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Articles 1 - 30 of 57
Full-Text Articles in Life Sciences
The Flora Of The Northern Part Of The Emirdağ Mountains, Yavuz Bülent Köse, Ati̇la Ocak
The Flora Of The Northern Part Of The Emirdağ Mountains, Yavuz Bülent Köse, Ati̇la Ocak
Turkish Journal of Botany
This study concerns the flora of the northern part of the Emirdağ nountains. According to Davis's grid system, the research area lies in B3 square. Four hundred and ninety-five taxa belonging to 279 genera and 70 families were found after the identification of 754 plant specimens collected from the research area. Forty-seven taxa are new records for the B3 grid square. The rate of endemism is 15.9%. The distribution of taxa within the phytogeographical regions is as follows: Irano-Turanian elements 17.9%, Mediterranean elements 8.8% and Euro-Siberians 5.4%. Moreover, 67.9% of total taxa are undetermined as far as the phytogeographic area …
The Sections Of Astragalus L. With Bifurcating Hairs In Iran, Farrokh Ghahremaninejad
The Sections Of Astragalus L. With Bifurcating Hairs In Iran, Farrokh Ghahremaninejad
Turkish Journal of Botany
In this study Astragalus L. with bifurcating hairs in sectional levels in Iran are discussed (subgenus Cercidothrix Bunge). This subgenus has more than 800 species in the Old World. Major areas of species endemism for this subgenus occur in Turkey, Iran, Kazakhstan and Afghanistan. Iran, with 158 species, is one of the most important centres of diversity. Leucocercis Bunge is the only endemic section for Iran, although other sections have some species in different countries. Thus having a knowledge of the distribution quality in other areas is also important. In reality, this study is part of the author's research on …
Additional Records Of Raphanus L. And Campanula L. For The Flora Of Turkey, Bi̇rol Mutlu
Additional Records Of Raphanus L. And Campanula L. For The Flora Of Turkey, Bi̇rol Mutlu
Turkish Journal of Botany
Two taxa, R. raphanistrum L. subsp. rostratus (DC.) Thell. and Campanula hierosolymitana Boiss., collected from southern Anatolia are described as a second records for the Flora of Turkey. Different taxonomists had collected these taxa from Turkey, but these collected samples were not identified correctly. These samples were not recorded in the first or sixth volume or the other 2 supplements of the Flora of Turkey in spite of these misidentifications having been published in different papers. In this paper, these taxa that are added to the Flora of Turkey are presented for the interest of Turkish taxonomists. The descriptions and …
Gypsophila Sphaerocephala Fenzl Ex Tchihat.: A Boron Hyperaccumulator Plant Species That May Phytoremediate Soils With Toxic B Levels, Mehmet Babaoğlu, Sai̇t Gezgi̇n, Ali̇ Topal, Bayram Sade, Hüseyi̇n Dural
Gypsophila Sphaerocephala Fenzl Ex Tchihat.: A Boron Hyperaccumulator Plant Species That May Phytoremediate Soils With Toxic B Levels, Mehmet Babaoğlu, Sai̇t Gezgi̇n, Ali̇ Topal, Bayram Sade, Hüseyi̇n Dural
Turkish Journal of Botany
Analyses were carried out to identify boron (B) hyperaccumulating plant species in an actively B- mined area of Kırka, Eskişehir, Turkey. Only 4 plant species, Gypsophila sphaerocephala Fenzl ex Tchihat. var. sphaerocephala (Caryophyllaceae), Gypsophila perfoliata L. (Caryophyllaceae), Puccinellia distans (Jacq.) Parl. subsp. distans (Gramineae) and Elymus elongatus (Host) Runemark subsp. turcicus (McGuire) Melderis (Gramineae), were identified in the highest B- containing sections of the mine. The species were found growing successfully under high total (8900 mg kg^{-1}) and available (277 mg kg^{-1}) soil B concentrations. Among these plant species, G. sphaerocephala contained considerably higher B concentrations in its above-ground parts …
Effect Of Thidiazuron On Shoot Regeneration From Different Explants Of Lentil (Lens Culinaris Medik.) Via Organogenesis, Khalid Mahmood Khawar, Cengi̇z Sancak, Serkan Uranbey, Sebahatti̇n Özcan
Effect Of Thidiazuron On Shoot Regeneration From Different Explants Of Lentil (Lens Culinaris Medik.) Via Organogenesis, Khalid Mahmood Khawar, Cengi̇z Sancak, Serkan Uranbey, Sebahatti̇n Özcan
Turkish Journal of Botany
