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Could The European Thanasimus Formicarius Outcompete The North American Thanasimus Dubius (Coleoptera: Cleridae) If Introduced To North America?, Robert A. Haack, Robert K. Lawrence

The Great Lakes Entomologist

A federal quarantine was enacted in the United States (US) in the fall of 1992 in response to discovery of the Eurasian pine shoot beetle, Tomicus piniperda (L.) (Coleoptera: Scolytinae), in July 1992 in Ohio. The quarantine regulated movement of pine products (e.g., Christmas trees, nursery stock, and logs) at the county level from infested to uninfested US states. A classical biological program was considered for T. piniperda starting in 1995, focusing on the Eurasian clerid beetle Thanasimus formicarius (L.), an important predator of T. piniperda. Concerns were raised in 1996 that if T. formicarius were released in the …

Initial Changes In Ground Beetle (Carabidae: Coleoptera) Assemblages After Creating Gaps Of Varying Sizes In Mature Northern Hardwoods Forests In Wisconsin, Robert A. Haack, Robert E. Acciavatti, Toby R. Petrice, Robert L. Davidson, Terry F. Strong

The Great Lakes Entomologist

Managers of northern hardwood forests in the Great Lakes region create gaps in the forest canopy of various sizes during harvesting to achieve different silvicultural objectives. Single-tree harvesting favors recruitment of shade-tolerant tree species by creating small gaps in the canopy, whereas group selection creates larger gaps that can favor shade-intolerant tree species. We evaluated changes in carabid beetle abundance and diversity in harvest-created gaps of five sizes (gap diameters of 6, 10, 20, 30, and 46 m) and uncut forests in northern Wisconsin where sugar maple (Acer saccharum) was the dominant tree species. Gaps were cut during …

The Distribution Of Some Soil Arthropods In A Manipulated Ecosystem, James H. Shaddy, James W. Butcher

The Great Lakes Entomologist

[Excerpt] Soil cultivation practices and agricultural utilization affect soil animal populations in various ways. Gisin (1955) and Schalk (1968) concluded that the effect was minimal. Others observed a deleterious effect (Karg, 1956; Sheals, 1956; Strebel, 1957; Aleinikova and Utrobina, 1975; Edwards and Lofty, 1975). However, some soil animals appear to be benefited (Aleinikova and Utrobina, 1975; Edwards and Lofty, 1975). The effects of cultivation and many other factors have been discussed thoroughly in reviews by Christiansen (1964), Burges and Raw (1967), Butcher, et al. (1971), Ghilarov (1975), and Wallwork (1976).

Invertebrate Predators Of Michigan Lepidoptera, M. C. Nielson

The Great Lakes Entomologist

[Excerpt]

For many years this author has been actively collecting Michigan Lepidoptera and generally observing insects in the field. Almost every season some predation of Lepidoptera by insects and their allies is seen. Unfortunately, many early observations went unrecorded in my field notebooks, but since the late 1950's I have made notations of these interesting encounters.

The Distribution Of The Pentatomoidea In The Northeastern Quarter Of The United States (Hemiptera), J. E. Mcpherson

The Great Lakes Entomologist

The distributions of the pentatomoids occurring in the northeastern quarter of the United States as listed by Van Duzee in 1917 are updated.

Diversity And Activity Of Ground-Dwelling Spiders (Araneae) In Four Sub-Communities In A Degraded Oak Woodland At The Chicago Botanic Garden, Cook County, Illinois, James F. Steffen, Michael L. Draney

The Great Lakes Entomologist

Ground-dwelling spiders were sampled using pitfall traps over a twelve-month period from four sub-communities within a 40-ha oak woodland complex in northern Cook County, Illinois. Mesic and dry mesic oak woodland, mesic upland forest, buckthorn dominated mesic oak savanna and a mesic woodland control, where no restoration management is planned, were sampled. Fifty-five species within 44 genera and 18 families were trapped. The results suggest negative correlations between non-native spider abundance and spider species richness. Also, phenology of the various species indicates that a year-round inventory is necessary to adequately sample ground-dwelling spider assemblages. Possibilities of using Bathyphantes concolor (Wider) …

Simulation Of The Effects Of Stand Factors On Spruce Budworm (Lepidoptera: Tortricidae), W. P. Kemp, J. P. Nyrop, G. A. Simmons

The Great Lakes Entomologist

A model was developed to simulate the life history of instar-I through instar-I1 spruce budworm, Choristoneura fumiferana (Clemens). Particular emphasis was given to the redistribution phases during spring and fall and the factors affecting budworm survival. Analyses included the effects of aggregated stand types, temperature, cloud cover, open areas, and the species distribution, on redistribution mortality. Of those stand factors examined, open area and non-host percentages were the most important in regulating redistribution survival.

