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Articles 1 - 30 of 331
Full-Text Articles in Life Sciences
A New Locality Of Mesobuthus Eupeus Thersites (C. L. Koch, 1839) (Scorpiones: Buthidae) In East Kazakhstan, Alexander A. Fomichev
A New Locality Of Mesobuthus Eupeus Thersites (C. L. Koch, 1839) (Scorpiones: Buthidae) In East Kazakhstan, Alexander A. Fomichev
Euscorpius
A new locality of Mesobuthus eupeus thersites (C. L. Koch, 1839) is reported, found during the fieldtrip to East Kazakhstan, one of the most northern areas where scorpions are found in Asia. Notes on the habitats, map and photographs of specimens are given.
A Checklist Of The Scorpions (Arachnida: Scorpiones) Of Panama, With Two New Records, Rolando Teruel, Michiel A.C. Cozijn
A Checklist Of The Scorpions (Arachnida: Scorpiones) Of Panama, With Two New Records, Rolando Teruel, Michiel A.C. Cozijn
Euscorpius
In the present note, we compile a checklist of all scorpion species recorded from Panama. A total of three families, five genera and 14 species are confirmed to occur in the country, and previous records of two other families, four genera, and nine species are discarded as misidentifications, mislabelings, or accidental introductions. Two Buthidae species are herein recorded for the first time from Panama: Tityus tayrona Lourenço, 1991 (so far known only from northern Colombia) and one apparently undescribed species of Ananteris Thorell, 1891.
On Centruroides Margaritatus (Gervais, 1841) And Closely Related Species (Scorpiones: Buthidae), Luis F. De Armas, Rolando Teruel, František Kovařík
On Centruroides Margaritatus (Gervais, 1841) And Closely Related Species (Scorpiones: Buthidae), Luis F. De Armas, Rolando Teruel, František Kovařík
Euscorpius
Centruroides edwardsii (Gervais, 1843) comb. nov. is restored as a valid species, and a male neotype from Riohacha, La Guajira Department, Colombia is herein designated; this species ranges from Mexico through Colombia, but there are no reliable records from Guatemala, Belize, and Panama; introduced populations also occur in Cuba (West Indies) and Senegal (Africa). We regard the following species as its junior synonyms: Scorpio (Atreus) degeerii Gervais, 1844, Centrurus gambiensis Karsch, 1879, Centruroides margaritatus septentrionalis Hoffmann, 1932, and Rhopalurus danieli Prado et Rios-Patiño, 1940. We also consider Centruroides margaritatus morenoi Mello-Leitão, 1945 as a junior synonym of …
Scorpions Of Iran (Arachnida: Scorpiones). Part Vii. Kerman Province, Shahrokh Navidpour, Majeed Ezatkhah, František Kovařík, Michael E. Soleglad, Victor Fet
Scorpions Of Iran (Arachnida: Scorpiones). Part Vii. Kerman Province, Shahrokh Navidpour, Majeed Ezatkhah, František Kovařík, Michael E. Soleglad, Victor Fet
Euscorpius
Thirteen species of scorpions belonging to two families are reported from the Kerman Province of Iran. Of these, the species Compsobuthus kaftani Kovařík, 2003, Mesobuthus macmahoni (Pocock, 1900), Orthochirus farzanpayi (Vachon et Farzanpay, 1987), Polisius persicus Fet, Capes et Sissom, 2001, Sassanidotus gracilis (Birula, 1900), and Hemiscorpius lepturus Peters, 1861 are recorded from the province for the first time. Kerman Province contains type localities of six species of scorpions, of which Kraepelinia palpator (Birula, 1903) and Orthochirus gruberi Kovařík et Fet, 2006 are valid. Prionurus crassicauda orientalis Birula, 1900 is a synonym of Androctonus crassicauda (Olivier, 1807), Buthus eupeus kirmanensis …
An Anomaly Of Pectinal Organs In Vaejovis Lapidicola (Scorpiones: Vaejovidae), Richard F. Ayrey
An Anomaly Of Pectinal Organs In Vaejovis Lapidicola (Scorpiones: Vaejovidae), Richard F. Ayrey
Euscorpius
Among specimens of Vaejovis lapidicola Stahnke, one female was found to have abnormally shaped pectines, both with only distal pectinal teeth. This rare, possibly teratological anomaly is documented and discussed.
