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1981

Ecology and Evolutionary Biology

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Full-Text Articles in Life Sciences

Sensory Head Pores And Canals In Goodeid Fishes, John Michael Fitzsimons Dec 1981

Sensory Head Pores And Canals In Goodeid Fishes, John Michael Fitzsimons

Occasional Papers of the Museum of Natural Science, Louisiana State University

No abstract provided.


Geographic Variation In The Scrub Euphonia, Robert W. Dickerman Dec 1981

Geographic Variation In The Scrub Euphonia, Robert W. Dickerman

Occasional Papers of the Museum of Natural Science, Louisiana State University

No abstract provided.


Distribution And Taxonomic Status Of Blarina Hylophaga Elliot (Insectivora: Soricidae), Sarah B. George, Jerry R. Choate, Hugh H. Genoways Dec 1981

Distribution And Taxonomic Status Of Blarina Hylophaga Elliot (Insectivora: Soricidae), Sarah B. George, Jerry R. Choate, Hugh H. Genoways

University of Nebraska State Museum: Mammalogy Papers

Systematic relationships of southern populations of short-tailed shrews (genus Blarina) are assessed on the basis of univariate and multivariate statistics. Populations are separated into two phena; southwestern short-tailed shrews are significantly larger morphometrically than southeastern forms. The two phena apparently represent distinct species. The name Blarina hylophaga is available for southwestern populations, and the name Blarina carolinensis is here restricted to short-tailed shrews in the southeastern United States.


Systematic Review Of The Texas Pocket Gopher, Geomys Personatus (Mammalia: Rodentia), Stephen L. Williams, Hugh H. Genoways Dec 1981

Systematic Review Of The Texas Pocket Gopher, Geomys Personatus (Mammalia: Rodentia), Stephen L. Williams, Hugh H. Genoways

University of Nebraska State Museum: Mammalogy Papers

The Texas pocket gopher (Geomys personarus), which occupies a range in southern Texas and extreme northeastern Tamaulipas, was examined for morphological variation. Univariate and multivariate analyses were used to determine age, secondary sexual, individual, and geographic variation. Significant differences were found among the three age classes and between the sexes for 12 of 13 cranial measurements. Males displayed higher individual variation than females. Distributions of the six previously recognized subspecies (fallax, fuscus, maritimus, megapotamus, personatus, and streckeri) were examined. An additional subspecies is recognized and described. Of the seven subspecies of …


The Ecology Of The Raccoon (Procyon Lotor) In Cades Coves, Great Smoky Mountains National Park, Alan Robert Rabinowitz Dec 1981

The Ecology Of The Raccoon (Procyon Lotor) In Cades Coves, Great Smoky Mountains National Park, Alan Robert Rabinowitz

Doctoral Dissertations

From September 1979 through September 1980, 288 captures of 153 raccoons were made in Cades Cove, Great Smoky Mountains National Park, Tennessee. Fourteen raccoons were equipped with radio-transmitters to monitor movements, activity patterns, and denning behavior. Measurements, blood samples, and ectoparasites were taken from all animals.

Population density was estimated to be 1 raccoon/17 ha with sex and age ratios indicating a stable population. Parturition occurred in early June with a mean litter size of 2.8 young. Average longevity of raccoons was 30 months with a maximum longevity of approximately 86 months. There were indications of increased male mortality in …


The Paleoecology Of Carbonaceous (Algal) Material In The Middle Devonian Rockport Quarry Limestone Of The Northeastern Portion Of Michegan's Lower Peninsula, Jeffrey Dean Spruit Dec 1981

The Paleoecology Of Carbonaceous (Algal) Material In The Middle Devonian Rockport Quarry Limestone Of The Northeastern Portion Of Michegan's Lower Peninsula, Jeffrey Dean Spruit

Masters Theses

The Rockport Quarry Limestone of the Lower Traverse Group represents a sequence of laterally contemporaneous carbonate platform sediments characterized in outcrop by four facies: (l) stromatoporoid biolithite, (2) organic-mud packstone, (3) biolithite-micrite transition, and (4) micrite. In facies (l), (2) and (3) dark brown 4-micron microspheres are found grouped or singly in wispy seams of brown carbonaceous material. Three species of filamentous blue-green algae were discovered in insoluble residues of facies (l) and (2). Numerous lamellar stromatoporoids and solitary corals occur with the carbonaceous-rich sediment in facies (l). Coalescing lenses of crinoid and bryozoan debris occur with the carbonaceous sediment …


