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Stem Development, Medullary Bundles, And Wood Anatomy Of Croton Glandulosus Var. Septentrionalis (Euphorbiaceae), Sheila M. Hayden, W. John Hayden Jan 1994

Stem Development, Medullary Bundles, And Wood Anatomy Of Croton Glandulosus Var. Septentrionalis (Euphorbiaceae), Sheila M. Hayden, W. John Hayden

Biology Faculty Publications

Anatomy and development of vascular tissues in the annual stems of Croton glandulosus var. septentrionalis are described. In primary stages of growth the stem possesses a eustele of bicollateral bundles; international phloem is notably more extensive than the external. In addition to a vascular cambium and secondary xylem that form in the usual fashion, additional cambia add cells to the internal phloem portion of the bicollateral bundles, forming well-marked medullary bundles at the perimeter of the pith. At first, the perimedullary cambial strands produce only internal secondary phloem; later, internal secondary xylem is present, the medullary bundles have an inverted …


Systematic Anatomy Of Euphorbiaceae Subfamily Oldfieldioideae I. Overview, W. John Hayden Jan 1994

Systematic Anatomy Of Euphorbiaceae Subfamily Oldfieldioideae I. Overview, W. John Hayden

Biology Faculty Publications

The biovulate subfamily Oldfieldioideae of Euphorbiaceae, characterized by spiny pollen, is an otherwise apparently diverse assemblage of mostly Southern Hemisphere trees and shrubs that traditionally have been allied with genera of Phyllanthoideae and Porantheroideae sensu Pax and Hoffmann. Although fairly diverse anatomically, the following structures characterize the subfamily with only a few exceptions: pinnate brochidodromous venation with generally randomly organized tertiary and higher order venation; foliar and petiolar glands absent; unicellular or unbranched uniseriate trichomes; latex absent; mucilaginous epidermis or hypodermis; brachyparacytic stomata; vessel elements with simple perforation plates and alternate, often very small, intervascular pits; thick-walled nonseptate imperforate tracheary …