Breeding Biology Of Oryzomys Palustris, The Marsh Rice Rat, In Eastern Virginia, Robert K. Rose, Erin A. Dreelin
Breeding Biology Of Oryzomys Palustris, The Marsh Rice Rat, In Eastern Virginia, Robert K. Rose, Erin A. Dreelin
Virginia Journal of Science
The objectives of our study were to determine the age of maturity, litter size, and the timing of the breeding season of marsh rice rats (Oryzomys palustris) of coastal Virginia. From May 1995 to May 1996, monthly samples of rice rats were live-trapped in two coastal tidal marshes of eastern Virginia, and then necropsied. Sexual maturity was attained at 30-40 g for both sexes. Mean litter size of 4.63 (N= 16) did not differ among months or in mass or parity classes. Data from two other studies conducted in the same county, one of them contemporaneous, also were …
A Review Of The Subgenus Pandinus Thorell, 1876 With Descriptions Of Two New Species From Uganda And Ethiopia (Scorpiones: Scorpionidae), František Kovařík
A Review Of The Subgenus Pandinus Thorell, 1876 With Descriptions Of Two New Species From Uganda And Ethiopia (Scorpiones: Scorpionidae), František Kovařík
Euscorpius
Pandinus (Pandinus) ugandaensis sp. n. from Uganda and P. (P.) mazuchi sp. n. from Ethiopia are described and compared with other species of the subgenus. P. ugandaensis sp. n. is characterized by 3-4/3: 3-4/3: 4/3: 4/3 spiniform formula of tarsomere II and only two spines on the inclined anteroventral surface of tarsomere II; eight ventral trichobothria on the chela; 10–11 pectinal teeth in females and 13–14 in males; and 1.6–1.7 length to depth ratio of the fourth metasomal segment. P. (P.) mazuchi sp. n. is characterized by 7/4: 6-7/4: 6-7/5-6: 8/5 spiniform formula of tarsomere II and only …
Redescription Of Centruroides Granosus (Thorell, 1876) And Identity Of Centrurus Granosus Simplex Thorell, 1876 (Scorpiones: Buthidae), Luis F. De Armas, Rolando Teruel, František Kovařík
Redescription Of Centruroides Granosus (Thorell, 1876) And Identity Of Centrurus Granosus Simplex Thorell, 1876 (Scorpiones: Buthidae), Luis F. De Armas, Rolando Teruel, František Kovařík
Euscorpius
Centruroides granosus (Thorell, 1876) is redescribed based on several specimens of both sexes from Panama. We demonstrate that the original description of this species was based on an immature (subadult) male, which is herein designated as the lectotype. We conclude that Centrurus granosus simplex Thorell, 1876 is a senior synonym of Centruroides testaceus arubensis (Bakker, 1963), and elevate it to species level: Centruroides simplex (Thorell, 1876), comb. nov.
Buthus Awashensis Sp. N. From Ethiopia (Scorpiones: Buthidae), František Kovařík
Buthus Awashensis Sp. N. From Ethiopia (Scorpiones: Buthidae), František Kovařík
Euscorpius
Buthus awashensis sp. n. from the Awash region of Ethiopia, is described and compared with B. berberensis Pocock, 1900 from Somaliland. The new species is characterized chiefly by the total length of 50–65 mm. B. berberensis reaches only 45–55 mm and differs in coloration, with adults having the chela of pedipalp and chelicerae entirely yellow without dark reticulations, and by mophometric characters, mainly the shape of the chela of pedipalp. Pectinal marginal tips extend the to proximal end of the fourth sternite in males of B. awashensis sp. n. and to proximal end of the fifth sternite in males of …
Efecto Del Almacenamiento De Huevo Incubable Con La Cámara De Aire Hacia Abajo, Sobre La Incubabilidad, Según La Edad De Las Reproductoras, Para Tres Días De Rotación, Marco Guío, Jorge Briñez, Jorge Amado, Andrés Martínez
Efecto Del Almacenamiento De Huevo Incubable Con La Cámara De Aire Hacia Abajo, Sobre La Incubabilidad, Según La Edad De Las Reproductoras, Para Tres Días De Rotación, Marco Guío, Jorge Briñez, Jorge Amado, Andrés Martínez
Revista Ciencia Animal
Actualmente, las líneas de conformación han reducido presión sobre la producción de huevos por parte de las abuelas y reproductoras pesadas, mientras se han concentrado en el rendimiento del pollo de engorde, evidenciándose mejores conversiones alimenticias en ambos sexos. Este contexto expone el hecho de que, en Colombia, las empresas más exitosas, que se encuentran entre el 5% y el 25% de productividad del país, con dificultad podrán superar los resultados históricos. Se hace necesario analizar todas las oportunidades de mejora en el manejo que contribuyan a maximizar los rendimientos zootécnicos de las aves, para obtener la mayor cantidad posible …