A Checklist Of Illinois Centipedes (Chilopoda), Gerald Summers, J. A. Beatty, Nanette Magnuson

The Great Lakes Entomologist

Records for 45 centipede species occurring in Illinois are given with a brief discussion of general habitat features and the effects of certain environmental factors on specific distribution patterns. The chilopod fauna of Illinois includes species from southeastern, western, and northern physiographic regions and represents a richer mixture of chilopod species than is found in any adjacent state.

Instar Head Widths, Individual Biomass, And Development Rate Of Forest Tent Caterpillar, Malacosoma Disstria (Lepidoptera: Lasiocampidae), At Two Densities In The Laboratory, Jocelyn M. Muggli, William E. Miller

The Great Lakes Entomologist

Two colonies of forest tent caterpillar were reared at different densities on trembling aspen foliage in an environment chamber programmed for natural temperatures. Based on direct observation of molting, both colonies underwent five instars which overlapped negligibly in head width. Instar head widths were relatively stable between colonies, but individual biomass was sensitive to the different densities. Instars 3-5 in the less dense colony were heavier than those of the more dense by factors of 1.3-1.9. Development to 10-20% of larvae spinning cocoons took 40 days in the less dense colony and 45 days in the more dense. Size of …

A Five-Year Study Of The Ground-Dwelling Beetles (Coleoptera) From A Grassland And An Adjacent Woods In Southern Québec (Canada), Claire Levesque, Gilles-Yvon Levesque

The Great Lakes Entomologist

We present results of the first long-term study on spatiotemporal variations in the composition and structure of ground-dwelling beetle assemblages from a grassland and adjacent woods in eastern Canada. It is the third part of an investigation on the beetle biodiversity using diverse methods between 2006 and 2012. During the entire snow-free season (April/May to October) in 2006–2010, we collected by pitfall trapping 67694 individuals representing 386 Coleoptera species and morphospecies belonging to 35 families in southern Québec. Adventive species represented at least 51.9% of individuals. Carabidae and Staphylinidae were the most abundant families and had the highest species richness …

Changes In Ground Beetle (Coleoptera: Carabidae) Species Assemblages After Removal Of Red Pine (Pinus Resinosa) Overstory On Dry Sand Prairie Sites In Michigan, Robert A. Haack, Toby Petrice, Robert L. Davidson, Robert E. Acciavatti

The Great Lakes Entomologist

Ground beetle (Coleoptera: Carabidae) species assemblages were studied at sites on the Huron-Manistee National Forests in Newaygo County, Michigan. The goal was to monitor changes in carabid diversity on areas where mature red pine (Pinus resinosa Sol. ex Aiton) was removed during the 1996-1997 winter and compare with carabid diversity at nearby natural dry sand prairie sites, natural oak-pine barrens, and uncut red pine stands. The red pine had been planted on dry sand prairie sites during 1940-1950. Sampling with pitfall traps occurred during May-August in both 1996 and 1997, but only July-August in 1999. Overall, 16,259 carabid specimens …

Coleoptera Collected Using Three Trapping Methods At Grass River Natural Area, Antrim County, Michigan, Robert A. Haack, Bill Ruesink

The Great Lakes Entomologist

Abstract. Overall, 409 Coleoptera species (369 identified to species, 24 to genus only, and 16 to subfamily only), representing 275 genera and 58 beetle families, were collected from late May through late September 2017 at the Grass River Natural Area (GRNA), Antrim County, Michigan, using baited multi-funnel traps (210 species), pitfall traps (104 species), and sweep nets (168 species). All three collecting methods were used in three distinct habitats: a rich conifer swamp (cedar), near the edge of a red pine plantation (pine), and within a mesic northern hardwood forest (hardwoods). Additional collections were made along two trails and in …

Occurrence Of Ticks (Acari: Ixodidae) On Birds In Northwestern Lower Michigan, 2011-2019, William C. Scharf, Lisa Aukland, Gary W. Shugart, Sarah A. Hamer

