Digital Commons Network^™

Description Of A New Species Of Aleuroparadoxus Quaintance And Baker (Hemiptera: Aleyrodidae) From Guatemala, José Francisco García Ochaeta, Anil Kumar Dubey

Insecta Mundi

Puparia of a new whitefly, Aleuroparadoxus marisae García-Ochaeta and Dubey, new species (Hemiptera: Aleyrodidae), collected on leaves of Brosimum alicastrum Sw. in Santa Ana, Petén, Guatemala, and on Pimenta dioica (L). Merr. in Calzada Mopán, Dolores, Petén are described and illustrated. Drawings of morphological structures and differential diagnosis of the puparium are provided, and differences from congeners are discussed. A key to puparia of the Aleuroparadoxus Quaintance and Baker species is included.

New Distributional Records For Mexican Cleridae (Coleoptera) With The Description Of Three New Species, Jacques Rifkind

Insecta Mundi

New state records for 32 species of Mexican Cleridae (Coleoptera) are presented. In addition, 10 species of Cleridae are reported for Mexico for the first time (new country records). Cymatodera bezarki new species and Enoclerus sepultura new species are described from Chiapas, Mexico. Enoclerus primulus new species is described from Chiapas, Mexico and El Salvador.

Approximately 550 species of Cleridae are known from Mexico. These are divided among approximately 63 gen­era, of which the most speciose are Cymatodera Gray (ca. 110 species), Enoclerus Gahan (ca. 105 species) and Phyllobaenus Dejean (ca. 40 species). Faunistic studies are limited to Vaurie’s (1952) …

On The Firefly (Coleoptera: Lampyridae) Species Of Carl Linnaeus, Oliver Keller

Insecta Mundi

The firefly species described by Carl Linnaeus in 1758 and 1767 (Coleoptera: Lampyridae) were checked to determine the actual dates of publication. Nine out of twelve species were originally described in 1758 and not in 1767 as published in the majority of firefly literature. Lampyris hespera Linnaeus, 1767 as a junior synonym of Aspisoma lampyris (Linnaeus, 1758) is established.

Se revisaron las especies de luciérnagas descritas por Carl Linnaeus in 1758 y 1767 (Coleoptera: Lampyridae) para la determinación de fechas de publicación. Nueve de las doce especies fueron originalmente descritas en 1758 y no en 1767 como aparece publicado en …

Synopsis Of The Subfamily Carventinae In New Zealand (Heteroptera: Aradidae), Marie-Claude Larivière, André Larochelle

Insecta Mundi

The subfamily Carventinae (Heteroptera: Aradidae) is revised for New Zealand. Eight genera and fifteen species are recognized.

One genus and six species are described as new: Carventaptera hallae Larivière and Larochelle new species, Lissaptera heissi Larivière and Larochelle new species, Modicarventus kirmani Larivière and Larochelle new species, Neocarventus montanus Larivière and Larochelle new species, Neocarventus northlandicus Larivière and Larochelle new species, Neocarventus potterae Larivière and Larochelle new species, Tuataraptera Larivière and Larochelle new genus.

One new combination is established: Neocarventus uncus Kirman, 1989 = Tuataraptera unca (Kirman, 1989).

One new synonymy is made: …

First Host Record, Nesting Behavior, And Taxonomic Position Of The Spider Wasp Genus Hesperopompilus Evans And Some Other Evans Genera (Hymenoptera: Pompilidae), Frank E. Kurczewski, Rick C. West, James P. Pitts

Insecta Mundi

First host record, prey transport, and burrow excavation are described for Hesperopompilus sp., an undescribed, rare spider wasp (Hymenoptera: Pompilidae) from Texas. Taxonomic, ecological, and behav­ioral examination of the genus subsequently led to an investigation of the previously related Perissopompilus Evans and Xerochares Evans. Taxonomic, host preference, nesting behavior, and phylogenomic relationships of the three taxa are discussed along with those of Xenopompilus Evans. The molecular connection of Perisso­pompilus and Allochares Banks is supported by their common use of host species of Filistatidae.