Hemiscorpius Novaki Sp. N. From Somaliland (Scorpiones: Hemiscorpiidae), František Kovařík, Tomáš Mazuch
Hemiscorpius Novaki Sp. N. From Somaliland (Scorpiones: Hemiscorpiidae), František Kovařík, Tomáš Mazuch
Euscorpius
Hemiscorpius novaki sp. n. from Somaliland is described and compared with other African species of the genus. The new species is characterized chiefly by the total length of 40–46 mm; yellow to yellowish-brown color with pedipalp fingers reddish brown to black, darker than the chela; slightly longer metasoma in the male than in the female. These characters distinguish H. novaki sp. n. from H. somalicus Lourenço, 2011. The third African species of the genus, H. tellinii Borelli, 1904, is known only from the female holotype, which differs from females of H. novaki sp. n. in having much less pronounced granulation …
Redescription Of Tityus Pictus Pocock, 1893 And Tityus Smithii Pocock, 1893, With Notes On The Tityus Species From The Lesser Antilles (Scorpiones: Buthidae), Rolando Teruel
Euscorpius
In the present paper, the taxonomic status of Tityus pictus Pocock, 1893 and all of its junior synonyms is revised. Tityus smithii Pocock, 1893 is restored as a valid species, and the subspecies Tityus smithii microdon Pocock, 1893 (currently T. pictus microdon) is regarded as its junior synonym. Both T. pictus and T. smithii are redescribed, supplementary information on their morphological variability are given, and their geographical distribution is up- dated, including a new locality record for the latter. Also, the taxonomic position and relationships of all Lesser Antillean species of the genus are clarified: it is demonstrated that …
Pectine Development In Scorpion Embryos And First And Second Instars, Roger D. Farley
Pectine Development In Scorpion Embryos And First And Second Instars, Roger D. Farley
Euscorpius
The pectines are complex sensory organs that extend from the ventral surface of the anterior opisthosoma (mesosoma) in all extant scorpions and nearly all the fossil scorpions that have been examined. The pectines are synapomorphic for the Order Scorpiones. In this investigation, the scanning electron microscope (SEM) was used to study the development of the pectines in representatives from five scorpion families. In the more basal families (e.g., Vaejovidae) with apoikogenic development, the pectines start to develop early with enlargement of the limb buds on the third opisthosomal segment. The primordial pectines become elongate lobes attached to the ventro-lateral surface …
In Situ Infrared Videography Of Sand Scorpion Nighttime Surface Activity, Douglas D. Gaffin
In Situ Infrared Videography Of Sand Scorpion Nighttime Surface Activity, Douglas D. Gaffin
Euscorpius
Sand scorpions make burrows from which they emerge at night to seek mates and prey. Previous ecological studies have documented scorpion activities on the sand surface. Few studies, however, have videotaped scorpion movements in the context of understanding their sensory neurobiology. Our objective is to understand which signals guide scorpions as they emerge from and return to their burrows. Candidate stimuli include chemical trails, footsteps, celestial patterns, geomagnetic cues, humidity and/or temperature gradients, distinctive scents from the burrow, seismic echolocation, memory, or landmark orientation. We videotaped scorpions to learn whether they return to their burrows by the same path they …
Scorpions In The Modern Cuban Culture: An Introductory Iconography, Luis F. De Armas
Scorpions In The Modern Cuban Culture: An Introductory Iconography, Luis F. De Armas
Euscorpius
Some graphic examples of the role of scorpions in the present-day culture of Cuba are presented. They include tattoos, jewels, carpets, caps, and drawings by children, among other objects of personal or social usage. Increased use of this arachnid as a cultural element among Cuban people during last 50 years seems to be directly related with globalization, mostly through the TV and movies.