Thidiazuron (TDZ) is among the most active cytokinin like substances and induces greater in vitro shoot proliferation than many other cytokinins in many plant species. Leaf, stem, stem node and cotyledonary node explants of 2 extensively cultivated Turkish lentil cultivars, Ali Dayı and Kayı 91, were cultured on Murashige and Skoog (MS) media supplemented with various concentrations of TDZ. The present study was conducted to develop a rapid and efficient shoot regeneration system suitable for the transformation of lentil (Lens culinaris Medik.) using TDZ. Cotyledonary nodes and stem nodes after the initial callus stage regenerated prolific adventitious shoots via organogenesis. …
Computer Technology For The Future Of Sw Asiatic Botany, R. J. Pankhurst
Computer Technology For The Future Of Sw Asiatic Botany, R. J. Pankhurst
Turkish Journal of Botany
Much computer software already exists for floristic and monographic studies. Database software can hold all the data required for a Flora, and can process it much more accurately. The fundamental data tables are those for names, authorities, people, specimens, bibliography, geography and distribution. To these can be added other data such as descriptions, keys, images, maps, chromosomes and vernacular names. The same data system can be used for collection management and making herbarium labels. It is very important to use data standards such as for books, journals and authorities. Computerised descriptions and keys can also be constructed and held in …
Morphological And Anatomical Investigations On The Saponaria Kotschyi Boiss. (Caryophyllaceae), Ebru Ataşlar
Morphological And Anatomical Investigations On The Saponaria Kotschyi Boiss. (Caryophyllaceae), Ebru Ataşlar
Turkish Journal of Botany
The morphological, anatomical and micromorphological features of Saponaria kotschyi Boiss. have been studied on plants collected from Osmangazi University-Meşelik Campus, Eskişehir. S. kotschyi is an endemic species and a new record from Eskişehir. In general, the results obtained from morphological studies were consistent with the description given in the Flora of Turkey, except for the calyx, as the calyx of S. kotschyi flowers not only had glandular hairs but also eglandular hairs. The anatomical studies on cross-sections of the root, stem, leaves and surface sections of leaves were presented and compared with previous findings. Furthermore, the micromorphological properties of the …
Value Of Trichome Characteristics For The Separation Of Bifurcating Hairy Astragalus L. (Fabaceae) At The Sectional Level, Farrokh Ghahremaninejad
Value Of Trichome Characteristics For The Separation Of Bifurcating Hairy Astragalus L. (Fabaceae) At The Sectional Level, Farrokh Ghahremaninejad
Turkish Journal of Botany
The attachment of hairs in sections of Astragalus L. may be either basal or lateral (bifurcate). This study focuses on Astragali with bifurcating hairs in the Old World (subgenus Cercidothrix Bunge). In this study, it appears that the ratio of hair arm length, and hair form and length are usually stable at the sectional level. The proposition of uniting some sections based on morphological characters is confirmed by hair characters; for example, uniting sections Tamias Bunge, Acantherioceras Rech. f. and Pseudoerioceras Ulzijchutag with Erioceras Bunge. The most important characters in the separation of the sections are macromorphological, but hair characters …
A Floristical Study On Poaceae Spp. Growing Naturally In Malatya Province, Turan Arabaci, Bayram Yildiz
A Floristical Study On Poaceae Spp. Growing Naturally In Malatya Province, Turan Arabaci, Bayram Yildiz
Turkish Journal of Botany
This study includes species of Poaceae (Gramineae) growing in Malatya province. According to the grid system used in the Flora of Turkey, the research area is situated in the B6, B7, C6 and C7 squares. Approximately 346 plant specimens were collected from the area. At the end of the study 89 species (101 taxa) belonging to 47 genera were identified. Species' totals of the larger genera in the study area as follows: Bromus L. 10, Elymus L. 6, Aegilops L. 6, Hordeum L. 5, Alopecurus L. 4, Poa L. 4, and Stipa L. 4. The phytogeographic elements represented in the …
Plant Life Of South-West Asia Symposia: Vi - Past And Future, Ian C. Hedge
Plant Life Of South-West Asia Symposia: Vi - Past And Future, Ian C. Hedge
Turkish Journal of Botany
No abstract provided.