The Great Lakes Entomologist

Abstract

Monitoring tick infestation of wildlife provides baseline tick

occurrence data that may have human or animal health implications. We collected 312 ticks of four species from 5,122 birds of 93 species while monitoring bird migration during 15 fall and spring seasons between 2011 and 2019 in the northern Lower Peninsula of Michigan. Twenty-seven of 93 bird species hosted ticks with an overall prevalence (=at least one tick) of 3.6% (185/5,122). Median burden was one tick/per infested bird with a range of 1-16 ticks per infested bird. Tick species collected were primarily Haemaphysalis leporispalustris (Packard) and Ixodes scapularis Say, with …

Reinstatement Of Andrena Vernalis Mitchell (Hymenoptera: Andrenidae) From Synonymy With A. Ziziae Robertson, Zachary M. Portman, Ian G. Lane, Gabriella L. Pardee, Daniel P. Cariveau

The Great Lakes Entomologist

Andrena (Micrandrena) ziziae Robertson, 1891 (Andrenidae) is a well-known species found in a variety of habitats in the eastern and central United States and adjacent southern Canada. Andrena (Micrandrena) vernalis Mitchell, 1960 was described from five female specimens in the eastern United States and was synonymized with A. ziziae by Ribble in 1968. Recently collected specimens from throughout Minnesota have revealed that A. ziziae sensu Ribble is actually two species, one of which matches A. vernalis. Here, we reinstate A. vernalis as a valid species and describe the previously unknown male. We provide diagnostic characters …

Epizooic Diatoms On The Cerci Of Ephemeroptera (Caenidae) Naiads, Daniel E. Wujek

The Great Lakes Entomologist

Using scanning electron microscopy, epizooic diatoms were observed growing on the cerci of Caenis amica Hagen naiads (Ephemeroptera, Caenidae). Meridion circulare (Greville) C. Agardh was the most abundant, followed by Synedra rumpens Kützing, then Cocconeis pediculus Ehrenberg. Other diatom species observed from substrates in Cedar Creek, Isabella County, Michigan were not observed on the cerci. No diatoms were observed on Ephemerellidae naiads.

Pentatomoidea (Hemiptera: Pentatomidae, Scutelleridae) Associated With The Dioecious Shrub Florida Rosemary, Ceratiola Ericoides (Ericaceae), A. G. Wheeler Jr.

The Great Lakes Entomologist

Florida rosemary (Ceratiola ericoides), an ericaceous shrub with needlelike leaves, is characteristic of Florida scrubs and certain other xeric plant communities of well-drained sands. The plant is dioecious, its inconspicuous male and female flowers borne on separate individuals. From 2003 to 2012 (mainly 2007–2012), rosemary was sampled periodically (with all months included at least once during the period) in sand ridges of peninsular Florida and two riverine dunes in southeastern Georgia. Branches of male and female plants were tapped separately into a beating net. Sampling during the final three years was female-plant-biased to facilitate work on fruit- and seed-feeding …

Activity And Diversity Of Collembola (Insecta) And Mites (Acari) In Litter Of A Degraded Midwestern Oak Woodland, James F. Steffen, Joan Palincsar, Florrie M. Funk, Daniel J. Larkin

The Great Lakes Entomologist

Litter-inhabiting Collembola and mites were sampled using pitfall traps over a twelve-month period from four sub-communities within a 100-acre (40-ha) oak-woodland complex in northern Cook County, Illinois. Sampled locations included four areas where future ecological restoration was planned (mesic woodland, dry-mesic woodland, mesic upland forest, and buckthorn-dominated savanna) and a mesic woodland control that would not be restored. Fifty-eight mite and 30 Collembola taxa were identified out of 5,308 and 190,402 individuals trapped, respectively. There was a significant positive relationship between litter mass and both mite diversity and the ratio of Oribatida to Prostigmata and a significant negative relationship between …

A New Species Of Aphis In Minnesota (Hemiptera: Aphididae) On Narrow-Leaved Purple Coneflower, Echinacea Angustifolia, D. M. Lagos, D. J. Voegtlin

The Great Lakes Entomologist

Aphis echinaceae, n. sp. is described from specimens collected in Douglas County, Minnesota, on Echinacea angustifolia (DC.). The aphid is illustrated with biometric data for alate and apterous vivipara. Its short cauda and siphunculi, and presence of secondary sensoria on antennal segments III, IV, and V in both morphs makes this Aphis morphologically similar to A. debilicornis (Gillette and Palmer), A. lugentis Williams and A. caliginosa Hottes and Frison, and root feeding aphids of the subgenus Pseudoprotaphis Kadyrbekov and genus Protaphis Börner. A diagnosis is provided.