On Fabre’S Traces: An Important Contributor To The Knowledge Of Buthus Occitanus (Amoreux, 1789), Marco Colombo
On Fabre’S Traces: An Important Contributor To The Knowledge Of Buthus Occitanus (Amoreux, 1789), Marco Colombo
Euscorpius
Jean-Henri Casimir Fabre (1823–1915) has probably been one of the most important entomologists of the world in the last two centuries, leaving to posterity a huge amount of manuscripts and books. The Languedoc yellow scorpion, Buthus occitanus (Amoreaux, 1789), has been one of his objects of study: many of its behaviours, including the famous “promenade a deux”, have been described by the careful eye of the French entomologist, giving an interesting contribution to modern scorpiology
Scorpions In Ancient Egypt, Hisham K. El-Hennawy
Scorpions In Ancient Egypt, Hisham K. El-Hennawy
Euscorpius
The ancient Egyptians knew the scorpion and its toxicity, and venerated it since pre-dynastic era. They used the scorpion as a king's name, a name of a nome (county), and a symbol to their goddess, Serqet, that protects the body and the viscera of the dead, and that accompanies them in their journey to the afterlife. They had medical prescriptions and magical spells to heal the stings. Since the 5th dynasty, the title of a “Follower of Serket” was given to clever physicians. Scorpions are most famously depicted on Horus Cippus, a talisman featuring Horus the Child holding in his …
The First Record Of Upper Permian And Lower Triassic Scorpions From Russia (Chelicerata: Scorpiones), Victor Fet, Dmitry E. Shcherbakov, Michael E. Soleglad
The First Record Of Upper Permian And Lower Triassic Scorpions From Russia (Chelicerata: Scorpiones), Victor Fet, Dmitry E. Shcherbakov, Michael E. Soleglad
Euscorpius
Several small fragments of fossil scorpions are reported from two localities in Vologda Province, Russia, representing the Upper Permian (Severodvinian, correlated to Wuchiapingian) (Isady) and Lower Triassic just above the Permian-Triassic boundary (Induan) (Nedubrovo). Most observed structures are not diagnostic at genus or family level. The Isady leg fragment possesses ungues (claws), which are both denticulated and setaceous, and resembles a Carboniferous Eobuthus sp. (Eobuthidae). It is the latest record of this type of ungues, which are known in some Paleozoic scorpions (extinct suborder Mesoscorpiones); all extant scorpions have smooth claws without denticulation or setation.
Further Considerations On The Species Of The Genus Orthochirus Karsch, 1891 From Africa, With Description Of Three New Species (Scorpiones: Buthidae), Wilson R. Lourenço, Elise-Anne Leguin
Further Considerations On The Species Of The Genus Orthochirus Karsch, 1891 From Africa, With Description Of Three New Species (Scorpiones: Buthidae), Wilson R. Lourenço, Elise-Anne Leguin
Euscorpius
New considerations are proposed regarding the African species of the genus Orthochirus Karsch, 1891. Two species, Orthochirus aristidis (Simon, 1882) and Orthochirus innesi Simon, 1910 have been the subject of several publications in the past decades; however, doubts remain about their exact identity and range of geographical distribution. In this note, their taxonomic status is reinvestigated. The type material is revised and the lectotype and paralectotypes are designated for O. aristidis. Revised diagnoses and illustrations are proposed for both species, and these are confirmed as valid. Three new species are described from Algeria, Morocco, and Mauritania. The total number …
Polymorphism And Hybridization In Species Of Hottentotta Birula, 1908 (Scorpiones: Buthidae), Wilson R. Lourenço, Eric Ythier, Mark Stockmann, John L. Cloudsley-Thompson
Polymorphism And Hybridization In Species Of Hottentotta Birula, 1908 (Scorpiones: Buthidae), Wilson R. Lourenço, Eric Ythier, Mark Stockmann, John L. Cloudsley-Thompson
Euscorpius
A new and well documented case of hybridization among scorpions is presented. It was obtained under laboratory conditions between Hottentotta jayakari (Pocock) and Hottentotta salei (Vachon) specimens of which had been collected in the northern and southern regions of Oman. Hybrids were successfully produced not only from F0 males and females, but also from F1 males and females, thereby attesting to the fact that the first generation obtained (F1), was completely fertile. Both F1 and F2 broods were composed of dark and pale morphs, indicating that the juveniles could inherit either one or …
The First Record Of The Family Euscorpiidae (Arachnida: Scorpiones) From Central China, With A Key Of Chinese Species Of The Genus Scorpiops, Zhi-Yong Di, Ya-Wen He, Zhi-Jian Cao, Ying-Liang Wu, Wen-Xin Li
The First Record Of The Family Euscorpiidae (Arachnida: Scorpiones) From Central China, With A Key Of Chinese Species Of The Genus Scorpiops, Zhi-Yong Di, Ya-Wen He, Zhi-Jian Cao, Ying-Liang Wu, Wen-Xin Li
Euscorpius
The genus Scorpiops (Euscorpiidae) is recorded for the first time in Central China. Two immature specimens of a form belonging to Scorpiops hardwickii (Gervais, 1843) “complex” were collected from Huzhaoshan Mountains in Hubei Province. A discussion of Chinese species of genus Scorpiops is provided, as well as a key of Scorpiops from China.
