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Neuronal And Behavioral Response Relationships In Virgin Females Of Acheta Domesticus, Brandon Shin Apr 2019

Neuronal And Behavioral Response Relationships In Virgin Females Of Acheta Domesticus, Brandon Shin

Honors Theses

Animals that communicate acoustically serve as models for investigating how organisms respond to sound. For instance, in field crickets, females exhibit phonotaxis (directed movement towards the sound's source) to calls with a specific frequency, intensity, and syllable period (SP) - the time between the beginning of one sound pulse and the beginning of another within the same chirp. Additionally, auditory neurons in females produce action potentials in direct response to the sound pulses of a male 's call. A previous study on the species Gryllus bimaculatus demonstrated a relationship between an individual female's behavioral and neuronal responses; however, no such …


Sex Differences In Behavioral Responses To Repeat Subconcussive Events, Rebecca J. Wilson Mar 2019

Sex Differences In Behavioral Responses To Repeat Subconcussive Events, Rebecca J. Wilson

College of Science and Health Theses and Dissertations

Although concussions, especially those in athletes and military, have become a popular focus of neurotrauma research, subconcussions occur with higher frequency and are less well-studied. A subconcussion is loosely defined as an impact to the head that does not result in a diagnosable concussion but can result in neuronal alterations. Repeat subconcussions have been shown to produce behavioral impairments along with neuropathology that is similar to or worse than those seen in a single concussion injury. These studies have primarily included male subjects. Given the potential effects of hormones and NIH’s call for sex-inclusion in biomedical research, assessing female responses …


Evaluation Of Reproductive Phenology And Ecology Of Eastern Wild Turkey (Meleagris Gallopavo Silvestris) Across The Southeastern United States, Nicholas W. Bakner Mar 2019

Evaluation Of Reproductive Phenology And Ecology Of Eastern Wild Turkey (Meleagris Gallopavo Silvestris) Across The Southeastern United States, Nicholas W. Bakner

LSU Master's Theses

Avian reproduction is known to be a driver of population dynamics across species and systems. Behavioral decisions during incubation such as habitat selection and nest attentiveness are thought to affect nest success. The incubation process is a costly period during which individuals are sedentary and must balance survival with reproductive success and overall fitness. Current understanding of Eastern wild turkey incubation behavior provides a limited view of the incubation period. Using GPS data collected from Eastern wild turkeys (n = 220), I evaluated nest fate as it relates to recess frequency, distance travelled during recess, and habitat selection during …


Ecology, Monogamy, And The Evolution Of Animal Families, Jacqueline Rae Dillard Jan 2019

Ecology, Monogamy, And The Evolution Of Animal Families, Jacqueline Rae Dillard

Theses and Dissertations--Biology

Family-living has been recognized as a necessary prerequisite for the evolution of advanced cooperative societies, yet the evolutionary and ecological processes that drive the coupling of different forms of cooperation in family-based societies are still poorly understood. In my dissertation, I investigate the correlated evolution of parental care, monogamy, and cooperative breeding in a variety of family-based taxa. I explore the mating and social behavior of family-living beetles with incipient cooperation to better understand the factors driving these social traits. Specifically, I evaluate different causes of extra-pair mating in socially monogamous beetles, the potential benefits that young adult offspring may …


Determinants Of Social Behavior In Captive North American River Otters (Lontra Canadensis), Olivia R. Beasley Dec 2018

Determinants Of Social Behavior In Captive North American River Otters (Lontra Canadensis), Olivia R. Beasley

Graduate Theses

Field Studies of North American River Otters (Lontra canadensis), documented social behavior atypical among its mustelid relatives. In the wild, river otters are most active during crepuscular hours and males have been shown to be more social than females, as they cooperatively forage within bachelor groups (Blundell et al. 2002). Most social behavior occurs at latrine sites, where feces act as a means of communication between conspecifics (Blundell et al. 2005). While scientists have conducted some behavioral research on wild river otters, detailed studies of interactions are difficult for a species that ranges widely. A full understanding of river otter …


Bubble Stream Production By Belugas (Delphinapterus Leucas), Megan Slack Aug 2018