Evans (1951), in his taxonomic study of the spider wasp tribe Pompilini (Hymenoptera: Pompilidae: Pompilinae), described the …

Host Selection And Nesting Behavior Of Nearctic Trapdoor Spider-Hunting Spider Wasps (Hymenoptera: Pompilidae: Pepsinae, Pompilinae), Frank E. Kurczewski, Rick C. West

Insecta Mundi

Host records and nesting behavior of the Nearctic trapdoor spider-hunting spider wasps (Hy­menoptera: Pompilidae) Calopompilus Ashmead and Priocnemissus Haupt (Pepsinae: Pepsini) and Aporus Spinola and Psorthaspis Banks (Pompilinae: Aporini) are reviewed, investigated, compared, and discussed. First time incidental trapdoor spider host records for Priocnemis (Priocnemissus) minorata Banks (Pepsi­nae: Pepsini), Anoplius (Lophopompilus) carolina (Banks) (Pompilinae: Pompilini), and Notocyphus dorsalis dorsalis Cresson (Notocyphinae: Notocyphini) are included, although they are not typical trapdoor spider-hunting spider wasp species. The Palearctic Aporus (Aporus) unicolor Spinola, A. (Aporus) bicolor Spinola and A. (Aporus) planiceps (Latreille) are …

Acmaeodera Moesta Dugès, 1891 (Coleoptera: Buprestidae), A Poorly Known Species From Mexico, Richard L. Westcott, Jesus Romero-Napoles

Insecta Mundi

Two new state records are reported for the Mexican Acmaeodera moesta Dugès (Coleoptera: Bu­prestidae), a lectotype is designated for it, and clarifying discussion is provided.

According to Westcott (2020) there are 152 described valid species of Acmaeodera Eschscholtz in Mexico. Acmaeodera moesta Dugès (Fig. 1–3) was described from the state of Guanajuato, and it has not since been men­tioned in the literature except for listings in catalogs and checklists. Herein we record it from the states of México (MEX) and San Luis Potosí (SLP), designate a lectotype, and provide a detailed discussion.

A New Species Of Chrysina Kirby (Coleoptera: Scarabaeidae: Rutelinae) From The Sierra Madre Occidental Of Mexico, With Notes On The Type Locality Of Chrysina Adelaida (Hope, 1841), David C. Robacker, David C. Hawks, Naidely Castillo, Brianna Miranda

Insecta Mundi

This paper presents a taxonomic review of the genus Boreocanthon Halffter, restored generic sta­tus, a group of ball-rolling (telocoprid) dung beetles (Coleoptera: Scarabaeidae: Scarabaeinae) endemic to North America. The genus, heretofore treated as a subgenus of Canthon, comprises 13 species, each keyed, diagnosed, illustrated and presented with information on distribution, relationships, and biology (*signifies restored generic combination): *Boreocanthon ateuchiceps (Bates, 1887); B. coahuilensis (Howden, 1966); *B. depressipennis (LeConte, 1859c); *B. ebenus (Say, 1823); *B. forreri (Bates, 1887); *B. integricollis (Schaef­fer, 1915); *B. lecontei (Harold, 1868); …

Taxonomic Review Of The North American Dung Beetle Genus Boreocanthon Halffter, 1958 (Coleoptera: Scarabaeidae: Scarabaeinae: Deltochilini), W. D. Edmonds

Insecta Mundi

This paper presents a taxonomic review of the genus Boreocanthon Halffter, restored generic sta­tus, a group of ball-rolling (telocoprid) dung beetles (Coleoptera: Scarabaeidae: Scarabaeinae) endemic to North America. The genus, heretofore treated as a subgenus of Canthon, comprises 13 species, each keyed, diagnosed, illustrated and presented with information on distribution, relationships, and biology (*signifies restored generic combination): *Boreocanthon ateuchiceps (Bates, 1887); B. coahuilensis (Howden, 1966); *B. depressipennis (LeConte, 1859c); *B. ebenus (Say, 1823); *B. forreri (Bates, 1887); *B. integricollis (Schaef­fer, 1915); *B. lecontei (Harold, 1868); …

Lasioglossum (Dialictus) Semicaeruleum (Cockerell, 1895) (Hymenoptera: Halictidae) In Maryland: A Disjunct Population In Eastern North America?, Eugene J. Scarpulla

Insecta Mundi

Since 2004, three specimens of Lasioglossum (Dialictus) semicaeruleum (Cockerell, 1895) (Hyme­noptera: Halictidae) have been collected in Maryland. Other than three specimens from Wisconsin, there are no additional records of this western United States species known east of the Mississippi River. I document the three Maryland records and offer possible scenarios of how the specimens could have arrived in Mary­land.

