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Full-Text Articles in Biological and Physical Anthropology

Meta-Analysis Of Scent Detection Canines And Potential Factors Influencing Their Success Rates, Molly Marie Jaskinia Jan 2023

Meta-Analysis Of Scent Detection Canines And Potential Factors Influencing Their Success Rates, Molly Marie Jaskinia

Graduate Student Theses, Dissertations, & Professional Papers

Objective: This is a meta-analysis focused on the success rates of scent detection canines and potential factors that could influence their accuracy. A series of statistical analyses were conducted to determine if certain demographic factors, such as the dog’s gender, age, and breed, have an effect on a scent dog’s accuracy during a search. Or if more circumstantial factors, like the dog’s level of experience in scent work, the type of target scent, and their handler’s awareness of the target’s location, affect the outcome of the search.

Materials and Methods: A dataset was created from 37 different articles consisting of …


Analysis And Observation Of Decomposition Of Immature Pigs In The Minnesota Winter/Early Spring, Rachael Herbes Jan 2023

Analysis And Observation Of Decomposition Of Immature Pigs In The Minnesota Winter/Early Spring, Rachael Herbes

All Graduate Theses, Dissertations, and Other Capstone Projects

This study analyzes the multiple factors affecting the decomposition of pigs in the Minnesota Winter/Early Spring within the first twelve weeks postmortem. The winters in the Minnesota River Valley can vary in regard to weather patterns and temperature changes. Adding the variable of location and accessibility to the study creates a mix of important research conducted in a less populated area. Pig carcasses were used in lieu of human remains to allow the data to be as close to a real scenario as possible. One pig was placed next to a farm site and the other about a half of …


Canine Microwear In Relation To Diet In Sumatran Primates And African Great Apes, Putu Pujiantari Dec 2021

Canine Microwear In Relation To Diet In Sumatran Primates And African Great Apes, Putu Pujiantari

Graduate Theses and Dissertations

This thesis considers canine microwear in relation to diet in five Sumatran primates (Pongo abelii, Hylobates lar, Hylobates agilis, Presbytis thomasi, and Macaca fascicularis) and two African great apes (Pan paniscus and Pan troglodytes schweinfurthii) using both microwear texture analysis and microwear feature analysis techniques. Statistical results for texture analysis show that there are significant differences in scale of maximum and heterogeneity. This indicates that some species have large pits on their canine surfaces, having these dominated by deep features at coarse scale yet have a slight microwear heterogeneity. For feature analysis, all variables show statistically significant variation. Variance in …


An Assessment Of The Neurovascular Structures Of The Trigeminal Nerve And Their Relationship To Diet In Primates, Caitlin Yoakum Dec 2021

An Assessment Of The Neurovascular Structures Of The Trigeminal Nerve And Their Relationship To Diet In Primates, Caitlin Yoakum

Graduate Theses and Dissertations

The inferior alveolar nerve (IAN) enters the mandible via the mental foramen, supplies nervous sensation to the mandibular teeth as it travels through the mandibular canal, and exits the mandibular foramen to send information to the brain to maintain chewing cycles and protect the teeth from damage. Although bony canals and foramina have been shown to form around soft-tissue structures, there are some examples (e.g., the hypoglossal nerve/canal) where the nervous structures do not comprise most of the canal/foramina space. It is important to know the size of nerves because it has been established that larger nerves convey more information …


Attitude Toward Companion And Guard Dogs In Hawaii: Health And Welfare Implications, Lynn Morrison, Julie Ann Luiz Adrian, Marina Kelley, Johana Hill, Zachariah Tman, Dana-Lynn Ko'omoa-Lange Oct 2021

Attitude Toward Companion And Guard Dogs In Hawaii: Health And Welfare Implications, Lynn Morrison, Julie Ann Luiz Adrian, Marina Kelley, Johana Hill, Zachariah Tman, Dana-Lynn Ko'omoa-Lange

People and Animals: The International Journal of Research and Practice

The island of Hawaii exhibits extremes in dog welfare ranging from dogs as family members to dogs used as commodities, either as guard or hunting dogs, with many lacking appropriate care. This study offers a preliminary exploration of people’s attitudes toward companion and noncompanion dogs and the health and welfare implications for humans and dogs. Data collection included interviews and surveys conducted among 20 companion dog/human and eight guard dog/human dyads. Blood pressure (BP) was monitored during interviews to assess stress. The results of the interview data led to five themes. The first two themes describing contexts of dog welfare …


Playing With Relatives: Genetic Kinship And Play Behavior In Juvenile Tibetan Macaques, Riley Derby Jan 2021

Playing With Relatives: Genetic Kinship And Play Behavior In Juvenile Tibetan Macaques, Riley Derby

All Master's Theses

This thesis is composed of a journal-ready article and an accompanying appendix containing data and figures. In this thesis, I statistically analyzed the relationship between player age, player sex, genetic kinship and social play duration in juvenile Tibetan macaques.