Modern Approaches To Floristics And Their Impact On The Region Of Sw Asia, Vernon Hilton Heywood
Modern Approaches To Floristics And Their Impact On The Region Of Sw Asia, Vernon Hilton Heywood
Turkish Journal of Botany
Attitudes to floristics have changed considerably during the past few decades as a result of increasing and often more focused consumer demands, heightened awareness of the threats to biodiversity, information flow and overload, and the application of electronic and web-based techniques to information handling and processing. This paper will examine these concerns in relation to our floristic knowledge and needs in the region of SW Asia. Particular reference will be made to the experience gained from the Euro+Med PlantBase project for the preparation of an electronic plant-information system for Europe and the Mediterranean, with a single core list of accepted …
On The Distribution And Relationships Of The South-West Asian Species Of Eryngium L. (Apiaceae-Saniculoideae), Arno Worz
Turkish Journal of Botany
This paper deals with the Old World species of Eryngium L. which grow mostly in regions with a Mediteranean type climate. Two centres of diversity are visible: one in the Western Mediterranean (Iberian Peninsula, Morocco), and the other one in South-West Asia. Distribution maps are presented for several species from South-West Asia, which is probably the genus' most important centre of diversity. A preliminary new classification of the genus Eryngium is presented, including a new combination (Eryngium subgenus Fruticosum (Wolff) Wörz stat. nov.). Both the species and the sectional diversity are conspicuously present in Turkey, where 9 of 14 sections …
Possibility Of An Information System On Plants Of South-West Asia With Particular Reference To The Turkish Plants Data Service (Tubives), M. Teki̇n Babaç
Possibility Of An Information System On Plants Of South-West Asia With Particular Reference To The Turkish Plants Data Service (Tubives), M. Teki̇n Babaç
Turkish Journal of Botany
For quick and accurate access to information about a flora of a country it is becoming increasingly important to develop databases. Databases developed and provided by international organisations are expensive to use and difficult to maintain without outside help. It is therefore important for developing countries to develop their own plant databases best suited to their requirements. One such example is the Turkish Plant Data Service (TUBIVES), designed by Turkish systematists. This database is described with some samples, and the possibility of creating a South-West Asian Plants Information Service (SWAP-IS) is briefly discussed.
Flora-Writing Exemplified By Classical, Conservational And Unconventional Models, Kit Tan
Flora-Writing Exemplified By Classical, Conservational And Unconventional Models, Kit Tan
Turkish Journal of Botany
There are many ways to compile a Flora. The Flora of Turkey is a remarkable 10-volume work completed within a quarter-century of its inception. A review of the factors contributing to the Flora's success is provided. The Flora of Turkey is a classical Flora with an additional supplement recently produced by Turkish botanists. Flora Hellenica is another example of a classical Flora. Some aspects of its background and production are briefly mentioned. Conservationists consider the endemic flora of a country needs protection for all time, and an opportunity arose to prepare a three-volume work on the endemic flora of Greece. …
Some Poisonous Fungi Of East Anatolia, Kenan Demi̇rel, Yusuf Uzun, Abdullah Kaya
Some Poisonous Fungi Of East Anatolia, Kenan Demi̇rel, Yusuf Uzun, Abdullah Kaya
Turkish Journal of Botany
This study was carried out on some poisonous macrofungi samples collected from the East Anatolia region. As a result of field and laboratory studies, 36 poisonous taxa belonging to Ascomycetes and Basidiomycetes were identified. The identified taxa are given with their distributions, habitat and substrates, collection dates and collectors' numbers. One of the species, Amanita eliae Qúel. (Amanitaceae), is a new record for the poisonous fungi of Turkey.