Small Mammals In Openings In Virginia's Dismal Swamp, Robert K. Rose Dec 1981

Small Mammals In Openings In Virginia's Dismal Swamp, Robert K. Rose

Biological Sciences Faculty Publications

In a study of small mammals of openings in the Dismal Swamp of Virginia, seven species were obtained using pitfall traps. Samples included several species rarely caught in the Swamp - seven specimens of the Dismal Swamp subspecies of the southern bog lemming, Synaptomys cooperi helaletes, the first collected in this century; two least shrews, Cryptotis parva; and 15 southeastern shrews, Sorex longirostris fisheri . Results are compared to previous studies, conducted primarily in forested habitats, in which the white-footed mouse, Peromyscus leucopus, and the golden mouse, Ochrotomys nuttalli, were numerically dominant.


Electrophoretic And Immunological Studies On The Relationship Of The Brachyphyllinae And The Glossophaginae, Robert J. Baker, Rodney L. Honeycutt, Michael L. Arnold, Vincent M. Sarich, Hugh H. Genoways Nov 1981

Electrophoretic And Immunological Studies On The Relationship Of The Brachyphyllinae And The Glossophaginae, Robert J. Baker, Rodney L. Honeycutt, Michael L. Arnold, Vincent M. Sarich, Hugh H. Genoways

University of Nebraska State Museum: Mammalogy Papers

Electrophoretic and albumin immunological data indicate that the Brachyphyllinae as currently conceived is a natural assemblage, with Erophylla sezekorni and Phyllonycteris aphylla being more closely related to each other than either is to Brachyphylla cavernarum. In both data sets, values that distinguish Erophylla from Phyllonycteris are in the general range of values that characterize congeneric species of mammals. Immunological distance values for the species Glossophaga soricina, Monophyllus redmani, Anoura caudifer, Leptonycteris sanborni, Choeroniscus minor, and Hylonycteris underuoodi indicate that these taxa are approximately equidistant from the Brachyphyllinae. Immunological comparisons of Glossophaga and Monophyllus to Anoura, Leptonycteris, Choeroniscus, …


Social, Spacing, And Cooperative Behavior Of The Collared Peccary, Tayassu Tajacu, John A. Byers, Marc Bekoff Nov 1981

Social, Spacing, And Cooperative Behavior Of The Collared Peccary, Tayassu Tajacu, John A. Byers, Marc Bekoff

Ethology Collection

Social behavior of the collared peccary was studied on the lower, eastern slopes of the Mazatzal Mountains, Arizona. The social unit in this species is a cohesive herd, in which small inter-individual distances are maintained. Two conspicuous acts, one olfactory and one auditory, functioned to maintain close spacing. Social interactions were brief but tended to synchronize the activities of animals and also to bring them closer together. Amicable and neutral actions occurred far more frequently than agonistic interactions. Most agonistic behavior did not involve physical contact. Cooperative nursing, predator defense, and feeding occurred; all adults were tolerant of young, and …


Mice Of The Genus Peromyscus In Guadalupe Mountains National Park, Texas, John E. Cornely, David J. Schmidly, Hugh H. Genoways, Robert J. Baker Oct 1981

Mice Of The Genus Peromyscus In Guadalupe Mountains National Park, Texas, John E. Cornely, David J. Schmidly, Hugh H. Genoways, Robert J. Baker

University of Nebraska State Museum: Mammalogy Papers

Mice of the genus Peromyscus are found in virtually every habitat type in Guadalupe Mountains National Park in West Texas. Because of their abundance and wide distribution, they comprise an important component of the park's ecosystem. The first known specimens of Peromyscus from the area now included in the park were collected by Vernon Bailey in 1901 (Bailey, 1905). He collected specimens of Peromyscus boyIii in Dog and McKittrick canyons. Davis (1940) collected P. leucopus at Frijole in 1938 and P. boylii in The Bowl in 1938 and 1939. Davis and Robertson (1944) reported collecting P. pectoralis from along Bell …


Activity Of Moose And White-Tailed Deer At Mineral Springs, D. Fraser, H. Hristienko Oct 1981

Activity Of Moose And White-Tailed Deer At Mineral Springs, D. Fraser, H. Hristienko