Effects Of Pitfall Trap Preservative On Collections Of Carabid Beetles (Coleoptera: Carabidae), Kenneth W. Mccravy, Jason E. Willand

The Great Lakes Entomologist

Effects of six pitfall trap preservatives (5% acetic acid solution, distilled water, 70% ethanol, 50% ethylene glycol solution, 50% propylene glycol solution, and 10% saline solution) on collections of carabid beetles (Coleoptera: Carabidae) were studied in a west-central Illinois deciduous forest from May to October 2005. A total of 819 carabids, representing 33 species and 19 genera, were collected. Saline produced significantly fewer captures than did acetic acid, ethanol, ethylene glycol, and propylene glycol, while distilled water produced significantly fewer captures than did acetic acid. Significant associations between numbers of captures and treatment were seen in four species: Amphasia interstitialis …

Why Are There So Few Insect Predators Of Nuts Of American Beech (Fagus Grandifolia)?, Charles E. Williams

The Great Lakes Entomologist

American beech, Fagus grandifolia Ehrh., is a common nut-bearing tree of eastern North America. Compared to other North American nut-bearing tree species of comparable geographic range, the nut-infesting insect fauna of American beech is species-poor: only the filbertworn, Cydia latiferreana (Wlsm.) (Lepidoptera: Tortricidae), infests nuts of American beech. Why are there so few insect predators of nuts of American beech? Using data from published studies, I explore two hypotheses that may help to explain the species-poor nut-infesting insect fauna of American beech. First, might chemical defense of beechnuts, and/ or low nutritional value, restrict the number of insect predators that …

A Friend Unmasked: Notes On Chauliognathus Pensylvanicus (Coleoptera: Cantharidae) And The Nature Of Natural History, Andrew H. Williams

The Great Lakes Entomologist

Data from field observations from 1992 through 2006 and from museum specimens of adult Chauliognathus pensylvanicus (DeGeer) from Wisconsin are presented here, with information from the literature. Beetles were found using flowers of 183 species in 29 families growing outside cultivation in Wisconsin and in every Wisconsin county. All evidence supports the characterization of C. pensylvanicus as a prairie insect that also lives in other, early successional plant communities.

Tomicus Piniperda (Coleoptera: Scolytidae) Reproduction And Development In Scots, Jack, Red And Eastern White Pine Under Laboratory Conditions, Hui Ye, Robert A. Haack, Toby R. Petrice

The Great Lakes Entomologist

The pine shoot beetle, Tomicus piniperda (L.) (Coleoptera: Scolytidae), is an exotic bark beetle in North America that was first found in the Great Lakes region in 1992. We evaluated T. piniperda reproduction and development in one Eurasian pine (Scots pine, Pinus sylvestris L.) and three North American pines (jack pine, P. banksiana Lamb.; red pine, P. resinosa Ait.; and eastern white pine, P. strobus L.) under laboratory conditions. We introduced one pair of adults into individual pine bolts, allowed development, collected brood, and later debarked all bolts and measured galleries. Reproduction and development occurred in all pine species tested. …

Ecological Notes On Species Of Cleridae (Insecta: Coleoptera) Associated With The Prairie Flora Of Central North America, Jonathan R. Mawdsley

The Great Lakes Entomologist

The association of eighteen species of Cleridae (Coleoptera) with tallgrass and shortgrass prairie ecosystems in central North America is reported for the first time. New flower visitation, habitat association, distribution, and/or rearing records are reported for Enoclerus analis (LeConte), E. coccineus coccineus (Schenkling), E. cordifer (LeConte), E. rosmarus (Say), E. zonatus (Klug), Placopterus thoracicus pallipes (Wolcott), Trichodes bibalteatus LeConte, T. bicinctus Green, T. nutalli (Kirby), Phyllobaenus dubius (Wolcott), P. knausii (Wickham), P. pubescens (LeConte), P. subfasciatus (LeConte), Isohydnocera albocincta (Horn), I. brunnea (Horn), I. curtipennis (Newman), I. tricondylae (LeConte) and Wolcottia pedalis (LeConte). Diagnostic characters are presented to separate adults …

New Records Of Rhopalosomatidae (Hymenoptera: Vespoidea) From Wisconsin, Steven J. Krauth

The Great Lakes Entomologist

The rhopalosomatid Olixon banksii is recorded from Wisconsin for the first time.