Jasione L. Taxonomy And Phylogeny, Fatima Sales, Ian C. Hedge, William Eddie, Jill Preston, Michael Moeller
Jasione L. Taxonomy And Phylogeny, Fatima Sales, Ian C. Hedge, William Eddie, Jill Preston, Michael Moeller
Turkish Journal of Botany
Jasione L., a genus with small flowers in compact ebracteolate heads subtended by involucral bracts, is very isolated in the Campanulaceae. It is mainly a Mediterranean-European genus with c. 15, mostly polymorphic, species, which show evidence of recent speciation, possibly as recent as the last glaciation. In the west of its distributional range, where the greatest morphological variation occurs, there has been much recent investigation on morphology, anatomy, phenetics, molecular evidence and nomenclature. This research has provided considerable evidence for a reduction in the number of taxa that should be recognised. On-going research is focused on more detailed molecular studies …
Seasonal Distribution Of Phytoplankton In Orduzu Dam Lake (Malatya, Turkey), Ahmet Kadri̇ Çeti̇n, Bülent Şen
Seasonal Distribution Of Phytoplankton In Orduzu Dam Lake (Malatya, Turkey), Ahmet Kadri̇ Çeti̇n, Bülent Şen
Turkish Journal of Botany
The species composition and seasonal distribution of phytoplankton in Orduzu Dam Lake was studied for a year. Diatoms (Bacillariophyta) were most diverse, followed by green algae (Chlorophyta), blue-green algae (Cyanophyta), euglenoids (Euglenophyta) and dinoflagellates (Dinophyta). A total of 117 taxa were recorded and the phytoplankton of the lake contained a large number of detached benthic algae. Phytoplankton assemblages were dominated by diatoms in all the periods investigated and centric diatoms were the most abundant. Overall phytoplankton density was high during the spring and summer months and the highest phytoplankton density was observed in August. The abundance of phytoplankton was positively …
Floristic Characteristics Of Beydağı (Malatya), Bayram Yildiz, Zeli̇ha Bahçeci̇oğlu, Turan Arabaci
Floristic Characteristics Of Beydağı (Malatya), Bayram Yildiz, Zeli̇ha Bahçeci̇oğlu, Turan Arabaci
Turkish Journal of Botany
The floristic characteristics of Beydağı are given and the vascular plant species growing there are documented. Beydağı is situated in the south of Malatya province and is in the Irano-Turanian phytogeographical region. The research was carried out between 1986 and 2000. Two thousand one hundred plant specimens were collected from the area. At the end of the study, 873 vascular plant species (897 taxa, 185 subsp., 112 var.) and 357 genera belonging to 73 families were identified: 140 taxa (16.9%) are endemic to Turkey. The phytogeographic elements are represented in the study area as follows: Irano-Turanian 364 (40.44%), Mediterranean 98 …
Flora Of The Natural Conservation Area In Adana-Yumurtalık Lagoon (Turkey), Haşi̇m Altinözlü
Flora Of The Natural Conservation Area In Adana-Yumurtalık Lagoon (Turkey), Haşi̇m Altinözlü
Turkish Journal of Botany
This study includes the natural and cultivated plants in the Natural Conservation Area in Yumurtalık Lagoon. In an evalution of 450 herbarium specimens collected from this area in 1997 and 1998, a total of 186 genera and 234 species belonging to 65 families were identified: 223 of the 234 species are natural and 11 are cultivated. Three of the 234 taxa are endemics (1.3%), 26 taxa (11.6%) are Mediterranean elements, 13 taxa (5.8%) are East Mediterranean elements, 6 taxa (2.6%) are Irano-Turanian elements, 5 taxa (2.2%) are Euro-Siberian elements, and 1 taxon (0.4%) is a Hyrcano-Euxine element. The other 172 …
Influence Of Some Plant Extracts And Microbioagents On Some Physiological Traits Of Faba Bean Infected With Botrytis Fabae, Yehia A.G. Mahmoud, Mohsen Ebrahim, Magda M. Aly
Influence Of Some Plant Extracts And Microbioagents On Some Physiological Traits Of Faba Bean Infected With Botrytis Fabae, Yehia A.G. Mahmoud, Mohsen Ebrahim, Magda M. Aly
Turkish Journal of Botany
Laboratory and greenhouse experiments were conducted to assess the efficacy of Eucalyptus citriodora Hook., Ipomoea carnea Jacq., Cuminum cyminum L., Allium sativum L. and Hyoscyamus muticus L. leaf extracts, and of Streptomyces exfoliatus (Waksman & Curtis) Waksman & Henrici (S) and Trichoderma harzianum Rifai (T) in controlling Botrytis fabae, which causes chocolate spot disease in the faba bean. Laboratory studies supported the use of leaf extracts of E. citriodora (Ex. 1) and I. carnea (Ex. 2) in preference to other extracts for controlling the mycelial growth of B. fabae. In addition, the mixture S + T was the best of …
A Morphological And Anatomical Study On Endemic Alkanna Haussknechtii Bornm. (Boraginaceae), Critically Endangered In Turkey, Öznur Ergen Akçi̇n, Nezahat Kandemi̇r, Arzu Cansaran
A Morphological And Anatomical Study On Endemic Alkanna Haussknechtii Bornm. (Boraginaceae), Critically Endangered In Turkey, Öznur Ergen Akçi̇n, Nezahat Kandemi̇r, Arzu Cansaran
Turkish Journal of Botany
Alkanna haussknechtii Bornm., which occurs only in the vicinity of Amasya, is critically endangered. The morphological and anatomical properties of the species, an endemic plant of the Irano-Turanian phytogeographic region, were investigated. The root is perennial. The leaves are ecvifacial and have stomata cells that are anisocytic and anomocytic. The stomata index is 18.048 on the upper epidermis and 15.428 on the lower epidermis.
