data associated with specimens and observations. Nuthatches represent a well-defined clade occurring throughout the Northern Hemisphere, but are most diverse in southern Asia where 15 of the 24 species occur and where the lineage is believed to have evolved. Species richness was focused in a narrow east-west band corresponding to the forested parts of the Himalayas with a maximum number of nine species predicted present in these foci. The distributional predictions have a mid-elevation focus with highest species diversity between 1,000 …
E-Science @ Umass: Anticipating And Supporting E-Science Activities At The University Of Massachusetts, Maxine G. Schmidt, Rebecca Reznik-Zellen
E-Science @ Umass: Anticipating And Supporting E-Science Activities At The University Of Massachusetts, Maxine G. Schmidt, Rebecca Reznik-Zellen
Maxine G Schmidt
In March of 2008 an Ad Hoccommittee of Science Librarians from the University of Massachusetts Five Campus System convened to discuss the challenges of e-science and prepare the Libraries for their role in e-science initiatives. Three primary outcomes intended to support e-science activities emerged from the work of the Ad Hoc committee.
Road-Killed Bats, Highway Design, And The Commuting Ecology Of Bats, Amy L. Russell, Calvin M. Butchkoski, Leslie Saidak, Gary F. Mccracken
Road-Killed Bats, Highway Design, And The Commuting Ecology Of Bats, Amy L. Russell, Calvin M. Butchkoski, Leslie Saidak, Gary F. Mccracken
Amy L. Russell
Incorporating Fairness Motives Into The Impulse Balance Equilibrium And Quantal Response Equilibrium Concepts: An Application To 2x2 Games, Alessandro Tavoni
Incorporating Fairness Motives Into The Impulse Balance Equilibrium And Quantal Response Equilibrium Concepts: An Application To 2x2 Games, Alessandro Tavoni
Alessandro Tavoni
Substantial evidence has accumulated in recent empirical works on the limited ability of the Nash equilibrium to rationalize observed behavior in many classes of games played by experimental subjects. This realization has led to several attempts aimed at finding tractable equilibrium concepts which perform better empirically; one such example is the impulse balance equilibrium (Selten, Chmura, 2008), which introduces a psychological reference point to which players compare the available payoff allocations. This paper is concerned with advancing two new, empirically sound, concepts: equity-driven impulse balance equilibrium (EIBE) and equity-driven quantal response equilibrium (EQRE): both introduce a distributive reference point to …
Landscape Genetics Of California Mule Deer (Odocoileus Hemionus): The Roles Of Ecological And Historical Factors In Generating Differentiation, Katherine Pease, Adam Freedman, John Pollinger, John Mccormack, Wolfgang Buermann, Jeff Rodzen, Jim Banks, Erin Meredith, Vernon Bleich, Robert Schaefer, Ken Jones, Robert Wayne
Landscape Genetics Of California Mule Deer (Odocoileus Hemionus): The Roles Of Ecological And Historical Factors In Generating Differentiation, Katherine Pease, Adam Freedman, John Pollinger, John Mccormack, Wolfgang Buermann, Jeff Rodzen, Jim Banks, Erin Meredith, Vernon Bleich, Robert Schaefer, Ken Jones, Robert Wayne
John E. McCormack
Landscape genetics is an emerging discipline that utilizes environmental and historical data to understand geographic patterns of genetic diversity. Niche modelling has added a new dimension to such efforts by allowing species–environmental associations to be projected into the past so that hypotheses about historical vicariance can be generated and tested independently with genetic data. However, previous approaches have primarily utilized DNA sequence data to test inferences about historical isolation and may have missed very recent episodes of environmentally mediated divergence. We type 15 microsatellite loci in California mule deer and identify five genetic groupings through a Structure analysis that are …
A New Species Of Streptanthus (Brassicaceae) From Three Peaks In Lake County, California, Rebecca W. Dolan, Richard O'Donnell
A New Species Of Streptanthus (Brassicaceae) From Three Peaks In Lake County, California, Rebecca W. Dolan, Richard O'Donnell
Rebecca W. Dolan
Streptanlhus vernalis is a newly described species inhabiting serpentine rock outcrops in the Three Peaks area in Lake County, California. Morphological and allozyme data indicate that this taxon is related to the S. morrisonii complex.