Investigation Of In-Package Ozonation: The Effectiveness Of Ozone To Inactive Salmonella Enteritidis On Raw, Shell Eggs, Austin Donner
Investigation Of In-Package Ozonation: The Effectiveness Of Ozone To Inactive Salmonella Enteritidis On Raw, Shell Eggs, Austin Donner
The Journal of Purdue Undergraduate Research
Food production, handling, and distribution practices pose a constant threat to the quality and safety of food products. The objective of this research is to evaluate an innovative in-package ozonation process to reduce Salmonella enteritidis on raw, shell eggs. Previous research has shown that in-package ozonation eliminates contaminants from outside sources, reduces pathogens, and extends shelf life. In this study, raw, shell eggs were inoculated with Salmonella enteritidis and exposed to ozonation treatment. Microbial recoveries were then tested to determine bacterial reductions. Measurements included: relative humidity (34 percent at 5oC), surface temperatures (oC), ozone concentrations, bacterial …
Diel Distribution And Movement Of Sound Scattering Layer In Kuroshio Waters, Northeastern Taiwan, Ming-An Lee, Ming-Hsiu Chao, Jinn-Shing Weng, Yang-Chi Lan, Hsueh-Jung Lu
Diel Distribution And Movement Of Sound Scattering Layer In Kuroshio Waters, Northeastern Taiwan, Ming-An Lee, Ming-Hsiu Chao, Jinn-Shing Weng, Yang-Chi Lan, Hsueh-Jung Lu
Journal of Marine Science and Technology
A 26-hr acoustic observation was carried out on board of the “Ocean Research II” in the Kuroshio waters at 123°09.027'E, 25°05.019'N from August 4 to 5, 2007. Acoustic volume scattering strengths (SVs) were collected by a scientific echo sounder (EK500, 38 kHz) and diel variations of sound scattering layer were analyzed. The maximum SV of surface scattering layers (SSL) around 20~80 m was observed after dawn, and mean SVs of SSL in nighttime was about -68 dB, 8 dB higher than that of daytime. Two deep scattering layers (DSL) were found in daytime at depth of 400~500 m and 600~700 …
Echo Integration Near The Seabed, Masahiko Furusawa
Echo Integration Near The Seabed, Masahiko Furusawa
Journal of Marine Science and Technology
The echo integration (EI) method has long been used for fish abundance estimations. The method is applicable to any type of fish distribution and the analysis is easy to perform by advanced software. For fish near the seabed, however, sometimes it is difficult to perform an analysis because of so-called dead zones. In this paper, the EI near the seabed is considered, reviewing past studies and introducing some new ideas such as an unsampled zone. For the consideration, close inspection of the EI theory is necessary and a section is devoted to the purpose. As an application of the EI …
Fish Acoustics: Physics-Based Modeling And Measurement, Davis Benjamin Reeder
Fish Acoustics: Physics-Based Modeling And Measurement, Davis Benjamin Reeder
Journal of Marine Science and Technology
The U.S. Office of Naval Research has sponsored research in the area of marine organism acoustics for many years. The research program has included development of theoretical physics-based acoustic scattering models of single animals, high-resolution laboratory measurements of scattering by individual animals, and at-sea field experiments. The program has been focused on the backscattered signal, but has also included investigation of the forward-scattered signal. Downward-looking acoustic surveys using ship borne echosounders rely on the backscattered signal and provide non-invasive, non-destructive, rapid, high-resolution, large area survey capability compared to traditional net tows. Horizontally-oriented acoustic surveys provide the opportunity to investigate both …
Vertical Distribution Of Walleye Pollock Juveniles Before And After The Period Of Transition For Feeding In Funka Bay, Hokkaido, Japan, Yohei Kawauchi, Osamu Shida, Hiroya Okumura, Naoki Tojo, Hiroki Yasuma, Kazushi Miyashita