Social Behavior Collection

Activity of moose (Alces alces) and white-tailed deer (Odocoileus virginianus) was studied at two natural mineral springs (licks) in Ontario during the springtime and summer of 1977 to 1980. Most adult moose appeared to use a lick during a single 1- to 5-day period in a year; some returned at intervals during the season; and all seemed drawn principally by the mineral-rich water. Some young moose remained in the lick vicinity for 3 weeks or more, and often wandered in and out of the licks, grazing herbage as well as drinking. This suggested a social as well as a nutritional …


Dynamics, Movements, And Feeding Ecology Of A Newly Protected Wolf Population In Northwestern Minnesota, Steven H. Fritts, L. David Mech Oct 1981

Dynamics, Movements, And Feeding Ecology Of A Newly Protected Wolf Population In Northwestern Minnesota, Steven H. Fritts, L. David Mech

USGS Northern Prairie Wildlife Research Center

The gray wolf Canis lupus occupies only about 1 percent of its former range in the lower 48 states (Mech 1974a). Most of the range is in northern Minnesota, where the resident population is classified as "threatened" by the U.S. Department of the Interior. Wolves have been and will continue to be the subject of considerable controversy in Minnesota.

The first scientific study of wolves in Minnesota was conducted by Olson (1938a,b). That and all subsequent re- search was in the Superior National Forest (SNF) of northeastern Minnesota even though wolves inhabit approximately the northern third of the state. Consequently, …


The Ecology Of Maine's Intertidal Habitats : A Report Prepared For The Maine State Planning Office, Peter F. Larsen, Lee F. Doggett Oct 1981

The Ecology Of Maine's Intertidal Habitats : A Report Prepared For The Maine State Planning Office, Peter F. Larsen, Lee F. Doggett

Maine Collection

The Ecology of Maine's Intertidal Habitats : A Report Prepared for the Maine State Planning Office

by Bigelow Laboratory for Ocean Sciences (Authors: Peter F. Larsen and Lee F. Doggett) , 1981.

Contents: Acknowledgements / List of Tables / List of Figures / Chapter 1 - Introduction / Chapter 2 - Human Impact on Intertidal Fauna / Chapter 3 - A Primer of Intertidal Ecology / Chapter 4 - A Sampler of Intertidal Species / Chapter 5 - Methodology / Chapter 6 - Results of Faunal Studies / Appendix 1 - Habitats With Date Sampled / Appendix 2 - List …


The Prairie Naturalist Volume 13, Numbers 3 And 4. September - December 1981 Oct 1981

The Prairie Naturalist Volume 13, Numbers 3 And 4. September - December 1981

The Prairie Naturalist

PROVISIONAL CHECKLIST OF MAMMALS OF SOUTH DAKOTA ▪ J. R. Choate and J. K. Jones, Jr.

LABORATORY FEEDING PREFERENCES OF THREE SMALL MAMMALS FOR FIVE TREE SPECIES ▪ T. G. Barnes and T. A. Schaid

NESTLING DIETS OF RED-WINGED AND YELLOW-HEADED BLACKBIRDS ON PLAYA LAKES OF WEST TEXAS ▪ D. H. Fischer and E. C. Bolen

COMPARISON OF FOUR NORTH DAKOTA IMPOUNDMENTS AND FACTORS AFFECTING

THE DEVELOPMENT OF IMPOUNDMENT PARASITOFAUNA ▪ H. L. Holloway, Jr. and N. T. Hagstrom

FIRST STATE RECORD OF RICHARDSON'S GROUND SQUIRREL IN IOWA ▪ R. P. Lampe, J. B. Bowles, and R. Spengler

THE EFFECT …


The Effects Of Copper On Skeletonema Costatum (Greville) Cleve, Susanne Sybille Jackman Oct 1981

The Effects Of Copper On Skeletonema Costatum (Greville) Cleve, Susanne Sybille Jackman

Biological Sciences Theses & Dissertations

Copper levels and dominant phytoplankton species composition were examined from six sites in Kings Creek on Virginia's Eastern Shore from March to August 1980. Creek copper levels were determined by an APDCMIBK extraction procedure and ranged from 0.003 to 0.020 mg/1. Diatoms were the dominant phytoplankters with Skeletonema costatum (Greville) Cleve emerging as a major late summer species.