Cultural Disturbances And Local Ecological Knowledge Mediate Cattail (Typha Domingensis) Invasion In Lake Pátzcuaro, México, Steven J. Hall
Cultural Disturbances And Local Ecological Knowledge Mediate Cattail (Typha Domingensis) Invasion In Lake Pátzcuaro, México, Steven J. Hall
Steven J. Hall
The influence of local actors and socioeconomic constraints on biological invasions is often ignored. Wetland plant harvesters appeared to intentionally influence cattail (Typha domingensis) invasion around Lake Pátzcuaro, México, by altering their harvesting regimes, according to interviews with 44 expert respondents and botanical surveys. The oldest and most experienced harvesters reported controlling Typha initially, sometimes through organized eradication efforts, in order to protect Schoenoplectus californicus, an economically and culturally valuable wetland plant. Later, outsiders commoditized Typha by introducing new weaving designs popular with tourists, while industrial products and new livelihood activities reduced Schoenoplectus harvest. Harvesters from several communities began to …
E-Science @ The University Of Massachusetts, Maxine G. Schmidt, Rebecca Reznik-Zellen, Raquel Rivera, Cecilia P. Mullen
E-Science @ The University Of Massachusetts, Maxine G. Schmidt, Rebecca Reznik-Zellen, Raquel Rivera, Cecilia P. Mullen
Maxine G Schmidt
e-Science @ the University of Massachusetts Abstract: What is e-Science and how can libraries and librarians support it? The University of Massachusetts takes a proactive approach to support network-enabled research on its campuses and provides examples where e-Science is already at work. Statement: “e-Science” is a term commonly used to describe research in a networked environment, a growing trend not only in the sciences, but the arts and humanities as well. e-Science creates both opportunities and challenges for academic libraries. The opportunities lie in leveraging the basic skill set that libraries and librarians already possess: the knowledge of and practical …
The Value Of A Net‐Cage As A Fish Aggregating Device In Southern California, Daniel Pondella
The Value Of A Net‐Cage As A Fish Aggregating Device In Southern California, Daniel Pondella
Daniel Pondella
There is an urgent need for primary data that can be used to quantify the value of marine aquaculture facilities that also describe influences on the surrounding natural ecosystem and its wild fish communities. Divers completed 360 transect replicates below a net-cage and at nearby and distant rocky reefs off Catalina Island, California, estimating the species abundance and size class of all conspicuous fishes in the water column. We observed 10,234 fishes aggregating below the net-cage with a mean annual density of 142 (SE ± 30) per 100 m2 and diversity H′ 2.29. At the adjacent reference reef, we counted …
California Grasslands Alkali Specialist, Hemizonia Pungenspungens, Prefers Non-Alkali Soil, Kari E. Veblen
California Grasslands Alkali Specialist, Hemizonia Pungenspungens, Prefers Non-Alkali Soil, Kari E. Veblen
Kari E. Veblen
Edaphically severe habitats commonly support edaphic endemics, specialized plant species that do not occur elsewhere. The endemism of native plant species in edaphically specialized habitats suggests either (a) that these native endemic species are uniquely specialized to survive and grow better under the conditions prevalent in these harsh areas, or (b) that these areas represent refuges from competition with other (often exotic) species.
Behavioral Explanations Underlying The Lack Of Trap Effectiveness For Small-Scale Management Of Japanese Beetles (Coleoptera: Scarabaeidae), Paul V. Switzer, Patrick C. Enstrom, Carissa A. Schoenick
Behavioral Explanations Underlying The Lack Of Trap Effectiveness For Small-Scale Management Of Japanese Beetles (Coleoptera: Scarabaeidae), Paul V. Switzer, Patrick C. Enstrom, Carissa A. Schoenick
Paul V. Switzer
Traps containing a combination floral and synthetic pheromone lure are used to monitor and manage Japanese beetles, Popillia japonica Newman (Coleoptera: Scarabaeidae). One key factor limiting trap effectiveness for beetle control is the "trap spillover" phenomenon, in which the trap attracts beetles without capturing them, resulting in increased damage to surrounding host plants. We investigated the mechanisms underlying trap spillover by conducting two studies in a soybean field in east central Illinois. In the first study, we set up trap stations for 1 d and compared the sex, size, and egg load (for females) of beetles caught in the traps …
Size Matters: Scalar Phenomena And A Proposal For An Ecological Definition Of ‘Cave’., Max Moseley
Size Matters: Scalar Phenomena And A Proposal For An Ecological Definition Of ‘Cave’., Max Moseley
Max Moseley
Caves, as ecological units, are redefined in terms of the ecologically-significant scalar phenomena that, in contrast with such traditional criteria as perpetual darkness, environmental stability and oligotrophy, are the special characteristics of large cavities (macrocaverns) in rocks. When so defined, ‘caves’ are ecologically distinct from the mesocaverns and all other subterranean habitats. Some implications of this perspective for the ecological classification of animals found in subterranean habitats are briefly discussed.