Chapter I provides a general introduction and addresses current and past literature on the behavior, ecology and play among macaque genera and Tibetan macaques within this study. I review the fitness benefits of play behavior and the influence of kin selection on macaque social life. Additionally, I review the existing literature on Tibetan macaque play and how the current study will …


Do Actions Speak Louder Than Words? Communicative Frequencies And Multimodality In Ring-Tailed Lemurs (Lemur Catta), Hilary Hager Sep 2020

Do Actions Speak Louder Than Words? Communicative Frequencies And Multimodality In Ring-Tailed Lemurs (Lemur Catta), Hilary Hager

Electronic Thesis and Dissertation Repository

The study of multimodal communication in primatology has increased only recently. At present, there are no on-going investigations of multimodal communication in ring-tailed lemurs (Lemur catta), despite the body of research on this species. I investigated how different modes of L. catta inter-individual multimodal communication are socially coordinated and integrated by examining frequencies of occurrence within four potential biological and social factors: age, troop affiliation, sex, and dominance rank. Research was conducted over four months at the Duke Lemur Center, Durham, NC, on 14 individuals from three separate troops of captive, free-ranging L. catta. Results demonstrate communicative …


The Development Of Social Behavior In The Tibetan Macaque (Macaca Thibetana), Rose Amrhein Jan 2020

The Development Of Social Behavior In The Tibetan Macaque (Macaca Thibetana), Rose Amrhein

All Master's Theses

Social cognition is vital for the proper integration into adulthood for any highly social animal species. The development of social intelligence during the childhood and adolescence of a social organism affects the individual throughout its life. This social intelligence allows for the establishment and maintenance of bonds through the formation of empathy, the understanding of intention and emotion, and theory of mind in some species. Changes to the rate and effectiveness of social development could lead to an individual incapable of integrating into the social environment of adulthood. Yet, much still needs to be learned about the process and influences …


Size Estimation Of Pre‐Columbian Caribbean Fish, Sandrine Grouard, Sophia Perdikaris, Nídia Cristina Espíndola Rodrigues, Irvy R. Quitmyer Jan 2019

Size Estimation Of Pre‐Columbian Caribbean Fish, Sandrine Grouard, Sophia Perdikaris, Nídia Cristina Espíndola Rodrigues, Irvy R. Quitmyer

School of Global Integrative Studies: Faculty Publications

In this contribution, we present a methodological approach to the identification of pre‐Columbian Caribbean fisheries and examine the interrelationships of exploitation according to size for eight fish families, in a diachronic perspective for the Lesser Antilles. Based on the principles of size and growth allometries, biometric repositories have been reconstructed for modern families that represent different ecological environments: Holocentridae, Serranidae, Carangidae, Lutjanidae, Haemulidae, Scaridae, Acanthuridae, and Scombridae. The measured fish bone elements were selected based on their robustness and potential for recovery at archaeological sites. This resulted in a sample size totaling 563 modern osteological specimens, which provided reconstructed standard, …


Allomaternal Care By Conspecifics Impacts Activity Budgets Of Colobus Guereza Mothers, Dominique L. Raboin Aug 2018

Allomaternal Care By Conspecifics Impacts Activity Budgets Of Colobus Guereza Mothers, Dominique L. Raboin

Theses and Dissertations

In primate societies, caring for infants involves nursing, protection, provisioning, and carrying - all energetically taxing states for mothers. The cost of holding and carrying clinging infants often constrains mothers from moving and traveling, potentially reducing their food and energy intake. Alternatively, when an infant is physically separated from their mother they are at risk of predation from birds of prey or other large mammals. This requires a high level of vigilance from mothers, often further deterring them from acquiring the food and energy that they need. Allomaternal care (AMC) is hypothesized to provide mothers with a way to safely …