Lasioglossum (Dialictus) semicaeruleum (Cockerell, 1895) (Hymenoptera: Halictidae) is an abundant western North American species that ranges from the Canadian Prairie Provinces (Alberta, Manitoba, and Saskatchewan) through the United States west of the Mississippi River (Arizona, California, Colorado, …

Genomics Reveals A New Genus And Species From A Single Female Specimen (Lepidoptera: Hesperiidae: Hesperiinae: Hesperiini: Moncina), Ernst Brockmann, Jing Zhang, Qian Cong, Nick V. Grishin

Insecta Mundi

New taxa in Hesperiidae (Lepidoptera: Papilionoidea) are traditionally proposed after inspection of male genitalia, which largely form the basis for Hesperiidae taxonomy. However, with genomic DNA se­quencing, even a single female specimen can be placed in a phylogenetic context of existing classification and taxonomically assigned with confidence. Genomic sequencing of an unusually patterned Hesperiidae female from San Martin, Peru, characterized by pearly spots outlining an inverted heart pattern on the rust-colored ventral hindwing, reveals that it represents an undescribed genus and species named here as Gemmia buechei Brockmann and Grishin, new genus and new species.

Neotropical regions are rich …

The Multiform Genus Psyrassa Pascoe (Coleoptera: Cerambycidae: Elaphidiini): New Species, New Records, Synonyms And Transfers, Kimberly García, Antonio Santos-Silva

Insecta Mundi

This paper presents a taxonomic review of the genus Boreocanthon Halffter, restored generic sta­tus, a group of ball-rolling (telocoprid) dung beetles (Coleoptera: Scarabaeidae: Scarabaeinae) endemic to North America. The genus, heretofore treated as a subgenus of Canthon, comprises 13 species, each keyed, diagnosed, illustrated and presented with information on distribution, relationships, and biology (*signifies restored generic combination): *Boreocanthon ateuchiceps (Bates, 1887); B. coahuilensis (Howden, 1966); *B. depressipennis (LeConte, 1859c); *B. ebenus (Say, 1823); *B. forreri (Bates, 1887); *B. integricollis (Schaef­fer, 1915); *B. lecontei (Harold, 1868); …

Erratum To Leblanc (2022): The Dacine Fruit Flies (Diptera: Tephritidae: Dacini) Of Oceania, Luc Leblanc

Insecta Mundi

After publication of Leblanc (2022), it was noticed that the wrong graph was used for the mean (±SE) daily captures of Dacus perpusillus (Drew) in cue-lure traps (Fig. 135). The author and the Insecta Mundi editorial staff apologize for this error. The intended version of Figure 135 is reproduced here. Insecta Mundi has also released a revised version of the Leblanc (2022) manuscript, with this error corrected. However, the revised version is not an official peer-reviewed article, and anyone wishing to reference the findings of Leblanc (2022) should cite the original manuscript or this erratum.

Revision Of Aneflomorpha Casey And Neaneflus Linsley (Coleoptera: Cerambycidae) Of The United States With An Illustrated Key To Species, Steven W. Lingafelter

Insecta Mundi

This paper presents a taxonomic review of the genus Boreocanthon Halffter, restored generic sta­tus, a group of ball-rolling (telocoprid) dung beetles (Coleoptera: Scarabaeidae: Scarabaeinae) endemic to North America. The genus, heretofore treated as a subgenus of Canthon, comprises 13 species, each keyed, diagnosed, illustrated and presented with information on distribution, relationships, and biology (*signifies restored generic combination): *Boreocanthon ateuchiceps (Bates, 1887); B. coahuilensis (Howden, 1966); *B. depressipennis (LeConte, 1859c); *B. ebenus (Say, 1823); *B. forreri (Bates, 1887); *B. integricollis (Schaef­fer, 1915); *B. lecontei (Harold, 1868); …

Review Of The Cybocephalidae (Coleoptera) Of North America And The West Indies With Descriptions Of Two New Species Of Cybocephalus Erichson, Trevor Randall Smith

Insecta Mundi

The nineteen species of Cybocephalidae (Coleoptera) occurring in North America and the West Indies (including Trinidad) are listed and keyed. Cybocephalus skelleyi new species and Cybocephalus edmondsoni new species are described, and Cybocephalus carrabeus T. R. Smith is found to be a new syn­onym of Cybocephalus geoffereysmithi T. R. Smith. Illustrations of morphological features, including detailed drawings of male genitalia, host records and distribution data, are provided.