Bubble Stream Production By Belugas (Delphinapterus Leucas), Megan Slack

Theses

Bubble stream production in belugas has been poorly characterized and its function is not well understood. I examined behavioral states when producing bubble streams (“bubbling”), and when bubbling calls, to determine whether bubbling was significantly associated with a particular call category or behavioral state. Using 19 hours of video and audio recordings collected over a two-day period, I quantified bubble streams of a 4-month old calf and an unrelated adult female housed together. Based on the overall activity budgets and pool of vocalizations for both animals, I calculated the expected counts of bubble streams with and without vocalizations, assuming that …


A Note On The Onset Of Synchronous Egg Laying In A Seabird Behavior Model, Dorothea Gallos, Christiane Gallos Apr 2018

A Note On The Onset Of Synchronous Egg Laying In A Seabird Behavior Model, Dorothea Gallos, Christiane Gallos

Honors Theses

Protection Island, Washington hosts a large colony of Glaucous-winged gulls (Larus glaucescens). These birds are known to exhibit every-other-day egg-laying synchrony in dense areas of the colony in response to egg cannibalism, which is the main source of egg loss. Here we present an equilibrium analysis of a discrete-time animal behavior model for egg laying. We use Jury Conditions to find the stability criteria for the equilibrium as a function of the colony density and show that a 2-cycle bifurcation occurs when the equilibrium loses stability. The 2-cycle pattern in egg laying becomes increasingly synchronous as the colony density increases. …


Grooming Behaviors Of, Lauren N. Williams Mar 2018

Grooming Behaviors Of, Lauren N. Williams

USF Tampa Graduate Theses and Dissertations

The giant freshwater shrimp, Macrobrachium rosenbergii is a large species of prawn grown extensively in aquaculture settings. A social hierarchy exists within the males of this species, representing three distinct male morphotypes. These male morphotypes differ in their behavior, physiology, and morphology and include the largest blue-clawed males (BC males), moderately- sized orange-clawed males (OC males), and the undifferentiated small-clawed males (SM males). All individuals of this species perform grooming behaviors to rid themselves of body fouling which can impede important functions such as movement, respiration, chemoreception, and reproduction. Grooming behaviors in crustaceans often utilize specialized structures called setae, which …


Learning And Behavior In Hatchling Trachemys Scripta Exposed To Bisphenol-A During Embryonic Development, Justin E. Dillard Feb 2018

Learning And Behavior In Hatchling Trachemys Scripta Exposed To Bisphenol-A During Embryonic Development, Justin E. Dillard

Theses and Dissertations

Steroids play an integral role in orchestrating embryonic development, and can affect a suite of phenotypic traits, including learning and memory. Endocrine disrupting compounds (EDCs) can mimic the effects of steroids and can affect the same suites of phenotypic traits during embryonic development. Bisphenol-A (BPA) is an EDC that mimics the action of estrogen, and recent work is beginning to implicate BPA in effects on learning and behavior similar to those caused by estrogen treatment studies. Red-eared slider turtles (Trachemys scripta) are a good system to investigate the effects of BPA on learning and behavior, both because the molecular underpinnings …


Effects Of Roundup On Behavior, Growth, And Mortality Of Larval Blue Dashers, Pachydiplax Longipennis, Kayleen K. Parker, Claire A. Fuller Jan 2018

Effects Of Roundup On Behavior, Growth, And Mortality Of Larval Blue Dashers, Pachydiplax Longipennis, Kayleen K. Parker, Claire A. Fuller

Murray State Theses and Dissertations

The objective of this study was to determine if Roundup (active ingredient: Glyphosate) causes negative effects on behavior, growth, and mortality of larval Pachydiplax longipennis. Larvae were captured from rainwater-filled mesocosms at Hancock Biological Station in Murray, KY. Larvae were exposed to one of four concentrations of Roundup (0mg/L, 2.5mg/L, 5mg/L, or 10mg/L). Daphnia consumption, seek refuge, and anti-predator trials were conducted at 7 and 14 days post-exposure. Growth and survival trials were carried out for eight weeks using different larvae. There were no significant differences among treatments for whether or not larvae ate offered Daphnia for Day 7 …


Leaping Behavior In Silver Carp (Hypophthalmichthys Molitrix): Analysis Of Burst Swimming Speeds, Angle Of Escape, Height, And Distance Of Leaps, Ehlana Stell Jan 2018

Leaping Behavior In Silver Carp (Hypophthalmichthys Molitrix): Analysis Of Burst Swimming Speeds, Angle Of Escape, Height, And Distance Of Leaps, Ehlana Stell