Playing It Cool: Characterizing Social Play, Bout Termination, And Candidate Play Signals Of Juvenile And Infant Tibetan Macaques (Macaca Thibetana), Kaitlin R. Wright, Jessica A. Mayhew, Lori K. Sheeran, Jake A. Funkhouser, Ronald S. Wagner, Li-Xing Sun, Jin-Hua Li May 2018

Playing It Cool: Characterizing Social Play, Bout Termination, And Candidate Play Signals Of Juvenile And Infant Tibetan Macaques (Macaca Thibetana), Kaitlin R. Wright, Jessica A. Mayhew, Lori K. Sheeran, Jake A. Funkhouser, Ronald S. Wagner, Li-Xing Sun, Jin-Hua Li

All Faculty Scholarship for the College of the Sciences

Play behaviors and signals during playful interactions with juvenile conspecifics are important for both the social and cognitive development of young animals. The social organization of a species can also influence juvenile social play. We examined the relationships among play behaviors, candidate play signals, and play bout termination in Tibetan macaques (Macaca

thibetana) during juvenile and infant social play to characterize the species play style. As Tibetan macaques are despotic and live in groups with strict linear dominance hierarchies and infrequent reconciliation, we predicted that play would be at risk of misinterpretation by both the individuals engaged in the play …


Response Of Three Species Of Monkeys To Caregiver Use Of Species-Typical Behavior, Lillian Anna Stolar Jan 2018

Response Of Three Species Of Monkeys To Caregiver Use Of Species-Typical Behavior, Lillian Anna Stolar

All Master's Theses

Caregivers are a primary part of captive monkey environments. When addressing psychological well-being of captive nonhuman primates, social environment should be considered. Chimpanzees, gorillas, New World monkeys, and Old World monkeys responded positively to interactions with caregivers. Some species showed increased affiliative behaviors and decreased abnormal or self-directed behaviors after interactions. These studies showed that caregivers can affect the behavior of nonhuman primates. Caregivers are underutilized as a source of social interaction for captive nonhuman primates. Utilizing species-typical behaviors during interactions, caregivers and nonhuman primates can communicate and interact in different ways that may be beneficial to both caregiver and …


From Icon Of Empire To National Emblem: New Evidence For The Fallow Deer Of Barbuda, Sophia Perdikaris, Allison Bain, Sandrine Grouard, Karis Baker, Edith Gonzalez, A. Rus Hoelzel, Holly Miller, Reaksha Persaud, Naomi Sykes Jan 2018

From Icon Of Empire To National Emblem: New Evidence For The Fallow Deer Of Barbuda, Sophia Perdikaris, Allison Bain, Sandrine Grouard, Karis Baker, Edith Gonzalez, A. Rus Hoelzel, Holly Miller, Reaksha Persaud, Naomi Sykes

School of Global Integrative Studies: Faculty Publications

Barbuda and Antigua’s national animal is the fallow deer, Dama dama dama, a species native to the eastern Mediterranean that has been transported around the world by people during the last 8000 years. The timing and circumstances by which fallow deer came to be established on Barbuda are currently uncertain but, by examining documentary, osteological and genetic evidence, this paper will consider the validity of existing theories. It will review the dynamics of human–Dama relationships from the 1500s AD to the present day and consider how the meaning attached to this species has changed through time: from a …


Enrichment Assessment For Geriatric African Old World Monkeys Under Human Care, Amanda E. Osborne Jan 2018

Enrichment Assessment For Geriatric African Old World Monkeys Under Human Care, Amanda E. Osborne

All Master's Theses

I proposed a study that determines whether enrichment regimes used for geriatric African Old World monkeys living under human care are effectively eliciting affiliative and active behaviors. I wanted to determine if alternating enrichment types used by the zoo staff were eliciting different social behaviors and locomotion in non-human primates based on different factors including ages, species, and sexes. My data collection took place at the Association of Zoos and Aquarium (AZA) accredited, Oregon Zoo in Portland, Oregon. I conducted my research from 11 June to 5 August 2017. I collected data from eight individuals of varying ages from three …


Re-Evaluating Captive Chimpanzee “Dominance”: Dominance Hierarchy And Chimpanzee-Caregiver Relationships At Chimpanzee Sanctuary Northwest, Jake Funkhouser Jan 2018