The Referential Abbreviations Used By Linnaeus And Other Authors From 1758 Through 1779 For Describing Species Of Scarabaeoidea (Insecta: Coleoptera), Auke J. Hielkema

Insecta Mundi

A list of abbreviations regarding literature, collections and persons as used by early authors (1758–1779) of scarabaeoid beetles (Insecta: Coleoptera) is given together with modern referrals to the Literature Cited. Notes regarding referential errors are included. Hyperlinks to all mentioned and freely online available publications are provided.

The Dacine Fruit Flies (Diptera: Tephritidae: Dacini) Of Oceania, Luc Leblanc

Insecta Mundi

A key, an annotated checklist with detailed distribution, biological and host information, and color photographic plates are provided for the 91 species of dacine fruit flies (Diptera: Tephritidae: Dacini) known to occur in Oceania. Dacus virgatus Coquillett, previously a synonym of Bactrocera psidii (Froggatt), is in­stead considered a junior synonym of B. facialis (Coquillett). The species originally described in 1971 as Dacus (Asiadacus) perpusillus Drew, later reassigned as Bactrocera (Sinodacus) perpusilla (Drew) and in re­cent years as Zeugodacus (Sinodacus) perpusillus (Drew) actually belongs to genus Dacus, and is transferred back to Dacus, …

New Records Of Invasive Aphids (Hemiptera: Aphididae) On Garlic Mustard In The Usa, Doris Lagos-Kutz, Rebecah Troutman, Glen L. Hartman

Insecta Mundi

Lipaphis alliariae Müller (Hemiptera: Aphididae) was identified from specimens collected in Lake County, Ohio, on the invasive garlic mustard, Alliaria petiolata (Bieb.) Cavara & Grande (Brassicaceae). The identification was performed on apterous viviparae using morphological and molecular data. Body color and shape of the cauda discriminated L. alliariae and L. pseudobrassicae Davis. Measurements of morphological characters of both species were similar, but they have distinct characters distinguishing them from L. erysimi Kaltenbach. Neighbor-joining analysis of cytochrome oxidase 1 (Cox1) barcoding indicated a close relation­ship of the aphids that feed on Brassicaceae, and the range of pair-wise distances for Cox1 barcoding …

Natural History Of Cassida Sphaerula Boheman, 1854 (Coleoptera: Chrysomelidae: Cassidinae: Cassidini) On Arctotheca Prostrata (Salisb.) Britten (Asteraceae: Arctotidinae) In South Africa, With A Checklist Of South African Cassidinae (Leaf-Mining And Tortoise Beetles), Sally Adam, Mariana Campos, Hugh D.C. Heron, C. L. Staines, Rob Westerduijn, Caroline Simmrita Chaboo

Insecta Mundi

The tortoise beetle, Cassida sphaerula Boheman, 1854 (Coleoptera: Chrysomelidae: Cassidinae: Cassidini) is endemic to South Africa. Its endemic host, Arctotheca prostrata (Salisb.) Britten (Asteraceae) has been introduced in other countries where it is becoming invasive. Cassida sphaerula could provide a potential biocontrol of Arctotheca weeds as it spends the entire life cycle on this host. An intensive field study, with rearing, photography, and short films of C. sphaerula was conducted in its native habitat to document the life cycle. A checklist of Cassidinae genera in South Africa, along with 19 new host records for Cassidini species in South Africa are …

Synopsis Of The Tribe Amarotypini In New Zealand (Coleoptera: Carabidae), André Larochelle, Marie-Claude Larivière

Insecta Mundi

The tribe Amarotypini (Coleoptera: Carabidae: Migadopinae) is revised for New Zealand. Three genera and fourteen species are recognized.