Electronic Theses and Dissertations

Silver Carp have rapidly expanded their range exploiting vulnerable habitats, disrupting fisheries, and inflicting unknown ecological damage. These fish have continued to spread into the Middle Mississippi River and the Tennessee River Valley and great effort is being expended to prevent Silver Carp from entering the Great Lakes and expanding further into the Ohio, Illinois, Missouri, and Tennessee Rivers. Using boat-mounted cameras, we recorded in situ video of invasive Silver Carp, Hypopthalmichthys molitrix leaps to measure their horizontal distance, height, and angle of escape as well as their burst speed. Video tapes of fish leaps were obtained from populations of …


The Effects Of Environmental Variability And Latitude On Breeding Behaviors And Life History Traits Of Aves In North America, Elizabeth E. Tharman May 2017

The Effects Of Environmental Variability And Latitude On Breeding Behaviors And Life History Traits Of Aves In North America, Elizabeth E. Tharman

Master's Theses

Breeding systems among Aves have garnered attention in the last several decades as many species considered monogamous have been found to exhibit an alternative breeding system such as polygyny, polyandry, cooperative breeding, or colonial nesting. My thesis focuses on assessing the correlations between type of breeding system and resource availability by using environmental variables as proxy variables. It also assess the correlations between life history trait variables and environmental variables. Both breeding system and life history traits are analyzed at the species and population levels while controlling for phylogenetic relationships and geographic location when appropriate. Breeding system among species was …


Determinants Of Host Use In Tachinid Parasitoids (Diptera: Tachinidae) Of Stink Bugs (Hemiptera: Pentatomidae) In Southwest Ohio, Matthew W. Duncan Jan 2017

Determinants Of Host Use In Tachinid Parasitoids (Diptera: Tachinidae) Of Stink Bugs (Hemiptera: Pentatomidae) In Southwest Ohio, Matthew W. Duncan

Browse all Theses and Dissertations

Tachinid parasitoids in the subfamily Phasiinae are important natural enemies of heteropteran bugs. Host location by these flies occurs via antennal reception to the pheromones of their hosts; however little is known regarding the mechanisms which underlie host selection. Halyomorpha halys, the invasive brown marmorated stink bug, represents a potential novel host species in North America. This study was conducted to determine the suitability of H. halys as a host for phasiine species, and to assess cues used in host selection by the species Gymnoclytia occidua. Field attraction to pentatomid pheromones by both phasiines and pentatomids in Southwest Ohio were …


Barn Owls (Tyto Alba) Crossing The Road - Examining The Interplay Among Occupancy, Behavior, Habitat Selection, And Roadway Mortality In Southern Idaho, Tempe Regan Dec 2016

Barn Owls (Tyto Alba) Crossing The Road - Examining The Interplay Among Occupancy, Behavior, Habitat Selection, And Roadway Mortality In Southern Idaho, Tempe Regan

Boise State University Theses and Dissertations

Barn Owls (Tyto alba) are killed by vehicle collisions in greater numbers than any other North American bird of prey. Interstate-84 (I-84) in southern Idaho, USA has among the world’s highest known rates of Barn Owl-vehicle collisions. Little is known about Barn Owl occupancy in this region, so it is unclear if owls are killed in proportion to their abundance, or if they are equally abundant in segments with lower mortality and somehow escape collisions. Furthermore, studies of Barn Owl movements and behavior are limited. I was interested in understanding (1) factors that affect Barn Owl occupancy in …


The Effect Of Auditory Stimulation On Sleep Disruption In West Indian Manatee (Trichechus Manatus Latirostris), Natalija Lace Aug 2016

The Effect Of Auditory Stimulation On Sleep Disruption In West Indian Manatee (Trichechus Manatus Latirostris), Natalija Lace

Dissertations

Florida manatees inhabit waterways where motorized boats are common. Although manatee mortalities resulting from boat strikes are well documented, the effect of boat noise on some manatee behaviors, including rest, has not been investigated. This study focuses on rest behavior and used a playback experiment with four manatees at the Lowry Park Zoo in Florida. We tested their responses to playback stimuli of either boat noise, silence, or manatee calls. A playback trial was initiated when the focal animal showed behavioral characteristics of rest.