Re-Evaluating Captive Chimpanzee “Dominance”: Dominance Hierarchy And Chimpanzee-Caregiver Relationships At Chimpanzee Sanctuary Northwest, Jake Funkhouser

All Master's Theses

This thesis is composed of two journal-ready articles and an accompanying appendix with additional data and interpretation. Overall, this thesis describes and statistically analyzes dominance relationships in two nonhuman primate groups with novel methods, possible correlations between dominance and testosterone, and uncovers the prominent connection of caregivers to captive chimpanzee social networks. Chapter I addresses current and past variability in behavioral measures and statistical methods to derive dominance rankings. I propose a novel approach to using existing statistical techniques to analyze dominance ranks, context-dependent dominant structures (agonistic competitions, lack of agonism, privileged role, priority access to resources), the reliability of …


Does Genotype Correlate With Phenotype? Evaluating Ruffed Lemur (Varecia Spp.) Color Vision Using Subject Mediated Automatic Remote Testing Apparatus (Smarta), Raymond Vagell May 2017

Does Genotype Correlate With Phenotype? Evaluating Ruffed Lemur (Varecia Spp.) Color Vision Using Subject Mediated Automatic Remote Testing Apparatus (Smarta), Raymond Vagell

Theses and Dissertations

Ruffed lemur (Varecia spp.) color vision research was conducted using a multidisciplinary approach: psychophysics, genetic analysis, technology, and animal training. The behavioral manifestation of Varecia spp. trichromacy was shown using a touchscreen apparatus (SMARTA). Trichromats performed better than dichromats when discriminating red from green (G2 = 78.10, p < 0.001).


Do Non-Human Primates Have Gender?, Aaron Pelchat May 2017

Do Non-Human Primates Have Gender?, Aaron Pelchat

Senior Honors Projects

As activism for trans rights and gender equality becomes ever more prevalent in the current American political discourse, so too has there been a rise in questions about gender. Are sexuality and gender linked? Aren’t there only two genders? What is the difference between gender and sex? Is there a difference? How does one DO gender? Isn’t gender just something you are born with? Helping the public understand these questions is important to transgender and gender non-conforming individuals in a time when more people are “coming out of the closet” and identifying as genders other than cisgender. As an anthropologist, …


Family Dynamics In Eastern Hoolock (Hoolock Leuconedys), Javan (Hylobates Moloch), Pileated (H. Pileatus) And Northern White-Cheeked Gibbons (Nomascus Leucogenys) At The Gibbon Conservation Center, Alexandra Sheldon Jan 2017

Family Dynamics In Eastern Hoolock (Hoolock Leuconedys), Javan (Hylobates Moloch), Pileated (H. Pileatus) And Northern White-Cheeked Gibbons (Nomascus Leucogenys) At The Gibbon Conservation Center, Alexandra Sheldon

All Master's Theses

Few studies have explored the continuing relationship and behaviors within primate groups as the offspring mature and continue to depend on parental care. Between June 7 and July 7, 2016, I studied the family dynamics of 5 gibbon groups from 4 species: hoolock (Hoolock leuconedys), Javan (Hylobates moloch), pileated (H. pileatus) and northern white-cheeked (Nomascus leucogenys) gibbons housed at the Gibbon Conservation Center (GCC) in Santa Clarita, CA. I recorded all occurrences of approach, retreat, groom and play behaviors as well as the proximity individuals maintain between each other. I predicted: 1. …


Enrichment Use & Social Interactions In A Mixed-Species Enclosure Of Sumatran (Pongo Abelij) & Bornean Orangutans (P. Pygmaeus) & Northern White-Cheeked Gibbons (Nomascus Leucogenys), Emily Veitia Jan 2017

Enrichment Use & Social Interactions In A Mixed-Species Enclosure Of Sumatran (Pongo Abelij) & Bornean Orangutans (P. Pygmaeus) & Northern White-Cheeked Gibbons (Nomascus Leucogenys), Emily Veitia

All Master's Theses

Enrichment is an aspect of captive husbandry that has been shown to positively impact animals’ well-being and can be designed to encourage species-typical behaviors, such as foraging and arboreality. Enrichment can include housing together multiple compatible species. Orangutans are sympatric with siamangs and agile and Bornean gibbons, so several zoos house these four Asian ape species in the same enclosure, in part, as social enrichment. In my study, I observed enrichment use and social interactions in a mixed-species enclosure at the Oregon Zoo, which housed two Sumatran (Pongo abelii) and two Bornean (P. pygmaeus) orangutans with …