A Sustainable Future, David A. Bainbridge
A Sustainable Future, David A. Bainbridge
David A Bainbridge
A key step toward a more sustainable future is beginning the discussion of what this would entail. What are the defining characteristics of sustainability? How could we get there? What benefits and costs might this involve?
How Fast Is The World's Center Of Co2 Emissions Moving Eastwards?, Nicole Andréa Mathys, Grether Jean-Marie
How Fast Is The World's Center Of Co2 Emissions Moving Eastwards?, Nicole Andréa Mathys, Grether Jean-Marie
Nicole Andréa Mathys
Borrowing from physics the concept of center of mass and applying it to the distribution of CO2 anthropogenic sources provided by the EDGAR data base, we can draw on the Earth’s surface the trajectory of the world’s pollution center of gravity over the 1970-2005 period. It is strongly heading to the East, and more so than GDP, which suggests that Asian production is getting more CO2 intensive than Western production.
Waste Recycling: Utilization Of Coffee Grounds And Kitchen Waste In Vermicomposting., Adi Ainurzaman Jamaludin, Noor Zalina Mahmood
Waste Recycling: Utilization Of Coffee Grounds And Kitchen Waste In Vermicomposting., Adi Ainurzaman Jamaludin, Noor Zalina Mahmood
Adi Ainurzaman Jamaludin
Vermicomposting using Lumbricus rubellus for 49 days was conducted after 21 days of pre-composting. Three different combination of treatments were prepared with eight replicates for each treatment namely cow dung : kitchen waste in 30:70 ratio (T1), cow dung : coffee grounds in 30:70 ratio (T2), and cow dung : kitchen waste : coffee grounds in 30:35:35 ratio (T3). The multiplication of earthworms in terms of numbers and weight were measured at the end of vermicomposting. Consequently, only T2 showed significant increase (from it initial stage) compared to other treatments. The presence of coffee grounds in T2 and T3 showed …
Wild Bird’S-Eye View Of Influenza Virus A(H1n1), Larry Clark
Wild Bird’S-Eye View Of Influenza Virus A(H1n1), Larry Clark
Larry Clark
Wild bird fecal samples collected and characterized by the USDA as part of a national surveillance effort were sequenced to study the genetic relatedness of avian, swine, and human H1 and N1 subtypes. Our results find that the 2009 H1N1 human outbreak is closely related to swine virus, but falls into different clades in the H1 and N1 trees. Further, there is evidence of multiple viral genetic exchanges between birds and swine. Ongoing research across host species contributes to an understanding of the circulation of influenza viruses.
Waste Recycling: Feasibility Of Saw Dust Based Spent Mushroom Substrate And Goat Manure In Vermicomposting, Noor Zalina Binti Mahmood
Waste Recycling: Feasibility Of Saw Dust Based Spent Mushroom Substrate And Goat Manure In Vermicomposting, Noor Zalina Binti Mahmood
Noor Zalina Binti Mahmood
Vermicomposting for 140 days by using Lumbricus rubellus was conducted after 21 days of natural pre-composting. Five treatments in different ratio of goat manure: spent mushroom substrate were prepared as feed materials with four replicates for each treatment namely; 20:80 (TA), 40:60 (TB), 50:50 (TC), 60:40 (TD) and 80:20 (TE). As for control, each treatment without earthworm was prepared. On the basis of nutrient elements, goat manure and spent mushroom substrate can be decomposed through both methods of vermicomposting and natural composting. Findings of this study indicated that the higher usage of goat manure with longer duration resulted in the …
Parental Behavior Of A Precocial Species: Implications For Juvenile Survival, Victoria Dreitz
Parental Behavior Of A Precocial Species: Implications For Juvenile Survival, Victoria Dreitz
Victoria Dreitz
No abstract provided.
A New Method For Collecting Clean Stable Fly (Diptera: Muscidae) Pupae Of Known Age., Dennis R. Berkebile, Anthony P. Weinhold, David B. Taylor
A New Method For Collecting Clean Stable Fly (Diptera: Muscidae) Pupae Of Known Age., Dennis R. Berkebile, Anthony P. Weinhold, David B. Taylor
David B Taylor
Stable flies, Stomoxys calcitrans L., are important pests of confined and pasture cattle. They have been reared in the laboratory to study their biology and to test new methods of control. Research on rearing modifications has concentrated on developing larval diets from materials locally abundant. Under current protocols, pupae form in the medium. Aggregations of pupae were located and removed, often with a considerable amount of extraneous material. Various methods have been developed to separate the pupae from waste material. We describe a method by which wandering larvae are enticed to leave the medium prior to pupariation. The larvae were …