Results showed that rest was interrupted in response to the playback of boat noise for each …


The Molecular And Cellular Basis For Cold Sensation, Daniel Brenner May 2016

The Molecular And Cellular Basis For Cold Sensation, Daniel Brenner

Arts & Sciences Electronic Theses and Dissertations

The ability to sense changes in temperature is crucial to surviving harsh environments. Over the last decade several ion channels that have been proposed to be cold sensitive have been identified, most notably TRPM8 and TRPA1. Although these molecules have been extensively studied in vitro, their exact roles in cold sensation in vivo are still debated. This uncertainty is in large part due to problems with the standard methods of testing cold sensitivity in vivo, which often rely on subjective measures of cold responsiveness. Experiments using these subjective measures have been repeated by different groups and have yielded conflicting results, …


Anti-Predator Responses Of Squid Throughout Ontogeny, Carly Anne York Apr 2016

Anti-Predator Responses Of Squid Throughout Ontogeny, Carly Anne York

Biological Sciences Theses & Dissertations

Multiple sensory modalities and a complex array of escape behaviors have evolved as components of anti-predator responses in squids. The goals of this study include: (1) examine the role of the lateral line analogue and vision in successful predator evasion; (2) measure kinematics of escape jetting; (3) document how chromatic patterning, posturing and inking in squid change in response to predators; and (4) investigate escape jet hydrodynamics of squid. Given that squids undergo considerable morphological, ecological, and behavioral changes throughout ontogeny, the goals above were all investigated across different life history stages. To test the respective roles of vision and …


Behavioral Responses By Cottonmouths (Agkistrodon Piscivorus) To Chemical And Visual Predator Cues, Kristen Lee Kohlhepp Jan 2016

Behavioral Responses By Cottonmouths (Agkistrodon Piscivorus) To Chemical And Visual Predator Cues, Kristen Lee Kohlhepp

MSU Graduate Theses

Predator detection and assessment of predation risk have important survival consequences for animals. However, responses to predatory threats can vary with different stimuli. Chemical cues are important for predator recognition but visual cues may elicit defensive responses. Cottonmouths (Agkistrodon piscivorus) exhibit an array of antipredatory behaviors that have been thoroughly characterized in response to human aggressors, but their responses to cues from other predators are poorly known. I conducted three experiments to evaluate cottonmouth responses to visual and chemical stimuli from predators and non-predators. Snakes exposed to predator chemical cues exhibited elevated tongue-flick rates compared to controls but did not …


The Wetted Ramp As A Useful Tool To Service Smaller-Bodied Finfishes At Low-Head Acquatic Barriers, Levi Kivari Jan 2016

The Wetted Ramp As A Useful Tool To Service Smaller-Bodied Finfishes At Low-Head Acquatic Barriers, Levi Kivari

Master's Theses and Doctoral Dissertations

This study examined the effects of finfish length and behavior on passage success on the wetted ramp, a proposed passage device for finfishes at low-head aquatic barriers. There were two hypotheses: there is a size effect to passage success because larger fish have relatively less propulsive surface area in partial submersion, and fish respond to changes in ramp angle of inclination during a passage attempt. Creek chub (Semotilus atromaculatus) and white sucker (Catostomus commersonii) were observed, using a high speed camera, as they attempted to scale the wetted ramp. Smaller fish traveled farther and were also able to accelerate on …


Examining Sociological Differences And The Influence Of Prey Distribution And Environmental Variability In The Distribution Of A Top Marine Predator, The Bottlenose Dolphin (Tursiops Truncatus), Stefanie K. Gazda Dec 2015

Examining Sociological Differences And The Influence Of Prey Distribution And Environmental Variability In The Distribution Of A Top Marine Predator, The Bottlenose Dolphin (Tursiops Truncatus), Stefanie K. Gazda

Graduate Doctoral Dissertations

The purpose of this dissertation was to examine the influence of environmental variability on the distribution of prey, and the influence of prey spatial structure and habitat variability may have on the distributions of bottlenose dolphins (Tursiops truncatus). Additionally I examined how sociological differences (behavior type and the changes in a foraging behavior specific to Cedar Key Florida) influences the relative roles of bottlenose dolphins within the population.