Subsequent laboratory studies involved 96-hour experiments using Skeletonema costatum (clone Skel) grown in enriched natural seawater. The water was collected from Kings Creek and enriched to Guillard and Ryther's (1962) f/2 media levels. Cupric sulfate was added to the experimental culture media …


Estimation Of Harvest Rate And Vulnerability From Age And Sex Data, J. E. Pahoheimo, David Fraser Oct 1981

Estimation Of Harvest Rate And Vulnerability From Age And Sex Data, J. E. Pahoheimo, David Fraser

Wildlife Population Management Collection

A nonlinear least-squares procedure is developed to estimate harvest rate and differential vulnerability in wildlife populations that are harvested with 1 segment of the population more able than another. The method requires age data by category from a series of harvests, plus information on accumulated harvest effort, and assumes that the nonharvest mortality rate is the same for the categories. The data need not be from consecutive harvests, as long as the effort is known for missing Monte Carlo simulations were done to verify the estimation procedure. An example shows the application of the method to moose (Alces alces) harvest …


Radial-Maze Performance In The Rat Following Lesions Of Posterior Neocortex, Melvyn A. Goodale, Robert H.I. Dale Sep 1981

Radial-Maze Performance In The Rat Following Lesions Of Posterior Neocortex, Melvyn A. Goodale, Robert H.I. Dale

Scholarship and Professional Work - LAS

The present experiment was designed to investigate the role of posterior neocortex (areas 17, 18 and 18a) in the maintenance of performance on the radial maze. Following training to criterion on the 8-arm radial maze, rats received either sham operations, bilateral eye enucleations, lesions of posterior neocortex, or combined enucleations and lesions of posterior neocortex. While the enucleated animals with intact brains showed a slight, but significant performance decrement relative to the sham-operated group, the other two groups, with lesions of areas 17, 18 and 18a, each showed a massive deficit. This large deficit was observed even in the group …


Remembrance Of Places Lasts: Proactive Inhibition And Patterns Of Choice In Rat Spatial Memory, William A. Roberts, Robert H.I. Dale Aug 1981

Remembrance Of Places Lasts: Proactive Inhibition And Patterns Of Choice In Rat Spatial Memory, William A. Roberts, Robert H.I. Dale

Scholarship and Professional Work - LAS

A series of experiments was carried out to evaluate the notion that rats given a sequence of massed daily trials on the radial maze reset working memory at the end of each trial by deleting its contents. Although curves presented by D. S. Olton [Scientific American, 1977, 236, 82-98: In S. H. Hulse, H. Fowler, & W. K. Honig (Eds.), Cognitive processes in animal behavior, 1978, Hillsdale, N.J.: Erlbaum] show that rats return to errorless performance at the beginning of each trial after the first, the fact that accuracy falls less rapidly over choices on Trial 1 …


The Effect Of Storm Sewer Drainage On An Urban Lake, Jose Ignacio Aizpurua Aug 1981

The Effect Of Storm Sewer Drainage On An Urban Lake, Jose Ignacio Aizpurua

Masters Theses

The present study was undertaken to evaluate the impact from normal precipitation (ra in , snow) into an urbanized lake during 1978- 1980. Several stations were chosen, and various limnological parameters were investigated. Chemical analyses were used to estimate the annual load of nutrients input into the lake by the storm sewer and surface runoff. Excessive phosphorus and nitrogen loads from these sources were discharged into the lake.

Fecal coliform distribution in the lake before, during, and after several storm events was studied. In general, fecal coliform counts increased after storm events. Peak concentrations decreased markedly with distance of the …


Behavioural Budgeting By Wild Coyotes: The Influence Of Food Resources And Social Organization, Marc Bekoff, Michael C. Wells Aug 1981

Behavioural Budgeting By Wild Coyotes: The Influence Of Food Resources And Social Organization, Marc Bekoff, Michael C. Wells

Ethology Collection

Daytime behavioural budgets of coyotes (Canis latrans) living in the Grand Teton National Park Jackson, Wyoming, were analysed in order to determine how activity patterns ' ere influenced by food resources and social organization. In winter coyotes rested more-and hunted less than in other seasons. Pack-living coyotes rested more and travelled less than resident pairs or solitary resident or transients during winter months when the major food resource was ungulate (predominantly elk, Cervus canadensis) carrion. A mated female living in a pack rested significantly more and travelled significantly less than a mated female living only with her mate (as a …