The Genus Crataegus L. (Rosaceae) With Special Reference To Hybridisation And Biodiversity In Turkey, Ali̇ Arslan Dönmez
The Genus Crataegus L. (Rosaceae) With Special Reference To Hybridisation And Biodiversity In Turkey, Ali̇ Arslan Dönmez
Turkish Journal of Botany
The genus Crataegus L. has approximately 200 species worldwide. However, the number of the species is increased to 1200 species by some taxonomists. Traditionally, the genus is placed in the subfamily Maloideae and it is closely related to the genera Pyracantha M.Roem., Mespilus L. and Hesperomeles Lindl. The latest phylogenetic analysis of the genus shows that it is closest to the genus Osteomeles Lind. Crataegus is divided into 40 sections by various taxonomists. The genetic diversity centre of the section Crataegus ranges from Turkey to Iran. According to field observations in Turkey, new diversity centres are proposed. The latest studies …
Recent Developments In International Biological Nomenclature, Werner Greuter
Recent Developments In International Biological Nomenclature, Werner Greuter
Turkish Journal of Botany
After a period of several decades of general neglect, biological systematics and taxonomy are again surfacing to the awareness of the scientific community and of policy makers. This we owe to the fact that they are the core disciplines of biodiversity research, now perceived to be a priority task in view of the impending threat of loss through extinction of a great number of species, and the wholesale destruction of ecosystems. Yet the renaissance of systematics and taxonomy is hampered by the inadequacy of the traditional rules governing the formation and use of scientific names of organisms. Names are a …
The High Mountain Vegetation Of Turkey - A State Of The Art Report, Including A First Annotated Conspectus Of The Major Syntaxa, Gerald Parolly
The High Mountain Vegetation Of Turkey - A State Of The Art Report, Including A First Annotated Conspectus Of The Major Syntaxa, Gerald Parolly
Turkish Journal of Botany
This contribution spotlights the present research on the (oreal) subalpine to subnival mountain vegetation of Turkey. It concisely compiles our present knowledge, which is very heterogeneous in geographical terms. The situation is comparatively good in NW and W Anatolia and the western half of the Taurus range, from where a first consolidation stage can be reported. By contrast, the E Taurus remains a largely unexplored area. The Pontic chains have hitherto attracted a few local researchers only, and those important pioneering accounts cover less than half of the vegetation types actually present. The results of the last years' field work …
Origin And Maintenance Of Rhododendron X Sochadzeae, A Fertile F1 Hybrid Which Occupies An Ecotone Between R. Ponticum And R. Caucasicum In Turkey, Richard I. Milne, Sali̇h Terzi̇oğlu, Richard J. Abbott
Origin And Maintenance Of Rhododendron X Sochadzeae, A Fertile F1 Hybrid Which Occupies An Ecotone Between R. Ponticum And R. Caucasicum In Turkey, Richard I. Milne, Sali̇h Terzi̇oğlu, Richard J. Abbott
Turkish Journal of Botany
Isolating barriers between sympatric interfertile species are maintained by processes that occur within their hybrid zones. Although the effects of intrinsic selection on hybrid fitness are well known, less is understood about extrinsic fitness variation. At Tiryal Dağı, NE Turkey, Rhododendron x sochadzeae Charadze & Davlianidze (R. ponticum L. x caucasicum Pall.) forms large populations in which neither segregation nor backcrossing occur, in habitats intermediate between those of its parents. Using single-copy species-specific RAPD and ISSR markers, it was determined that all R. x sochadzeae plants are F1s, and that there are many separate genets present. Through hand pollination experiments, …
The Asian Umbelliferae Biodiversity Database (Asium) With Particular Reference To South-West Asian Taxa, M. G. Pimenov, M. V. Leonov
The Asian Umbelliferae Biodiversity Database (Asium) With Particular Reference To South-West Asian Taxa, M. G. Pimenov, M. V. Leonov
Turkish Journal of Botany