The Gowans et al. scheme assumes that small groups form small communities and that foraging groups are small and rare as there are few foraging benefits to promote grouping. Using network analysis, …


The Behavioral Response Of Culex Erraticus To Different Snake Odors, Lindsey E. Wells Jun 2015

The Behavioral Response Of Culex Erraticus To Different Snake Odors, Lindsey E. Wells

Honors College Theses

Eastern equine encephalitis virus (EEEV) is an arbovirus that can cause fatal infections in humans and horses. Unfortunately, the transmission mechanisms of this virus are still largely unknown. Culex erraticus displays a strong potential for serving as a vector of EEEV because of its indiscriminate feeding pattern and abundance in areas with the highest prevalence of infection. However, Culex erraticus is incapable of over-wintering the virus, yet EEE recurs each spring. Snakes may play an important role in over-wintering the virus, and certain snake species may be infected more frequently than others. This study was conducted to determine if Culex …


Bats And Disease: Behavioral And Community Responses Of Southern Bat Populations During The White-Nose Syndrome Epizootic, Riley Fehr Bernard May 2015

Bats And Disease: Behavioral And Community Responses Of Southern Bat Populations During The White-Nose Syndrome Epizootic, Riley Fehr Bernard

Doctoral Dissertations

This dissertation investigates regional differences in the behavior and activity of bats in eastern North America during the white-nose syndrome epizootic, specifically in the understudied region of the Southeastern United States. An introductory section provides a brief review of the history of white-nose syndrome, an emerging infectious disease in bats, and its introduction into North America. Chapter one provides the first documented evidence of bat activity outside of hibernacula throughout winter. The research presented in chapter two attempts to explain the variation in load and prevalence of P. destructans among species, sites and between years. Finally, chapter three examines the …


Establishing The Dance Floor: Frame Manipulation Experiments, Peter D. Suich May 2015

Establishing The Dance Floor: Frame Manipulation Experiments, Peter D. Suich

Undergraduate Honors Theses

Past studies of honey bee populations, in both natural and laboratory settings have allowed researchers to elucidate the dance language of honey bees within the hive. While the intent and meaning of the waggle dance is thoroughly understood, the area within the hive on which the bees dance is poorly understood. Several factors that may contribute to waggle dancing were studied: substrate, scent and hive entrance proximity. Two separate honey bee colonies were placed in three-frame observation hives. After establishing the dance floor, new experimental conditions were introduced by changing the position of the frames and watching for three days …


The Effect Of Predator Positioning And Related Angles On The Flight-Initiation Distance And Refuge Choice Of The Eastern Gray Squirrel., Lindsay Nason May 2015

The Effect Of Predator Positioning And Related Angles On The Flight-Initiation Distance And Refuge Choice Of The Eastern Gray Squirrel., Lindsay Nason

College of Arts & Sciences Senior Honors Theses

The purpose of this study was to determine the effect that the relative positions of squirrel, predator, and refuges, as well as the predator’s SPT angle (See Figure 1), had on a squirrel’s choice of refuge and the length of its flight initiation distance. A running person was used to simulate an attacking predator, and the relative positions of squirrel, person, and refuges were measured using both distance and compass angle. Data were collected in September and October 2014 at several urban green spaces in Louisville, Kentucky. The SPT angle measurement was not found to be particularly useful in predicting …


Chemical And Behavioral Ecology Of The Coastal Giant Salamander (Dicamptodon Tenebrosus), David Reavill Jan 2015

Chemical And Behavioral Ecology Of The Coastal Giant Salamander (Dicamptodon Tenebrosus), David Reavill

All Master's Theses

Intraspecific interactions are an important factor in shaping the population structure of terrestrial salamanders. Both physical interactions and scent-marking are vital components to the establishment of territories and influence dispersion of individuals. However, little is known about the terrestrial interactions of many salamander species, particularly that of the Coastal Giant Salamander (Dicamptodon tenebrosus). In this study, I observed the first evidence for the use chemical cues by terrestrial D. tenerosus in regards to shelter selection. I also observed the first accounts of agonistic and display behavior of terrestrial D. tenebrosus. These results suggest that terrestrial D. tenebrosus …


Logging Debris Protects Sugar Maple (Acer Saccharum) Seedlings From White-Tailed Deer (Odocoileus Virginianus) Herbivory In Wolf-Occupied Forest, Ari Erin Sullivan Jan 2015