Results Of The Alcoa Foundation-Suriname Expeditions. V. Noteworthy Records Of Surinamese Mammals, Hugh H. Genoways, Stephen L. Williams, Jane A. Groen Jul 1981

Results Of The Alcoa Foundation-Suriname Expeditions. V. Noteworthy Records Of Surinamese Mammals, Hugh H. Genoways, Stephen L. Williams, Jane A. Groen

University of Nebraska State Museum: Mammalogy Papers

The occurrence of seven species of mammals previously unknown in Suriname is documented. The new taxa recorded include Didelphis albiventris, Peropteryx macrotis, Lonchorhina aurita, Micronycteris hirsuta, Vampyrodes caraccioli, Furipterus horrens, and Thyroptera discifera . Additional information is provided on several species already known to occur in Suriname, including Metachirus nudicaudatus, Peronymus leucopterus, Mimon bennettii, Tonatia carrikeri, T. schulzi, Anoura geoffroyi, Choerniscus intermedius, Mesophylla macconnelli, Neaeomys guianae, Holochilus brasiliensis, and Potos fiavus.


Results Of The Alcoa Foundation-Suriname Expeditions. Vi. Additional Chromosomal Data For Bats (Mammalia: Chiroptera) From Suriname, Robert J. Baker, Hugh H. Genoways, Paisley A. Seyfarth Jul 1981

Results Of The Alcoa Foundation-Suriname Expeditions. Vi. Additional Chromosomal Data For Bats (Mammalia: Chiroptera) From Suriname, Robert J. Baker, Hugh H. Genoways, Paisley A. Seyfarth

University of Nebraska State Museum: Mammalogy Papers

As part of ongoing studies of the bats of Suriname, karyotypic information is presented for seventeen species. Chromosomal data are presented for the first time for Peronymus leucopterus, Peropteryx macrotis, Mimon bennettii, Artibeus concolor, Furipterus horrens, and Thyroptera discifera. Additional chromosomal data are presented for eleven other species of bats for which some information was available previously.


Deer Social Organization And Wolf Predation In Northeastern Minnesota, Michael E. Nelson, L. David Mech Jul 1981

Deer Social Organization And Wolf Predation In Northeastern Minnesota, Michael E. Nelson, L. David Mech

USGS Northern Prairie Wildlife Research Center

The white-tailed deer Odocoileus virginianus has been subject to intensive research and management, yet we are just beginning to understand its social organization. Little is known about home range formation, migration, social bonds, and traditions in this deer, what functions they serve, and what selective forces have affected them.

Predation by wolves Canis lupus, in particular, has not been examined as a factor in deer evolution, yet the intimate interactions between deer and wolf through the millennia no doubt strongly influenced major morphological and behavioral adaptations in both species. It is a reasonable assumption that wolf predation has been …


Diagenesis Of Organic Matter In Las Vegas Bay And Bonelli Bay, Lake Mead, James W. Murray, Carolyn J. Jones, Kathy Kuivila, Jeff Sawlan Jul 1981

Diagenesis Of Organic Matter In Las Vegas Bay And Bonelli Bay, Lake Mead, James W. Murray, Carolyn J. Jones, Kathy Kuivila, Jeff Sawlan

Publications (WR)

The interstitial water chemistry of the sediments of Las Vegas Bay and Bonelli Bay in Lake Mead has been studied as part of a comprehensive water quality study of those locations. Pore water and solid phase analyses were completed from four stations in Las Vegas Bay and two stations in Bonelli Bay. At both locations the pore water compositions and organic matter diagenesis in the sediments are dominated by sulfate reduction. This major role of sulfate reduction is unusual for lake sediments and reflects the fact that SO4 is the major anion in the lake water. In addition, gypsum …


The Seasonality Of Occurrence Of Larval And Juvenile Sciaenids In A Virginia Seaside Estuary, James Howard Cowan Jr. Jul 1981

The Seasonality Of Occurrence Of Larval And Juvenile Sciaenids In A Virginia Seaside Estuary, James Howard Cowan Jr.

Biological Sciences Theses & Dissertations

The seasonality of occurrence and relative abundance of larval and juvenile fishes, particularly members of the family Sciaenidae, from a Virginia seaside estuary were determined from ichthyoplankton and trawl collections made from March 1979 to March 1980.