Use Of Vertical Enclosure Space And Species-Typical Locomotion By A Rehabilitating Spider Monkey (Ateles Fusciceps), Jake Funkhouser Jan 2017

Use Of Vertical Enclosure Space And Species-Typical Locomotion By A Rehabilitating Spider Monkey (Ateles Fusciceps), Jake Funkhouser

Undergraduate Honors Theses

With wild spider monkey populations in decline, investigations contributing to captive welfare, and successful rehabilitation and reintroduction knowledge is increasingly pressing. Quantifying and analyzing the appropriateness of naturalistic enclosure designs to foster species-typical behaviors is an effective way to address both of these needs. This study investigates enclosure space use, vertical space preference, substrate use, positional/postural modes, and interactions with human caregivers of a wild-caught, pet-trade rehabilitant Columbian black spider monkey (Ateles fusciceps rufiventris, N = 1). Video data collected daily from August to October 2015 via focal animal sampling (from 08:00 to 10:00) at Alouatta Sanctuary, Panama …


Chimpanzee (Pan Troglodytes) Space Use In A Sanctuary Setting, Holly Soubiea Jan 2017

Chimpanzee (Pan Troglodytes) Space Use In A Sanctuary Setting, Holly Soubiea

All Master's Theses

Studies of chimpanzee space use can provide information about chimpanzees’ preferences and welfare. This study examined the space use of 12 chimpanzees at Fauna Foundation, a chimpanzee sanctuary located in Carignan, Québec, Canada. Observers recorded the chimpanzees’ location, elevation, perimeter, and substrate using a scan sampling procedure. In total, there were 93 hours of data and 3,190 scans. The author used Geographical Information System (GIS) to plot the chimpanzees’ location for each scan. The chimpanzees used the largest rooms most often, were inside 61% (n = 1,942) of the time, an arm’s reach away from the perimeter 90% (n = …


The Social Behavior Of Rehabilitated Spider Monkeys (Ateles Geoffroyi), Anthony Denice Jan 2017

The Social Behavior Of Rehabilitated Spider Monkeys (Ateles Geoffroyi), Anthony Denice

All Master's Theses

The welfare of captive primates in laboratories, sanctuaries, and zoos is affected by various aspects of their environments. Although space restrictions increase aggression and stress-related behaviors in most captive animals, primates show diverse mechanisms for displacing stress and mitigating conflict. Many primates, including wild spider monkeys (genus Ateles), use these mechanisms flexibly to cope with social and environmental stressors. I investigated whether or not captive black-handed spider monkeys (Ateles geoffroyi) use behavioral strategies to cope with potential stressors in captivity. In particular, I tested whether an affiliative or avoidant strategy was used in response to changes in …


Questions On The Divergence Of Rhinopithecus Bieti, Melissa Reed Apr 2016

Questions On The Divergence Of Rhinopithecus Bieti, Melissa Reed

Georgia State Undergraduate Research Conference

No abstract provided.


Dama Dentition: A New Tooth Eruption And Wear Method For Assessing The Age Of Fallow Deer (Dama Dama), Frazer Bowen, Ruth F. Carden, Julie Daujat, Sandrine Grouard, Holly Miller, Sophia Perdikaris, Naomi Sykes Jan 2016

Dama Dentition: A New Tooth Eruption And Wear Method For Assessing The Age Of Fallow Deer (Dama Dama), Frazer Bowen, Ruth F. Carden, Julie Daujat, Sandrine Grouard, Holly Miller, Sophia Perdikaris, Naomi Sykes

School of Global Integrative Studies: Faculty Publications

Reliable aging techniques for wild animals are notoriously challenging to develop because of the scarcity of sizeable collections of known-age specimens. Without such techniques it is difficult to reconstruct hunting patterns, which is a significant problem for the examination of assemblages from pre-farming cultures. This paper presents a new method, based on mandibular tooth eruption and wear, for assessing the age of fallow deer. The method was developed from a large collection (n = 156) of known-age Dama dama specimens, has been blind tested by members of the zooarchaeological community and represents a user-friendly system with the potential to …


Reaction To Stimulus Figures In Chimpanzee Drawings, Alexandra Bobrinskoy Casti Jan 2016