Logging Debris Protects Sugar Maple (Acer Saccharum) Seedlings From White-Tailed Deer (Odocoileus Virginianus) Herbivory In Wolf-Occupied Forest, Ari Erin Sullivan

Browse all Theses and Dissertations

White-tailed deer are a species of great economic and ecological concern. Foresters sometimes leave logging debris known as slash on the forest floor with the intent to protect seedlings from deer herbivory and promote forest regeneration. I examined the effects of slash on rates of deer browsing on sugar maple seedlings in a forest of northern Wisconsin and measured deer foraging behavior using giving-up density and vigilance rates by employing trail cameras. Rates of browsed stems were almost twice as high in the open as within and adjacent to slash. These findings underscore the usefulness of slash for mitigating the …


Determining The Critical Window Of Influence Of Pcb Perinatally On Behavioral And Hormonal Development In Sprague-Dawley Rat Pups, Natalie Sommerville, Lee A. Meserve, Howard C. Cromwell Dec 2014

Determining The Critical Window Of Influence Of Pcb Perinatally On Behavioral And Hormonal Development In Sprague-Dawley Rat Pups, Natalie Sommerville, Lee A. Meserve, Howard C. Cromwell

Honors Projects

Polychlorinated biphenyls (PCB) had widespread use in the United States in commercial manufacturing in the United States until the late 1970s. Even though they were banned, measurable amounts can still be found in the environment and food sources. PCB has known effects on altering hormone development and behavior in the species Rattus norvegicus. To determine the most crucial developmental time of exposure to PCB in Sprague-Dawley rat pups, rat pups were exposed to PCB at differing weeks of either gestation period or the first postnatal week. Behavioral tests were performed for the different rat pups, as well as blood …


Genetic Background Influences Behavior And Responses To Epigenetic Changes Induced By A Methyl-Donor Diet, Kimberly R. Shorter Aug 2014

Genetic Background Influences Behavior And Responses To Epigenetic Changes Induced By A Methyl-Donor Diet, Kimberly R. Shorter

Theses and Dissertations

With recent strides in epigenetics, mainstream media informs the public that we can “beat our genes” by, for instance, changing our diet. Genetics, however, still plays a role in phenotype. Folate and other methyl-donor pathway components are widely supplemented due to their ability to prevent neural tube defects during prenatal development. In addition to vitamins, these compounds are also added to commercial flour, energy drinks, and other supplements. Several lines of evidence suggest that these supplements act through epigenetic mechanisms, including altering DNA methylation. Increasing evidence suggests potential deleterious effects of excessive folate. Given the benefits of these compounds, risk …


Early Developmental Impacts On Male Traits And Female Preference In Zebra Finches (Taeniopygia Guttata), Martyna Boruta Jun 2014

Early Developmental Impacts On Male Traits And Female Preference In Zebra Finches (Taeniopygia Guttata), Martyna Boruta

USF Tampa Graduate Theses and Dissertations

Some male sexually selected traits are sensitive to stressors early in life and provide females with information to discriminate among males with different developmental experiences. Moreover, female early life experiences could also impact which males they choose. Females might either choose honest traits indicative of male quality, no matter their own experiences, or they might choose mates to match or compensate for their own experiences. To determine how developmental stressors alter male sexually-selected traits and female preference thereof, I exposed zebra finches (Taeniopygia guttata, ZEFI) to i) lipopolysaccharide (LPS), an immunogenic, Gram-negative bacterial component, ii) corticosterone (CORT), an avian stress …


The Behavioral And Physiological Effects Of Long-Distance Translocation On Western Rattlesnakes (Crotalus Oreganus), Kory Hayden Heiken Dec 2013

The Behavioral And Physiological Effects Of Long-Distance Translocation On Western Rattlesnakes (Crotalus Oreganus), Kory Hayden Heiken

Master's Theses

Long-distance translocation (LDT), the relocation of an animal outside of its home range, is a popular strategy for mitigating conflict between humans and venomous snakes. While LDT has been demonstrated to prevent a snake’s return to the location of capture, it may result in increased mortality, magnitude and frequency of movements, and activity range sizes. Thus, it has generally been discouraged. However, the effects of LDT on stress physiology and thermoregulation have gone largely unstudied in reptiles. To elucidate these effects, we conducted an experimental LDT on Western Rattlesnakes (Crotalus oreganus) on Vandenberg Air Force base in California. …