Richness And Abundance Of Carabidae And Staphylinidae (Coleoptera), In Northeastern Dairy Pastures Under Intensive Grazing, R. A. Byers, G. M. Barker, R. L. Davidson, E. R. Hoebeke, M A. Sanderson

The Great Lakes Entomologist

Dairy cattle has become popular to dairy farmers in the Northeast looking for management schemes to cut production costs. Carabidae (ground beetles) and Staphylinidae (rove beetles) are indicators of habitat disturbances, such as drainage of wetlands, or grassland for grazing animals, and their monitoring could provide one measure of ecosystem sustainability if intensive management systems expand or intensify in the future. Our objective was assess the abundance and species richness of these two beetle families under intensive grazing throughout Pennsylvania, southern New York and Vermont. We collected 4365 ground beetles (83 species) and 4,027 rove beetles (79 species) by pitfall …

Dispersal And Re-Capture Of Marked, Overwintering Tomicus Piniperda (Coleoptera: Scolytidae) From Scotch Pine Bolts, A. V. Barak, D. Mcgrevy, G. Tokaya

The Great Lakes Entomologist

The pine shoot beetle (PSB), Tomicus piniperda is a recently established exotic pest of live pine in the southern Great Lakes of the U.S. and Canada. Scotch pine, Pinus sylvestris L. is the most susceptible pine species, but the adult also attacks several other North American species of Pinus. This research investigated the dispersal behavior of beetles emerging from overwintering sites to aid in the development of effective monitoring and management practices. Scotch pine logs with overwintering PSB were sprayed with fluorescent pigments to mark dispersing beetles. These logs were placed in piles in the centers of three circular …

Seasonal Shoot-Feeding By Tomicus Piniperda (Coleoptera: Scolytidae) In Michigan, Robert A. Haack, Robert K. Lawrence, George C. Heaton

The Great Lakes Entomologist

Seasonal shoot-feeding by Tomicus piniperda (L.) was monitored at 2­ week intervals on 15 Scotch pine, Pinus sylvestris L., trees from 8 April through 16 November 1994 in southern Michigan. All shoots that showed evidence of T. piniperda attack were removed every two weeks. In 1994, initial spring flight of T. piniperda began on 22 March. At least two live T. piniperda adults were found on the 15 trees on each sampling date from 8 April through 1 November 1994. In addition, at least one freshly attacked, beetle- free shoot was found on each sampling date except for 1 November. …

Tomicus Piniperda (Coleoptera: Scolytidae): Is Shoot Feeding Required For Reproductive Maturation?, Therese M. Poland, Robert A. Haack

The Great Lakes Entomologist

The pine shoot beetle, Tomicus piniperda (Coleoptera: Scolytidae), is a univoltine pest of pine in its native range of Europe and Asia. Tomicus piniperda is now widely established in the Great Lakes region and poses a potentially significant threat to other pine-producing areas in North America. An unusual aspect of the life history of T. piniperda is the extended period of maturation-feeding that takes place in the shoots of living pine trees and subsequent overwintering before adults reproduce the following year. We investigated the extent to which shoot-feeding is required by newly-emerged T. piniperda before introduction into Scotch pine (Pinus …

New Canadian Records Of Asilidae (Diptera) From An Endangered Ontario Ecosystem, Jeffrey H. Skevington

The Great Lakes Entomologist

The Asilidae (Diptera) of Bosanquet (northern Lambton County, Ontario) are surveyed. Forty-one species are recorded. Twelve species are published for the first time from Canada: Atomosia puella, Cerotainia albipilosa, Cerotainia macrocera, Holcocephala calva, Holopogon (Holopogon) oriens, Laphria canis, Laphria divisor, Laphria grossa, Lasiopogon opaculus, Machimus notatus, Machimus sadyates, and Neomochtherus auricomus. These species plus the following four are new to Ontario: Laphystia flavipes, Lasiopogon tetragrammus, Machimus novaescotiae, and Proctacanthella ca­copiloga.

Habitat Characterization Of Five Rare Insects In Michigan (Lepidoptera: Hesperiidae, Riodinidae, Satyridae; Homoptera: Cercopidae), Keith S. Summerville, Christopher A. Clampitt

The Great Lakes Entomologist

Over 80 species of insects are listed as endangered, threatened, or special concern under Michigan's endangered species act. For the majority of these species, detailed habitat information is scant or difficult to interpret. We describe the habitat of five insect species that are considered rare in Michigan: Lepyronia angulifera (Cercopidae), Prosapia ignipectus (Cercopidae), Oarisma poweshiek (Hesperiidae), Calephelis mutica (Riodinidae), and Neonympha mitchellii mitchellii (Satyridae). Populations of each species were only found within a fraction of the plant communities deemed suitable based upon previous literature. Furthermore, individuals of each species were observed to be closely affiliated with just a few vegetation …