Two genera and thirteen species are described as new: Amarophilus Larochelle and Larivière new genus, Amarophilus lomondensis Larochelle and Larivière new species, Amarophilus otagoensis Larochelle and Larivière new species, Amarophilus rotundicollis Larochelle and Larivière new species, Amarophilus wanakensis Larochelle and Larivière new species, Amarotypus fiordlandensis Larochelle and Larivière new species, Amarotypus glasgowensis Larochelle and Larivière new species, Amarotypus murchisonorum Larochelle and Larivière new species, Amarotypus simoninensis Larochelle and Larivière new species, Amarotypus takaheensis …

Whiteflies (Hemiptera: Aleyrodidae) Intercepted On Plant Product Imported To South Korea From 2013–2021, Suhyeon Lee, Soo-Jung Suh

Insecta Mundi

Current globalization and technological progress has facilitated and increased the international trade of plant products worldwide and has promoted the long-distance movement of immobile sucking pests such as whiteflies attached on plants. Therefore, being able to compile and update information on intercepted insect pests will help to improve the inspection procedures, to detect, identify and mitigate the damage caused by exotic invasive pests. Records of whiteflies (Hemiptera: Aleyrodidae) intercepted on import plants from 2013 to 2021 in the Pest Information System (PIS) database of South Korea were analyzed. A total of 32 species belonging to 19 genera were intercepted on …

A New Phyllophaga (Listrochelus) Timida Group Species From Baja California Sur (Coleoptera: Scarabaeidae: Melolonthinae), William B. Warner

Insecta Mundi

Phyllophaga (Listrochelus) baja, new species (Coleoptera: Scarabaeidae: Melolonthinae), is de­scribed and illustrated. Its relationship to other species in the “timida group” is discussed.

Morón (2002) reviewed the Phyllophaga Harris (Listrochelus Blanchard) species near P. timida (Horn) and included a key to the eight known species of the “timida group.” Many species in this group are difficult to dis­tinguish from one another via external characters; however, the male genital shape is diagnostic for each. Here I describe a new species from Baja California Sur, Mexico, that falls within that group.

A Review Of Host Plants For The Tortricid Tribe Grapholitini, With A Synopsis Of Host Utilization By Genus (Lepidoptera: Tortricidae), John W. Brown

Insecta Mundi

A database of larval host plants for the tortricid tribe Grapholitini (Lepidoptera: Tortricidae: Ole­threutinae) is presented, and larval hosts are summarized for each genus. Food plants have been reported for over 400 of the approximately 1,644 described species of Grapholitini. Of the 81 genera currently as­signed to the tribe, at least one larval host has been reported for 51. Ninety-seven different plant families have been reported at least once for a species of Grapholitini, with the greatest number of grapholitines recorded from Fabaceae (168 species), followed by Fagaceae (43 species), Pinaceae (43), Sapindaceae (36), Rosaceae (30), Asteraceae (30), Euphorbiaceae (15), …

A New Species Of Neostenoptera Meunier (Diptera: Cecidomyiidae: Winnertziinae) From Hawai‘I, John D. Plakidas, Nhu H. Nguyen, Michael L. Ferro

Insecta Mundi

A new paedogenetic midge (Diptera: Cecidomyiidae: Winnertziinae: Heteropezini) from O‘ahu Island, Hawai‘i, Neostenoptera hawaiiensis Plakidas, Nguyen, and Ferro, new species, is described and illus­trated. A key to all species in the genus is provided. Specimens were emergent from deadwood gathered at Waimea Arboretum and Botanical Garden. Neostenoptera appalachiensis Plakidas and Ferro were collected from the same set of samples in Hawai‘i, and additional specimens are reported from Georgia and South Carolina, three new state records. The discovery of two paedogenetic midges in Hawai‘i poses a unique set of questions as to their possible mode of arrival on an …

New Species, New Synonymies, A New Rank, And New Records In Desmiphorini, Saperdini, And Hemilophini (Coleoptera: Cerambycidae: Lamiinae), Antonio Santos-Silva, Robert A. Androw

Insecta Mundi

New information is presented for Neotropical Cerambycidae (Coleoptera). Dolichestola vittipennis Breuning, 1948 is synonymized with D. annulicornis Breuning, 1942, and the species is newly recorded from the Brazilian state of São Paulo. Dolichestola densepunctata Breuning, 1942 is newly recorded from Venezuela and Brazil, and the difference between it and D. annulicornis is reported. Mecas skillmani Santos-Silva and Androw, new species, is described from Mexico (Jalisco). Pannychella callicera (Bates, 1881) is illustrated, and notes on the genus and species are provided. Pannychis Thomson, 1864 is considered a genus different from Mecas LeConte, 1852, and notes on the genus …