The larvae of 20 species distributed among 15 families were identified from the ichthyoplankton. Larvae of the bay anchovy, Anchoa mitchilli, and the Atlantic silverside, Menidia menidia, dominated the samples and made up 57 and 34%, respectively, of all larvae collected. Peak occurrence began in May and continued through August. Fish larvae were present in the study area all year.

Twenty-eight …


The Prairie Naturalist Volume 13, Number 2. June 1981 Jun 1981

The Prairie Naturalist Volume 13, Number 2. June 1981

The Prairie Naturalist

HERPETOFAUNA OF MORMON ISLAND PRESERVE, HALL COUNTY, NEBRASKA ▪ S. M. Jones, R. E. Ballinger and J. W Nietfeldt

COYOTE USE OF PLAYAS IN THE TEXAS HIGH PLAINS ▪ R. W. Whiteside and F. S. Guthery

NEW RECORD OF THE LEAST WEASEL IN WYOMING ▪ M. R. Stromberg, D. E. Biggens and M. Bidwell

STATUS OF AMERICAN WOODCOCK IN NEBRASKA WITH NOTES ON A RECENT BREEDING RECORD ▪ G. R. Lingle

WINTER FOODS OF THE PINE GROSBEAK IN EASTERN NORTH DAKOTA ▪ R. C. Reigh

SEASONAL VARIATION IN HOME RANGE OF A FEMALE BADGER (Taxidea taxus) ▪ R. …


Rare Vascular Plants Of Maine : A Critical Areas Program Report, Susan C. Gawler, L. M. Eastman Jun 1981

Rare Vascular Plants Of Maine : A Critical Areas Program Report, Susan C. Gawler, L. M. Eastman

Maine Collection

Rare Vascular Plants of Maine : A Critical Areas Program Report

Based on Rare Vascular Maine Plants (1978) with Additional Herbarium Data and Rare Plants Information by L.M. Eastman.

Introduction, Criteria, Annotated Rare Plant List, Species Accounts, Additional Herbarium Data, County & Town Index Editing by Susan C. Gawler.

Executive Department, State Planning Office, 189 State Street, Augusta, Maine, 04333 (June 1981).

Contents: Acknowledgements / List of Tables / List of Figures / Part One: Introduction / Part Two: List of the Rare Vascular Plants of Maine / Part Three: Species Accounts / Part Four: Subspecific Taxa / Appendices / …


Vegetation Patterns A Half Century Following The Chestnut Blight In The Great Smoky Mountains National Park, Ernesto Arends Jun 1981

Vegetation Patterns A Half Century Following The Chestnut Blight In The Great Smoky Mountains National Park, Ernesto Arends

Masters Theses

Vegetation patterns approximately 50 years after the chestnut blight were examined in forest stands of the Great Smoky Mountains National Park. Specific objectives of this study were: (1) to determine the present composition of replacement stands following the elimination of chestnut, (2) to examine selected site factors and to relate them to tree species composition, (3) to project future trends in the composition of the replacement stands, and (4) to compare these results with earlier results and predictions of Wood and Shanks (1959) concerning chestnut replacement in the same area.

Data were analyzed from 38 sample plots ranging from 2000 …


Middlesex County Tidal Marsh Inventory, Walter L. Priest Iii, Gene M. Silberhorn Jun 1981

Middlesex County Tidal Marsh Inventory, Walter L. Priest Iii, Gene M. Silberhorn

Reports

No abstract provided.


An Observational Study Of Coyote (Canis Latrans) Scent-Marking And Territoriality In Yellowstone National Park, Joseph J. Allen, Marc Bekoff, Robert L. Crabtree May 1981

An Observational Study Of Coyote (Canis Latrans) Scent-Marking And Territoriality In Yellowstone National Park, Joseph J. Allen, Marc Bekoff, Robert L. Crabtree

Ethology Collection

Free-ranging coyotes (Canis latrans) living in neighboring packs were observed in the Lamar Valley of Yellowstone National Park, Wyoming, from Jan. to May 1997. Through direct observation, we recorded the location of coyote scent marks and information regarding the identity of the marking animal. Patterns of scent-marking were then analyzed spatially and demographically. All of the evidence from the present study supports a strong relationship between scent-marking and territoriality in these coyotes, and all predictions were met. A preponderance of scent marks was found in the periphery of territories. Most of those marks were raised-leg urinations (RLUs) and forward-lean urinations …