Reaction To Stimulus Figures In Chimpanzee Drawings, Alexandra Bobrinskoy Casti

All Master's Theses

Seven captive chimpanzees (Pan troglodytes) produced drawings at the University of Oklahoma between November 1971 and November 1972. Chimpanzees drew on sheets of paper that were either blank or had a stimulus. The stimulus was located in the center or offset from the center. These drawings were scanned and digitized. Analysis tested whether chimpanzee mark placement was contingent on the location of stimulus figures. Centroid locations significantly changed between stimulus type for all drawing categories and among participants for free choice and central figure drawings. Participants drew in the empty space opposite offset figure drawings. Findings support previous …


Raccoon Scavenging And The Taphonomic Effects On Early Human Decomposition And Pmi Estimation, Jacob K. Smith Aug 2015

Raccoon Scavenging And The Taphonomic Effects On Early Human Decomposition And Pmi Estimation, Jacob K. Smith

Masters Theses

Forensic investigators must consider the effects of numerous taphonomic variables when estimating the postmortem interval (PMI) in forensic cases, such as temperature and insect activity. Efforts to quantify the decomposition process have been made. Megyesi et al. (2005) modified the categories and stages of decomposition presented by Galloway et al. (1989), and introduced a total body scoring system that uses accumulated degree-days to estimate PMI. However, none of these systems include the potential effects of animal scavenging. Galloway et al. (1989) stated that carnivorous activity occurs mainly during advanced decomposition and mummification. Haglund et al. (1989) specifically investigated canid scavenging …


Becoming Rabbit: Living With And Knowing Rabbits, Margo Demello Jul 2015

Becoming Rabbit: Living With And Knowing Rabbits, Margo Demello

Margo DeMello, PhD

Rabbits, like all animals (human and non-human), have rich internal lives, as people who live intimately with rabbits can attest.1 Living with house rabbits—where rabbits live indoors, without a cage or with minimal caging, as part of the human family—is, to me, the best way to gain some understanding of the rabbit psyche. In addition, living closely with rabbits opens up the possibilities of the humanrabbit relationship—a relationship which, until very recently, was one-sided and based on exploitation. Today, however, with the rise of the house rabbit movement, the subjectivity of rabbits has been exposed, leading to the possibility of …


Pantomime In Great Apes: Evidence And Implications, Ann E. Russon, Kristin Andrews Apr 2015

Pantomime In Great Apes: Evidence And Implications, Ann E. Russon, Kristin Andrews

Kristin Andrews, PhD

We recently demonstrated, by mining observational data, that forest-living orangutans can communicate using gestures that qualify as Pantomime. Pantomimes, like other iconic gestures, physically resemble their referents. More elaborately, pantomimes involve enacting their referents. Holding thumb and finger together at the lips and blowing between them to mean balloon is one example. Here we sketch evidence of pantomime in other great apes, methodological concerns, and sophisticated cognitive capabilities that great ape pantomimes suggest.


The “Babe” Vegetarians: Bioethics, Animal Minds And Moral Methodology, Nathan Nobis Mar 2015

The “Babe” Vegetarians: Bioethics, Animal Minds And Moral Methodology, Nathan Nobis

Nathan M. Nobis, PhD

Here I discuss the role the film “Babe” has played in helping people address these challenges and make this moral progress. It is thought that a significant number of young people (mostly girls, now young women) became vegetarians due to their seeing “Babe.” These people are often called “Babe Vegetarians,” influence by what has been called “The Babe Effect.” Many of their stories are found on the internet.


Javan Gibbon (Hylobates Moloch) Non-Vocal Social Communication And Gesture Use With Conspecifics, Melanie Bell Jan 2015

Javan Gibbon (Hylobates Moloch) Non-Vocal Social Communication And Gesture Use With Conspecifics, Melanie Bell

All Master's Theses

I explored gestures used by captive Javan gibbons (Hylobates moloch) at the Gibbon Conservation Center (Santa Clarita, CA). I hypothesized that a sender gibbon’s gesture modality would vary with the recipient gibbon’s attentional state and the sender would be equally likely to use all modalities (tactile, visual, actions, and facial expressions) when the recipient was attending (facing the sender), but would use more tactile gestures and actions when the recipient was non-attending (oriented away from the sender). I collected data from 10 individuals using all-occurrences sampling and an ethogram to score behaviors from video recordings. In 1,143 interactions, …