Three New Species Of Cautethia Grote (Lepidoptera: Sphingidae) From The Lucayan Archipelago And Keys To West Indies Species, Jacqueline Y. Miller, Deborah L. Matthews, Riley J. Gott

Insecta Mundi

. Five species of Cautethia Grote (Lepidoptera: Sphingidae) occur in the Lucayan (Bahamas) Archipelago, three of which are new to science. Cautethia simoni Miller, Matthews, and Gott, new species, is described and illustrated from Mayaguana Island, Bahamas, and Providenciales and Grand Turk of the Turks and Caicos Islands. Cautethia gossi Miller, Matthews, and Gott, new species, is described and illustrated from Great Inagua, Bahamas. Cautethia geraceorum Miller, Matthews, and Gott, new species, is described from San Salvador Island. Diagnoses are provided and new island records are reported for the two previously described Bahamas species, Cautethia grotei Edwards …

A New Species Of Epeolus Latreille, 1802 (Hymenoptera: Apidae) From Western North America, Thomas M. Onuferko, Cory S. Sheffield

Insecta Mundi

A new species of Epeolus Latreille, 1802 (Hymenoptera: Apidae)—E. emiliae Onuferko and Shef­field, new species—is described from North America west of the Great Plains. It is morphologically most similar to Epeolus autumnalis Robertson, 1902, a species known exclusively from east of the Rocky Moun­tains. DNA barcode sequences from representatives of E. autumnalis and E. emiliae share a barcode index number (i.e., BIN: BOLD:AAF2361), but the two species exhibit marked and consistent differences in integu­ment coloration and the patterns of pubescence on the metasoma, and their distributional ranges, based on known specimens, show no overlap. This discovery increases …

Tomoxia Bucephala A. Costa (Coleoptera: Mordellidae), A Palearctic Tumbling Flower Beetle Established In North America, R. F. C. Naczi, Robert A. Androw, John Rosenfeld

Insecta Mundi

Tomoxia bucephala A. Costa (Coleoptera: Mordellidae), a Palearctic tumbling flower beetle native to Europe, Asia, and northernmost Africa, is now known from North America. The first known occurrences were in 2015 in Essex and Union counties, New Jersey, U.S.A. and in 2019 in Passaic County, New Jersey, all in the New York City metropolitan area. An additional collection documents the species in 2016 from Allegheny County, Pennsylvania, in the Pittsburgh metropolitan area. The multiple occurrences, the large distance between those in New Jersey and Pennsylvania, and multiple detections in natural areas indicate T. bucephala is established in North America and …

First Host Record For The Spider Wasp Cryptocheilus Severini Banks (Hymenoptera: Pompilidae: Pepsinae), Frank E. Kurczewski, Rick C. West, James P. Pitts, Cecilia Waichert

Insecta Mundi

The first host record for the North American spider wasp Cryptocheilus severini Banks (Hymenoptera: Pompilidae: Pepsinae) from Mazatlán, Sinaloa, México is introduced with pertinent observation information. The genus Cryptocheilus Panzer in North America is briefly described, its nesting habitat and prey transport outlined, and host specificity detailed.

The genus Cryptocheilus Panzer (Hymenoptera: Pompilidae: Pepsinae) comprises medium to rather large species of average stoutness (Townes 1957). The six Nearctic species of Cryptocheilus are all closely related (Townes 1957). In the Old World this genus is much richer, with 24 species and structural diversity that can present problems in identification from other …

First Record Of Ancognatha Erythrodera (Blanchard, 1846) (Coleoptera: Scarabaeidae: Dynastinae: Cyclocephalini) In The Altiplano Of Chile, José Mondaca, Sergio Rothmann

Insecta Mundi

The presence of Ancognatha erythrodera (Blanchard, 1846) (Coleoptera: Scarabaeidae) is confirmed for the first time in Chile based on male and female specimens collected in the extreme north of the country. This is the second species of Ancognatha Erichson, 1847 recorded in Chile. Morphological characters, illustrations of male genitalia, male and female habitus photographs of this species, and additional records in Argentina and Bolivia are provided. A map with the collection sites and montane habitats photograph in Chile are included.

The Dynastinae (Coleoptera: Scarabaeidae) that occur in Chile were comprehensively revised by Ratcliffe et al. (2021), who recorded nine species …