Vertical Distribution Of Walleye Pollock Juveniles Before And After The Period Of Transition For Feeding In Funka Bay, Hokkaido, Japan, Yohei Kawauchi, Osamu Shida, Hiroya Okumura, Naoki Tojo, Hiroki Yasuma, Kazushi Miyashita
Journal of Marine Science and Technology
We examined the vertical distributions of walleye pollock Theragra chalcogramma juveniles for two size groups (smaller and larger than 30 mm) during the period of transition for feeding (PTF) and after this period in Funka Bay, Hokkaido, Japan. Samplings were conducted in May and June in 2006 and 2007, and water temperature and salinity were measured. The correlation between juvenile sizes and distributed depth was observed using net sampling. From this result, size groups were divided by depth. In addition, the distribution and abundance of juveniles were examined using acoustic data. In the PTF (May), juvenile size increased with increasing …
In Situ And Ex Situ Target Strength Measurement Of Mesopelagic Lanternfish, Diaphus Theta (Family Myctophidae), Kouichi Sawada, Kazuhisa Uchikawa, Tomohiko Matsuura, Hiroya Sugisaki, Kazuo Amakasu, Koki Abe
In Situ And Ex Situ Target Strength Measurement Of Mesopelagic Lanternfish, Diaphus Theta (Family Myctophidae), Kouichi Sawada, Kazuhisa Uchikawa, Tomohiko Matsuura, Hiroya Sugisaki, Kazuo Amakasu, Koki Abe
Journal of Marine Science and Technology
Acoustic target strengths (TS) of myctophid fish Diaphus theta were measured in situ at 70 kHz off the east coast of Hokkaido by tethering an acoustic-optical system (the Japanese Quantitative Echosounder and Stereo-video Camera System or J-QUESTχ) at a depth of 150 m from a research vessel. Fish length distributions were obtained from daytime hauls taken with a frame trawl after the TS measurement. Measured average TS () and a standard length (SL) in cm gave the empirical length - average TS relationship, = 20logSL70.6, assuming that the linear TS is proportional to the square of the SL. A laboratory …
Preface, Kohji Iida, Ming-An Lee, Masahiko Furusawa
Preface, Kohji Iida, Ming-An Lee, Masahiko Furusawa
Journal of Marine Science and Technology
No abstract provided.
Technology Evolution And Advances In Fisheries Acoustics, Dezhang Chu
Technology Evolution And Advances In Fisheries Acoustics, Dezhang Chu
Journal of Marine Science and Technology
Application of sonar technology to fisheries acoustics has made significant advances over recent decades. The echosounder systems evolved from the simple analog single-beam and single-frequency systems to more sophisticated digital multi-beam and multi-frequency systems. In this paper, a brief review of major technological advances in fisheries acoustics is given, as well as examples of their applications.
Measurement And Modeling Of Three Dimensional Target Strength Of Fish For Horizontal Scanning Sonar, Muhammad Kurnia, Kohji Iida, Tohru Mukai
Measurement And Modeling Of Three Dimensional Target Strength Of Fish For Horizontal Scanning Sonar, Muhammad Kurnia, Kohji Iida, Tohru Mukai
Journal of Marine Science and Technology
This paper discusses the three-dimensional target strength (3DTS) of fish for a horizontal scanning sonar. Experiments were conducted in a tank using a defrosted Japanese mackerel. Measurements were made by changing the horizontal incident angle of a 50 kHz acoustic beam and the pitch angle of a tethered fish. Following the experiment, morphological characteristics, including the swimbladder size and shape, were measured using soft X-ray and digitized. The theoretical 3DTS was estimated using the prolate spheroid model and compared with the measured 3DTS. Results showed that the TS strongly depended on the fish orientation. The maximum and minimum TS were …