An original database, ASIUM, on Asian Umbelliferae has been compiled at the Moscow State University to include all accepted names and synonyms, principal citations in regional Floras, monographs and critical articles, protologue data and distribution in countries, their regions and provinces. A special option permits the arrangement of synomyms, which can be divided into several homotypic groups. It is also possible to extract information for analysis and comparison, for instance, of species sets from different regions to estimate their similarity. ASIUM is a mobile monograph of the Umbelliferae for a continent, with their maximal biodiversity at the generic and specific …
Wild Edible Plants Of The Bodrum Area (Muğla, Turkey), Füsun Ertuğ
Wild Edible Plants Of The Bodrum Area (Muğla, Turkey), Füsun Ertuğ
Turkish Journal of Botany
The town of Bodrum is situated on a peninsula in south-western Turkey in the province of Muğla. The research project on the Useful Plants of the Bodrum Area has been collecting data since October 1999 on the traditional uses of plants for food, medicine, fodder, fuel, handicrafts and other purposes. The information was collected over a two-and-a-half-year period from informants of various ages and background categories in Bodrum and the surrounding villages, with the help of about 25 volunteers. In addition to the informant-based research, year-round surveys of weekly markets in Bodrum, Milas and Muğla have been carried out to …
Some Morphological And Anatomical Investigations On Autumn Species Of Crocus L. Occurring In Şanlıurfa, Hasan Akan, İsmai̇l Eker
Some Morphological And Anatomical Investigations On Autumn Species Of Crocus L. Occurring In Şanlıurfa, Hasan Akan, İsmai̇l Eker
Turkish Journal of Botany
Some morphological and anatomical characteristics of Crocus pallasii Goldb. subsp. turcicus B.Mathew and Crocus cancellatus Herbert subsp. damascenus (Herbert) B.Mathew, which flower in autumn, were investigated. Detailed descriptions of these two taxa of Crocus L., which occur in Şanlıurfa, are provided and their morphological characters are compared with those given in the Flora of Turkey. The specimens were collected from different parts of Şanlıurfa within C7 of the grid system; their habitat and population status were also determined. Their anatomical structure are illustrated.
Taxonomical And Ecological Investigations On Some Geophytes Growing Around Denizli Province (Turkey), Ali̇ Çeli̇k, Mehmet Çi̇çek, Gürkan Semi̇z, Mustafa Karincali
Taxonomical And Ecological Investigations On Some Geophytes Growing Around Denizli Province (Turkey), Ali̇ Çeli̇k, Mehmet Çi̇çek, Gürkan Semi̇z, Mustafa Karincali
Turkish Journal of Botany
In this study, the taxonomical and ecological features of some geophytes which grow naturally around Denizli province have been investigated. From this research, which was performed between 1996 and 2000, 31 species belonging to 8 families have been determined. Thirteen soil parameters from 31 localities were analysed in order to establish plant and soil relations. The ecological spectra of the species are considered, comprising 16 species in macchie and phrygana (50%); 1 species in fields (1%); 7 species in high mountains (23%) and 8 species in forests (26%).
A New Record For The Flora Of Turkey: Ambrosia Tenuifolia Spreng. (Compositae), Lütfi̇ Behçet
A New Record For The Flora Of Turkey: Ambrosia Tenuifolia Spreng. (Compositae), Lütfi̇ Behçet
Turkish Journal of Botany
Ambrosia tenuifolia Spreng. (Compositae) is a new addition to the flora of Turkey from Malatya (in the East Anatolia region). The specimens collected from Malatya have different properties from the known diagnostic characteristics of the species. A. tenuifolia is a monoecious plant without glands. Its female capitula are in the axils of the upper leaves in its original description, but some of our specimens have only female capitula (female capitula in terminal racemes and lower parts of the racemes bracteate, but upper parts ebracteate) and there are many glands particularly on the tops of their fruits. The diagnostic characteristics, ecology …